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Pugnax

Rhynchonellata - Rhynchonellida - Pugnacidae

Taxonomy
Pugnax is a genus. It is not extant. It is the type genus of Pugnacidae.

It was assigned to Rhynchonellida by Sepkoski (2002); and to Pugnacidae by Williams et al. (2002).

Synonyms
Synonymy list
YearName and author
2002Physetorhynchia Sepkoski
2002Pugnax Sepkoski
2002Striatopugnax Sepkoski
2002Pugnax Williams et al. p. 1165

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RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Spiralia
superphylumLophotrochozoa
—Lophophorata
—PanbrachiopodaCarlson and Cohen 2020
RankNameAuthor
phylumBrachiopodaCuvier 1805
subphylumRhynchonelliformeaWilliams et al. 1996
classRhynchonellataWilliams et al. 1996
orderRhynchonellidaKuhn 1949
superfamilyPugnacoideaRzhonsnitskaia 1956
familyPugnacidaeRzonsnitzkaja 1956
genusPugnax

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Pugnax
show all | hide all
†Pugnax kayseri Tschernyschew 1902
†Pugnax keyserlingi Moeller 1862
†Pugnax latesinuata Feng 1986
†Pugnax morahensis Waagen 1883
†Pugnax osagensis Swallow 1908
†Pugnax sublaevis Mansuy 1913
†Pugnax utah Marcou 1858
Invalid names: Physetorhyncha [synonym], Striatopugnax [synonym]
Diagnosis
No diagnoses are available
Measurements
partNmeanminimummaximummedians.d.
shell height56.374.608.206.601.32

Composition: low Mg calcitep
Environment: marineuc
Locomotion: stationaryo
Attached: yesp
Life habit: epifaunalo
Diet: suspension feedero
Vision: blindc
Created: 2004-02-28 14:12:25
Modified: 2004-02-28 16:12:25
Source: o = order, c = class, p = phylum, uc = unranked clade
References: Nesnidal et al. 2013, Aberhan et al. 2004

Age range: base of the Emsian to the top of the Changhsingian or 407.60000 to 252.17000 Ma

Collections (50 total)


Time interval Ma Country or state Original ID and collection number
Helderbergian - Emsian419.2 - 393.3Austria P. sp. (26203)
Lochkovian - Emsian419.2 - 393.3Austria (Carnic Alps) P. preacuminatus, P. scupini (77909)
Pragian - Emsian410.8 - 393.3Tajikistan P. julii (45831)
Emsian407.6 - 393.3Australia (New South Wales) P. oepiki (39884 39888)
Middle Devonian393.3 - 382.7USA (Alaska) P. sp. (111901)
Givetian387.7 - 382.7Russian Federation (Kemerovskaya Oblast) P. ephippioides (84384)
Frasnian382.7 - 372.2Canada (Ontario) P. sp. (633)
Frasnian382.7 - 372.2China (Guangxi) P. lummatoensis (40574)
Middle Frasnian382.4 - 379.5France (Alsace) P. kayseri (65766)
Late/Upper Givetian380.2 - 382.7Germany (Nordrhein-Westfalen) P. sp., P. acuminatus (82561)
Late/Upper Frasnian379.5 - 376.1Russian Federation (Kemerovskaya Oblast) P. acuminata, P. anisodonta, P. tensa, P. voroni (84378)
Late/Upper Frasnian379.5 - 376.1Russian Federation (Novosibirskaya Oblast) P. tensa, P. mesogonia (84381)
Early/Lower Famennian376.1 - 370.6Canada (Alberta) P. rara (35541)
Early/Lower Famennian376.1 - 370.6Russian Federation P. sp. (61964)
Middle Famennian370.6 - 364.7Canada (Alberta) P. minutus (35543)
Late/Upper Famennian364.7 - 360.7USA (Colorado) P. sp. (644)
Tournaisian358.9 - 346.7Australia (New South Wales) P. minutus (5615)
Ivorian - Chadian353.8 - 340.0Ireland P. platylobus, P. sulcatus (76501)
Chadian346.7 - 340.0United Kingdom (England) P. acuminatus (5972) P. cordiformis, P. platilobus, P. plicatilis, P. reniformis (5983) P. sp. (5971)
Asbian340.0 - 336.0United Kingdom (England) P. accuminatus (5995)
Asbian - Brigantian340.0 - 330.9United Kingdom (England) P. acuminatus (28383) P. cordiformis, P. triplex (5987)
Brigantian336.0 - 330.9United Kingdom (England) P. acuminatus, P. codiformis, P. pseudopugnus, P. sp., P. pugnoides (76172) P. sp. (57131)
Bashkirian323.2 - 315.2USA (Utah) Terebratula uta (150953)
Gzhelian303.7 - 298.9China (Guangxi) P. latesinuata (134347)
Virgilian303.7 - 298.9USA (Ohio) P. utah (69720)
Gzhelian - Artinskian303.7 - 279.3Bolivia Rhynchonella sp. (120479)
Asselian298.9 - 295.5Bolivia P. sp. (128633)
Asselian - Sakmarian298.9 - 290.1Laos P. osagensis, P. sublaevis (233571)
Permian298.9 - 251.902USA (Alaska) P. utah (112199)
Sakmarian293.52 - 290.1China (Guangxi) P. sp. (134319)
Roadian272.3 - 268.8USA (Texas) P. osagensis (108803 108822)
Roadian - Capitanian272.3 - 259.51USA (Texas) P. osagensis (108829)
Wordian266.9 - 264.28Pakistan P. sp. (125854)
Wordian266.9 - 264.28USA (Texas) P. osagensis (108821)
Wuchiapingian259.9 - 254.17Pakistan P. morahensis (125895 125901 126007) P. sp. (125898 125910) Rhynchonella morahensis (119541 119850)
Wuchiapingian259.9 - 254.17Azerbaijan P. utah (132459)
Changhsingian254.17 - 252.17China (Guizhou) P. sp. (149293)
Changhsingian254.17 - 252.17China (Hubei) P. sp. (120809)