Basic info Taxonomic history Classification Included Taxa
Morphology Ecology and taphonomy External Literature Search Age range and collections

Osmyloidea

Insecta - Neuroptera

Taxonomy
Osmyloidea was named by Leach (1815). It is extant. Its type is Osmylus.

It was assigned to Neuroptera by Martins-Neto (2002); to Hemerobiiformia by Engel and Grimaldi (2008); and to Euneuroptera by Engel et al. (2018).

Synonymy list
YearName and author
1815Osmyloidea Leach
2002Osmyloidea Martins-Neto p. 38
2008Osmyloidea Engel and Grimaldi
2018Osmyloidea Engel et al.

Is something missing? Join the Paleobiology Database and enter the data

RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
Dicondylia
Paranotalia
subclassPterygota()
RankNameAuthor
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
Holometabola
Aparaglossata
Neuropteriformia
superorderNeuropterida
Metaxyneuroptera
Eidoneuroptera
orderNeuroptera
Euneuroptera
superfamilyOsmyloideaLeach 1815
superfamilyOsmyloideaLeach 1815

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. Osmyloidea Leach 1815
show all | hide all
Fm. †Archeosmylidae Riek 1953
+
G. †Archeosmylus Riek 1953
+
Archeosmylus costalis Riek 1955
G. †Babykamenia Ponomarenko and Shcherbakov 2004
+
Babykamenia eskovi Ponomarenko and Shcherbakov 2004
G. †Lithosmylidia Riek 1955
+
Lithosmylidia baronne Lambkin 1988
Lithosmylidia lineata Riek 1955
Lithosmylidia parvula Riek 1955
Fm. Nevrorthidae Nakahara 1958
+
G. †Balticoneurorthus Wichard 2016
+
Balticoneurorthus elegans Wichard 2016
G. †Cretarophalis Wichard 2017
+
G. †Electroneurorthus Wichard et al. 2010
+
Electroneurorthus malickyi Wichard et al. 2010
G. †Girafficervix Du et al. 2023
+
Girafficervix baii Du et al. 2023
Invalid names: Palaeoneurorthus Du et al. 2023 [replaced]
G. †Palaeoneurorthus Wichard 2009
+
Palaeoneurorthus eocaenus Wichard 2016
Palaeoneurorthus groehni Wichard et al. 2010
G. †Proberotha Krüger 1923
+
Proberotha dichotoma Wichard 2016
Proberotha prisca Krüger 1923
G. †Rophalis Pictet 1854
+
Rophalis relicta Pictet 1854
G. †Sisyroneurorthus Nakamine et al. 2023
+
Sisyroneurorthus aspoeckorum Nakamine et al. 2023
Fm. Osmylidae Leach 1815 [lance lacewing]
+
G. †Archaeosmylidia Makarkin et al. 2014
+
Archaeosmylidia fusca Makarkin et al. 2014
G. †Dimidiosmylus Khramov 2014
+
Dimidiosmylus ramosus Khramov 2014
G. †Dzhidosmylus Khramov 2020
+
Dzhidosmylus solus Khramov 2020
Subfm. Eidoporisminae Esben-Petersen 1917
+
G. †Stenosmylina Jepson et al. 2009
+
Stenosmylina medialis Jepson et al. 2009
G. †Erlikosmylus Khramov 2014
+
Erlikosmylus obscurus Khramov 2014
Subfm. Gumillinae Navas 1912
+
G. †Allotriosmylus Yang et al. 2010
+
Allotriosmylus uniramosus Yang et al. 2010
G. †Enodinympha Ren and Engel 2007
+
Enodinympha imperfecta Ren and Engel 2007
Enodinympha translucida Ren and Engel 2007
G. †Epiosmylus Panfilov 1980
+
Epiosmylus longicornis Panfilov 1980
Epiosmylus longus Khramov 2017
Epiosmylus panfilovi Ren and Yin 2002
G. Gumilla Navas 1912
G. †Kolbasinella Khramov 2014
+
Kolbasinella elongata Khramov 2014
G. †Nilionympha Ren and Engel 2007
+
Nilionympha pulchella Ren and Engel 2007
Nilionympha shantouensis Li et al. 2018
G. †Nuddsia Menon and Makarkin 2008
+
Nuddsia longiantennata Menon and Makarkin 2008
Nuddsia magnifica Myskowiak et al. 2016
Nuddsia repatriata Martins-Neto 2010
Invalid names: Burmaleon Myskowiak et al. 2016 [synonym]
G. †Osmylochrysa Jepson et al. 2012
+
Osmylochrysa anomala Jepson et al. 2012
Osmylochrysa fragilis Jepson et al. 2012
Osmylochrysa navasia Khramov 2020
G. †Tenuosmylus Wang et al. 2009
+
Tenuosmylus brevineurus Wang et al. 2009
Invalid names: Epiosmylinae Panfilov 1980 [synonym]
G. †Karaosmylus Makarkin 1990
+
Karaosmylus latus Panfilov 1980
Invalid names: Parosmylus Panfilov 1980 [replaced]
Subfm. Kempyninae Carpenter 1943
+
G. †Arbusella Khramov 2014
+
Arbusella bella Khramov 2014
Arbusella magna Khramov et al. 2017
Arbusella platyptera Ma and Wang 2022
G. †Cretosmylus Makarkin 1990
+
Cretosmylus sibiricus Makarkin 1990
G. †Euporismites Tillyard 1916
+
Euporismites balli Tillyard 1916
G. †Jurakempynus Wang et al. 2011
+
Jurakempynus arcanus Khramov 2014
Jurakempynus bellatulus Wang et al. 2011
Jurakempynus epunctatus Wang et al. 2011
Jurakempynus loculosus Ma et al. 2020
Jurakempynus sinensis Wang et al. 2011
Jurakempynus sublimis Khramov 2014
G. †Kempynosmylus Makarkin 2014
+
Kempynosmylus zherikhini Makarkin 2014
G. †Mirokempynus Ma et al. 2020
+
G. †Ponomarenkius Khramov et al. 2017
+
Ponomarenkius excellens Khramov et al. 2017
G. †Sauktangida Khramov 2014
+
Sauktangida aenigmatica Khramov 2014
G. †Kubekius Khramov 2017
+
Kubekius multiramosus Khramov 2017
G. †Mesosmylidus Jepson et al. 2012
+
Mesosmylidus vulgaris Jepson et al. 2012
Subfm. †Mesosmylininae Bode 1953
+
G. †Mesosmylina Bode 1953
+
Mesosmylina angusta Khramov 2014
Mesosmylina exornata Bode 1953
Mesosmylina falcifera Ansorge 1996
Mesosmylina mongolica Ponomarenko 1984
Mesosmylina shurabica Khramov 2014
Mesosmylina sibirica Ponomarenko 1985
Subfm. Osmylinae Leach 1815
+
G. †Lithosmylus Carpenter 1943
+
Lithosmylus columbianus Cockerell 1908
G. Osmylus Latreille 1802
G. †Vetosmylus Ma et al. 2020
+
Vetosmylus maculosus Ma et al. 2020
Vetosmylus tentus Ma et al. 2020
G. †Osmylopsis Handlirsch 1906
+
Osmylopsis duplicata Giebel 1856
+
Invalid names: Pterinoblattina binneyi Scudder 1886 [objective synonym]
G. †Palaeothyridosmylus Wang et al. 2009
+
G. †Petrushevskia Martynova 1958
+
Petrushevskia borisi Martynova 1958
G. †Pronymphites Panfilov 1980
+
Pronymphites elegans Panfilov 1980
Subfm. Protosmylinae Krüger 1913
+
G. †Juraheterosmylus Wang et al. 2010
+
Juraheterosmylus antiquatus Wang et al. 2010
Juraheterosmylus astictus Wang et al. 2010
Juraheterosmylus minor Wang et al. 2010
G. †Jurosmylus Makarkin and Archibald 2005
+
Jurosmylus atalantus Panfilov 1980
Jurosmylus parvulus Khramov 2014
Invalid names: Mesosmylus Panfilov 1980 [replaced]
G. †Osmylidia Cockerell 1908
+
Osmylidia donnae Makarkin et al. 2021
Osmylidia glastrai Makarkin et al. 2021
Osmylidia picta Hagen 1856
Osmylidia requieta Scudder 1890
Osmylidia taliae Makarkin et al. 2021
Invalid names: Oligosmylus Krüger 1913 [objective synonym], Protosmylus Krüger 1913 [synonym]
G. †Protosmylina Jepson et al. 2009
+
Protosmylina bifasciata Jepson et al. 2009
G. †Pseudosmylidia Makarkin 2017
+
Pseudosmylidia relicta Makarkin 2017
G. †Sogjuta Martynova 1958
+
Sogjuta shartegica Khramov 2011
Sogjuta speciosa Martynova 1958
G. †Scapoptera Panfilov 1980
+
Scapoptera recta Panfilov 1980
Subfm. Spilosmylinae Kruger 1913
+
G. †Ensiosmylus Khramov 2014
+
Ensiosmylus acutus Khramov 2014
G. †Imanosmylus Makarkin 1994
+
Imanosmylus ussuriensis Makarkin 1994
G. †Zakamnosmylus Khramov 2020
+
Zakamnosmylus elongatus Khramov 2020
G. †Stenochrysa Jepson et al. 2012
+
Stenochrysa gradata Jepson et al. 2012
G. †Tengriosmylus Khramov 2014
+
Tengriosmylus magnificus Khramov 2014
G. †Tetanoptilon Bode 1953
+
Fm. †Saucrosmylidae Ren and Yin 2003 [elegant lance lacewing]
+
G. †Bellinympha Wang et al. 2010
+
Bellinympha dancei Wang et al. 2010
Bellinympha filicifolia Wang et al. 2010
G. †Daohugosmylus Liu et al. 2014
+
Daohugosmylus castus Liu et al. 2014
G. †Huiyingosmylus Liu et al. 2013
+
Huiyingosmylus bellus Liu et al. 2013
G. †Laccosmylus Ren and Yin 2003
+
Laccosmylus calophlebius Ren and Yin 2003
Laccosmylus cicatricatus Fang et al. 2018
Laccosmylus latizonus Fang et al. 2018
G. †Lebanosmylus Azar and Nel 2022
+
Lebanosmylus leae Azar and Nel 2022
G. †Rudiosmylus Ren and Yin 2003
+
Rudiosmylus ningchengensis Ren and Yin 2003
G. †Saucrosmylus Ren and Yin 2003
+
Saucrosmylus sambneurus Ren and Yin 2003
G. †Thaumatomerobius Ponomarenko 1985
+
Thaumatomerobius mirabilis Ponomarenko 1985
G. †Ulrikezza Fang et al. 2015
+
Ulrikezza aspoeckae Fang et al. 2015
Fm. Sisyridae Banks 1905 [spongillafly]
+
Subfm. †Cretadilarinae Makarkin 2017
+
G. †Cretadilar Makarkin 2017
+
Cretadilar olei Makarkin 2017
G. †Microsisyra Makarkin 2022
+
Microsisyra carsteni Makarkin 2022
G. †Paleosisyra Nel et al. 2003
+
Paleosisyra eocenica Nel et al. 2003
Paleosisyra minor Wichard et al. 2016
Subfm. †Paradoxosisyrinae Makarkin 2016
+
G. †Buratina Khramov 2019
+
Buratina lusitanica Termier et al. 1985
Buratina truncata Khramov 2019
G. †Khobotun Khramov 2019
+
Khobotun elephantinus Khramov 2019
G. †Paradoxosisyra Makarkin 2016
+
Paradoxosisyra groehni Makarkin 2016
G. †Protosiphoniella Khramov 2019
+
G. †Sidorchukatia Khramov 2019
+
Sidorchukatia gracilis Khramov 2019
G. †Prosisyrina Perkovsky and Makarkin 2015
+
Prosisyrina sphinga Makarkin and Perkovsky 2016
Prosisyrina sukachevae Perkovsky and Makarkin 2015
G. Sisyra Burmeister 1839
+
Sisyra amissa Hagen 1856
G. †Stictosisyra Yang et al. 2018
+
Stictosisyra pennyi Yang et al. 2018
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Changhsingian to the top of the Messinian or 254.17000 to 5.33300 Ma
Minimum age of oldest fossil (stem group age): 252.17 Ma

Collections (127 total)


Time interval Ma Country or state Original ID and collection number
Changhsingian254.17 - 252.17Australia (New South Wales) Archeosmylus pectinatus (122433)
Changhsingian254.17 - 252.17South Africa (Kwazulu-Natal) Archeosmylus sp. (14361)
Changhsingian254.17 - 252.17South Africa (Natal) Archeosmylidae indet. (124428)
Griesbachian252.17 - 251.3Russian Federation (Kemerovo) Babykamenia eskovi (104713)
Anisian247.2 - 242.0Australia (Queensland) Lithosmylidia baronne (154203)
Norian228.0 - 208.5Australia (Queensland) Archeosmylus sp. (111558) Lithosmylidia parvula (154204) Lithosmylidia sp. A, Archeosmylus stigmatus, Archeosmylus costalis, Lithosmylidia lineata, Lithosmylidia parvula (106484)
Hettangian - Sinemurian201.3 - 190.8Kyrgyzstan (Tonskiy) Petrushevskia borisi, Sogjuta speciosa (105087)
Sinemurian199.3 - 190.8Kyrgyzstan (Tonskiy) Petrushevskia borisi (113412)
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) Mesosmylina exornata (105002) Tetanoptilon brunsvicense (107323)
Early/Lower Toarcian183.0 - 182.0Germany (Mecklenburg-Vorpommern) Mesosmylina falciferum (114993)
Toarcian182.7 - 174.1Kyrgyzstan (Batken) Erlikosmylus obscurus (118023) Mesosmylina shurabica (105003) Tengriosmylus magnificus, Mesosmylina angusta (110817)
Toarcian - Aalenian182.7 - 170.3Kyrgyzstan (Osh) Sauktangida aenigmatica (156499)
Bajocian170.3 - 168.3France Neuropora sp. (34712 34714)
Bajocian - Bathonian170.3 - 166.1Russian Federation (Krasnoyarsk) Osmylidae indet., Thaumatomerobius mirabilis, Mesosmylina sibirica, Kubekius multiramosus, Epiosmylus longus (118010)
Bajocian - Bathonian170.3 - 166.1Mongolia (Bayan-Khongor) Saucrosmylidae indet. (114615)
Bathonian168.3 - 166.1Mongolia Mesosmylina mongolica (119621)
Bathonian168.3 - 166.1Romania (Central Dobrogea) Neuropora spinosa (38190)
Late/Upper Bathonian167.7 - 164.7France Neuropora spinosa, Neuropora sp. erect form (37510)
Late/Upper Bathonian167.7 - 164.7India (Gujarat) Neuropora sp. (93561)
Late/Upper Bathonian167.7 - 164.7India (Kachchh) Neuropora sp. (24050)
Callovian - Oxfordian166.1 - 157.3China (Nei Mongol) Allotriosmylus uniramosus, Archaeosmylidia fusca (113365) Huiyingosmylus bellus, Daohugosmylus castus, Ponomarenkius sp., Ponomarenkius excellens, Arbusella magna, Palaeoneurorthus baii (113371) Jurakempynus sinensis, Jurakempynus bellatulus, Jurakempynus epunctatus, Bellinympha filicifolia, Bellinympha dancei, Juraheterosmylus antiquatus, Juraheterosmylus astictus, Juraheterosmylus minor (113366) Laccosmylus calophlebius, Laccosmylus cicatricatus, Laccosmylus latizonus (196979) Mirokempynus profundobifurcus, Jurakempynus loculosus, Arbusella platyptera (219840) Nilionympha shantouensis (122589) Palaeoneurorthus baii (229433) Saucrosmylus sambneurus, Rudiosmylus ningchengensis, Laccosmylus calophlebius, Enodinympha translucida, Nilionympha pulchella, Nilionympha imperfecta, Palaeothyridosmylus sp., Palaeothyridosmylus septemaculatus, Epiosmylus panfilovi, Tenuosmylus brevineurus (113367) Ulrikezza sp., Ulrikezza aspoeckae (172659) Vetosmylus tentus, Vetosmylus maculosus (210302)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Arbusella sp. (106279) Ensiosmylus acutus (113439) Jurosmylus parvulus, Jurakempynus arcanus, Arbusella sp. (105970) Jurosmylus parvulus, Kolbasinella elongata, Arbusella bella (105426) Mesosmylus atalantus, Epiosmylus longicornis (106101) Pronymphites elegans, Parosmylus latus (105971) Scapoptera recta, Jurosmylus parvulus (106102)
Early/Lower Callovian164.7 - 161.2India Neuropora sp. (67608)
Late/Upper Callovian164.7 - 161.2India Neuropora sp. (68291)
Middle Callovian - Early/Lower Kimmeridgian164.7 - 150.8Iran (Tabas Area) Neuropora sp. (41417)
Late/Upper Oxfordian161.2 - 155.7France Neuropora spinosa (67975)
Late/Upper Oxfordian161.2 - 155.7Romania Neuropora sp., Neuropora spinulosa (35199)
Late/Upper Oxfordian161.2 - 155.7Portugal Neuropora lusitanica (68246)
Late/Upper Oxfordian161.2 - 155.7Poland Neuropora sp. (60853 60854 60855)
Late/Upper Oxfordian161.2 - 155.7Germany (Bavaria) Neuropora sp. (25411)
Middle Oxfordian161.2 - 155.7Germany (Niedersachsen) Neuropora sp., Neuropora lusitanica (31435)
Middle Oxfordian161.2 - 155.7France Neuropora spinosa (68642 163585)
Late/Upper Oxfordian - Kimmeridgian161.2 - 152.1Portugal Neuropora lusitanica (68245)
Late/Upper Oxfordian - Valanginian161.2 - 132.9China (Xizang) Neuropora sp. (61414)
Kimmeridgian157.3 - 152.1Portugal (Centro) Neuropora spinosa, Neuropora lusitanica (68220)
Kimmeridgian157.3 - 152.1Portugal Neuropora lusitanica (68784 68787 68819) Neuropora sp. (68212) Neuropora spinosa (68207 68242)
Kimmeridgian - Early/Lower Tithonian157.3 - 145.0Romania Neuropora lusitanica (201710 201711)
Late/Upper Kimmeridgian155.7 - 150.8Austria Neuropora lusitanica (36223)
Tithonian152.1 - 145.0Mongolia (Dund-Gov’ Aymag) Saucrosmylidae indet. (122321)
Tithonian152.1 - 145.0Mongolia Jurakempynus sublimis (125978) Osmylidae indet., Sogjuta shartegica, Dimidiosmylus ramosus (104748)
Tithonian152.1 - 145.0Russian Federation (Chita) Mesosmylina sp. (125407)
Late/Upper Tithonian - Berriasian150.8 - 139.8Austria Neuropora lusitanica (144461)
Early/Lower Berriasian145.0 - 140.2United Kingdom (England) Orthoptera indet. (105966)
Late/Upper Berriasian145.0 - 140.2United Kingdom (England) Mesosmylidus vulgaris, Osmylochrysa anomala, Osmylochrysa fragilis, Stenochrysa gradata (122909)
Late/Upper Hauterivian136.4 - 130.0United Kingdom (England) Stenosmylina medialis (106417)
Early/Lower Barremian130.0 - 125.45United Kingdom (England) Protosmylina bifasciata (118921)
Aptian125.0 - 113.0Russian Federation (Buryatia) Cretosmylus sibiricus (119498) Kempynosmylus zherikhini (106103) Osmylochrysa navasia, Zakamnosmylus elongatus, Dzhidosmylus solus (113350)
Late/Upper Aptian122.46 - 112.03Brazil (Ceará) Nuddsia longiantennata (122557)
Late/Upper Aptian122.46 - 112.03Brazil (Ceara) Nuddsia repatriata (34901)
Early/Lower Cenomanian99.6 - 93.5Myanmar Buratina truncata, Sidorchukatia gracilis, Protosiphoniella anthophila, Khobotun elephantinus (190961) Burmaleon magnificus (161726) Cretarophalis patrickmuelleri (123053 167760 178159) Nevrorthidae indet. (184692 184693) Osmylidae indet. (111688) Paradoxosisyra groehni, Cretadilar olei, Microsisyra carsteni (172900) Sisyroneurorthus aspoeckorum (196491) Stictosisyra pennyi (165561)
Late/Upper Cenomanian99.6 - 93.5Lebanon Lebanosmylus leae (229466)
Late/Upper Santonian85.8 - 83.5Russian Federation Prosisyrina sp. (35055) Prosisyrina sukachevae, Prosisyrina sphinga (35056)
Maastrichtian72.1 - 66.0Russian Federation (Primorskiy) Imanosmylus ussuriensis (119699)
Ypresian56.0 - 47.8Canada (British Columbia) Osmylidia donnae (31677) Osmylidia sp. (104793)
Ypresian56.0 - 47.8Australia (Queensland) Euporismites balli (111780 125666 139941)
Ypresian56.0 - 47.8France (Oise) Paleosisyra eocenica (123850)
Ypresian56.0 - 47.8USA (Washington) Osmylidia glastrai (103206)
Bridgerian50.3 - 46.2USA (Colorado) Osmylidia taliae (181561)
Priabonian38.0 - 33.9Ukraine Rophalis relicta (122997)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Electroneurorthus malickyi, Palaeoneurorthus groehni, Proberotha prisca, Paleosisyra electrobaltica, Palaeoneurorthus bifurcatus, Proberotha dichotoma, Osmylidia picta (110735) Palaeoneurorthus eocaenus, Balticoneurorthus elegans (120814) Paleosisyra electrobaltica (140480) Paleosisyra minor (123928) Proberotha prisca (123766) Rophalis relicta (123439) Rophalis relicta, Electroneurorthus malickyi, Proberotha prisca, Paleosisyra electrobaltica, Palaeoneurorthus hoffeinsorum (124468) Rophalis relicta, Proberotha prisca (123420)
Priabonian38.0 - 33.9Russian Federation Palaeoneurorthus hoffeinsorum (123419)
Priabonian38.0 - 33.9Poland Sisyra relicta, Osmylus pictus, Proberotha prisca, Sisyra amissa (120811) Sisyra relicta, Proberotha prisca (123438)
Priabonian38.0 - 33.9United Kingdom (England) Sisyridae sp.A (21976)
Chadronian37.2 - 33.9USA (Colorado) Lithosmylus columbianus (151548) Osmylidia requieta (113749 124624) Osmylidia requietus (106379) Osmylus columbianus (106453) Osmylus requietus (106384) Pseudosmylidia relicta (106381)
Sarmatian12.7 - 11.608Romania Osmylus sp. (156756)
Messinian7.246 - 5.333France (Cantal) Sisyra sp. (122195)