PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Cycadophytanae

Taxonomy

Cycadophytanae was named by Doweld (2001). It is not extant.

It was assigned to Pteridobiotina by Doweld (2001).

Subtaxa

Aneurophytophyta (syn. Archaeopteridophyta), Cycadeoideophyta, Gnetophyta, Magnoliophyta, Moresnetiophyta, Noeggerathiophyta, Peltaspermophyta

Synonymy list

Year	Name and author
2001	Cycadophytanae Doweld

Is something missing? Join the Paleobiology Database and enter the data

Rank	Name	Author
kingdom	Plantae
phylum	Tracheophyta	(Sinnott 1935)
—	Pteridobiotina

Rank	Name	Author
superphylum	Cycadophytanae
superphylum	Cycadophytanae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. †Cycadophytanae Doweld 2001

show all | hide all

Ph. †Aneurophytophyta Bold 1973

Subph. †Aneurophytophytina Doweld 2001

Cl. †Aneurophytopsida Takhtajan 1978

Or. †Aneurophytales Beck 1957

Fm. †Aneurophytaceae Ananiev 1963

G. †Milleria Lang 1926

Subph. †Archaeopteridophytina Doweld 2001

Cl. †Archaeopteridopsida Takhtajan 1978

Or. †Archaeopteridales Zimmermann 1930

Fm. †Archaeopteridaceae Nicholson and Lydekker 1889

G. †Dzungaropteris Doweld 2001

Invalid names: Neuburgia Radtschenko 1969 [replaced]

G. †Pterispermotheca Carpentier 1919

G. †Archaeopteridium Kidston 1923

†Archaeopteridium tschermaki Stur 1923

Invalid names: Archaeopteridophyta Snigirevskaya 2000 [synonym]

Ph. †Cycadeoideophyta Taylor 1981

Cl. †Cycadeoideopsida Scott 1923

Or. †Cycadeoideales Berry 1916

G. †Bucklandia Brongniart 1828

†Bucklandia bucklandii Doweld 2012

Invalid names: Conites bucklandii Sternberg 1823 [objective synonym]

†Bucklandia florovia Watson and Sincock 1992

G. †Conites Sternberg 1823

†Conites rajmahalensis Sahni 1928

†Conites sessilis Sahni 1928

†Conites sripermaturensis Sahni 1928

Invalid names: Conites cutchensis Sahni 1926 [nomen nudum], Conites hobsoni Sahni 1932 [nomen nudum]

G. †Pseudocycadeoidea Aiyengar and Jacob 1952

†Pseudocycadeoidea indica Aiyengar and Jacob 1952

G. †Yabeiella Oishi 1931

†Yabeiella brackebushiana Kurtz 1921

†Yabeiella crassa Jones and de Jersey 1947

†Yabeiella hirsuta Bose and Sukh-Dev 1959

†Yabeiella indica Pal 1984

†Yabeiella mareyesiaca Geinitz 1876

Invalid names: Taeniopteris dunstoni Walkom 1917 [synonym]

†Yabeiella spatulata Oishi 1931

†Yabeiella wielandi Oishi 1931

Invalid names: Sturianthaceae Doweld 2001 [empty]

G. †Cycadeoidella Ogura 1930

†Cycadeoidella japonica Ogura 1930

Invalid names: Bennettitopsida Engler 1892 [synonym]

Ph. Gnetophyta Bessey 1907

Cl. Gnetopsida Kirpotenko 1884

G. †Gontriglossa Anderson and Anderson 1989

†Gontriglossa reinerae Nielsen 2005

G. †Pityophyllum Nathorst 1899

†Pityophyllum nordenskioeldi Heer 1899

†Pityophyllum staratschini Nathorst 1899

Ph. Magnoliophyta Cronquist et al.

G. †Andreisporis Belsky et al. 1965

†Andreisporis mariae Belsky et al. 1965

Unr. Anthophyta

G. †Aquilapollenites Rouse 1957

†Aquilapollenites minimus Belsky et al. 1965

†Aquilapollenites quadrilobus Rouse 1957

G. †Bevhalstea

G. †Cranwellia Srivastava 1966

Unr. †Liliopsida Kenrick and Crane 1997

Unr. †Alismatidae

Invalid names: Hydrocharitales Dumort 1829 [empty], Triuridales [empty]

G. †Limnocarpus Reid 1898

†Limnocarpus cooperi Chandler 1961

†Limnocarpus enormis Chandler 1961

†Limnocarpus forbesi Heer 1862

†Limnocarpus headonensis Gardner 1898

†Limnocarpus magnus Chandler 1961

Invalid names: Liliophyceae Kenrick and Crane 1997 [replaced]

G. †Loranthacites Mtchedlishvili 1961

†Loranthacites nataliae Salard-Cheboldaeff 1978

G. †Araripia Mohr and Eklund 2003

†Araripia florifera Mohr and Eklund 2003

G. †Asteropollis Hedlund and Norris 1968

†Asteropollis asteroides Hedlund and Norris 1968

†Asteropollis transitorius Vallati 2006

†Asteropollis vulgaris Singh 1983

G. †Auriculiidites Elsik 1964

†Auriculiidites hourcqui Boltenhagen 1967

†Auriculiidites laevigatus Reyre 1964

†Auriculiidites reticulatus Elsik 1964

G. †Auriculopollenites Legoux et al. 1970

†Auriculopollenites reticulatus Salard-Cheboldaeff 1978

G. †Bacumorphomonocolpites Sole de Porta 1971

†Bacumorphomonocolpites tausae Sole de Porta 1971

G. †Balanopollis Salard-Cheboldaeff 1978

†Balanopollis minutus Salard-Cheboldaeff 1978

G. Bignonia Linnaeus 1753

Bignonia aequinoctialis Linnaeus 1753

Bignonia caerulea Linnaeus 1753

Bignonia catalpa Linnaeus 1753

Bignonia crucigera Linnaeus 1753

Bignonia espreolata Linnaeus 1753

†Bignonia gigantifolia Engelhardt 1891

Bignonia indica Linnaeus 1753

Bignonia loucoxylon Linnaeus 1753

Bignonia paniculata Linnaeus 1753

Bignonia peruviana Linnaeus 1753

Bignonia radiata Linnaeus 1753

Bignonia radicans Linnaeus 1753

Bignonia sempervireus Linnaeus 1753

Bignonia unguiscati Linnaeus 1753

G. †Brenneripollis Juhász and Goczan 1985

†Brenneripollis pellitusreticulatus Juhász and Goczan 1985

G. †Brevicolporites Anderson 1960

†Brevicolporites guinetti Salard-Cheboldaeff 1978

†Brevicolporites molinae Salard-Cheboldaeff 1978

G. †Ceratocarpus Velenovsky and Viniklar 1931

†Ceratocarpus fendrychi Velenovsky and Viniklar 1931

G. †Constructipollenites van Hoeken-Klinkenberg 1964

†Constructipollenites ineffectus van Hoeken-Klinkenberg 1964

G. †Cornetipollis Pocock and Vasanthy 1988

†Cornetipollis herngreenii Deaf et al. 2014

†Cornetipollis perforata Dino 1992

G. †Crassitriapertites Herngreen 1972

†Crassitriapertites polymorphus Herngreen 1972

†Crassitriapertites semitectatus Herngreen 1972

†Crassitriapertites uesugii Herngreen 1972

†Crassitriapertites vanderhammeni Herngreen 1972

G. †Crassitricolporites Herngreen 1972

†Crassitricolporites brasiliensis Herngreen 1972

†Crassitricolporites subprolatus Herngreen 1972

G. †Cristaecolpites Schrank 1994

†Cristaecolpites echinaceus Schrank 1994

G. †Cristaeturites Schrank 1994

†Cristaeturites cristatus Schrank 1994

G. †Crusafontites Sole de Porta 1971

†Crusafontites megagemmatus Jaramillo and Dilcher 2001

G. †Cupuliferoidaepollenites Potonié et al. 1950

†Cupuliferoidaepollenites liblarensis Potonie 1960

†Cupuliferoidaepollenites minutus Singh 1971

†Cupuliferoidaepollenites parvulus Dettman 1973

Invalid names: Psilatricolpites parvulus Norris 1967 [synonym]

†Cupuliferoidaepollenites quisqualis Potonie 1960

G. †Dettmannaepollenites Kedves 1999

†Dettmannaepollenites clavatus Schrank and Mahmoud 2000

†Dettmannaepollenites intrabaculatus Kedves 1999

G. †Dicolpopollis Pflanzl 1956

†Dicolpopollis malesianus Muller 1968

G. †Dryadopollis Srivastava 1977

†Dryadopollis argus Srivastava 1975

†Dryadopollis inconspicuus Burger 1993

†Dryadopollis minima Barreda 1997

G. †Echitricolpites Regali et al. 1974

G. †Foveomonocolpites Wijmstra 1971

†Foveomonocolpites bauchiensis Adegoke 1978

G. †Foveotetradites Singh 1983

†Foveotetradites fistulosus Singh 1983

G. †Foveotricolpites Pierce 1961

†Foveotricolpites genuinus González-Guzmán 1967

†Foveotricolpites giganteus Du Chene 1978

†Foveotricolpites gigantoreticulatus Schrank 1987

†Foveotricolpites perforatus Van der Hammen and Garcia de Mutis 1966

†Foveotricolpites pomarius Van der Hammen and Garcia de Mutis 1966

†Foveotricolpites prolatus Frederiksen 1973

†Foveotricolpites santanderianus Van der Hammen and Garcia de Mutis 1966

†Foveotricolpites tienabaensis Schrank 1987

G. †Fraxinoipollenites Potonié 1960

†Fraxinoipollenites constrictus Chlonova 1976

†Fraxinoipollenites fragilis Burger 1993

†Fraxinoipollenites helbyi Burger 1993

†Fraxinoipollenites variformis Norvick and Burger 1975

†Fraxinoipollenites venustus Singh 1971

G. †Hammenia Ward 1986

G. †Hexacolpites Erdtman 1947

†Hexacolpites echinatus Salard-Cheboldaeff 1978

†Hexacolpites indicus Ramanujam 1966

G. †Holkopollenites Stover 1966

†Holkopollenites chemardensis Stover et al. 1966

†Holkopollenites forix Christopher and Prowell 2010

†Holkopollenites irregularis Christopher and Prowell 2002

†Holkopollenites propinquus Christopher and Prowell 2010

G. †Inaperturotetradites van Hoeken-Klinkenberg 1964

†Inaperturotetradites lacunosus van Hoeken-Klinkenberg 1964

†Inaperturotetradites psilatus Ramanujam 1966

†Inaperturotetradites reticulatus Salard-Cheboldaeff 1978

G. †Integritetradites Schrank and Mahmoud 2000

†Integritetradites porosus Schrank and Mahmoud 2000

G. †Intrareticulitis Kar 1985

†Intrareticulitis brevis Kar 1985

G. †Jugella Mchedlishvili and Shakmundes 1973

†Jugella caichiguensis Schrank 2010

†Jugella semistriata Schrank 2010

†Jugella sibirica Mtchedlishvili and Shakmundes 1973

G. †Lethomasites Ward 1989

G. †Microfoveotriporites van Hoeken-Klinkenberg 1966

†Microfoveotriporites cretaceus van Hoeken-Klinkenberg 1966

G. †Monocolpopollenites Pflug and Thomson 1953

†Monocolpopollenites ovatus Jaramillo and Dilcher 2001

†Monocolpopollenites sphaeroidites Jardiné and Magloire 1965

†Monocolpopollenites spheroidites Jardiné and Magloire 1965

†Monocolpopollenites tranquilloides Nichols 1973

G. †Multiporopollenites Pflug 1953

†Multiporopollenites pauciporatus Jaramillo and Dilcher 2001

G. †Nigeripollis Srivastava 1975

†Nigeripollis gemmatus Salami 1985

G. †Nyssapollenites Thiergart 1937

†Nyssapollenites albertensis Singh 1971

†Nyssapollenites barooahii Sah and Dutta 1968

†Nyssapollenites coromandelinum Ramanujam 1966

†Nyssapollenites endobalteus Mildenhall and Pocknall 1989

†Nyssapollenites incertus Dutta and Sah 1970

†Nyssapollenites kutchensis Venkatachala and Kar 1969

†Nyssapollenites lanosus Dettmann 1973

†Nyssapollenites lehmanii Schrank 1994

†Nyssapollenites pachyexinus Baldoni and Askin 1993

†Nyssapollenites scabratus De Benedetti et al. 2023

†Nyssapollenites squamosus Dettmann 1973

†Nyssapollenites triangulus Singh 1983

G. †Pediculisporis Belsky et al. 1965

†Pediculisporis reticularis Belsky et al. 1965

†Pediculisporis rnicrogranulatus Belsky et al. 1965

G. †Penetetrapites Hedlund and Norris 1968

†Penetetrapites incipiens de Lima 1989

†Penetetrapites mollis Hedlund and Norris 1968

G. †Phimopollenites Dettmann 1973

†Phimopollenites augathallaensis Dettman 1973

†Phimopollenites augathellaensis Dettman 1973

†Phimopollenites pannosus Dettmann 1974

†Phimopollenites pseudocheros Srivastava 1975

†Phimopollenites tectatus Singh 1983

G. †Pleurospermaepollenites Kulkova 1973

G. †Polycolpopollenites Nakoman 1967

G. †Polyporotetradites Salard-Cheboldaeff 1978

†Polyporotetradites laevigatus Salard-Cheboldaeff 1978

G. †Protrudodiporites van Hoeken-Klinkenberg 1966

†Protrudodiporites typicus van Hoeken-Klinkenberg 1966

G. †Psilabrevitricolpites van Hoeken-Klinkenberg 1966

†Psilabrevitricolpites flexibilis van Hoeken-Klinkenberg 1966

†Psilabrevitricolpites rotundus van Hoeken-Klinkenberg 1966

G. †Psilabrevitricolporites Van der Kaars 1983

†Psilabrevitricolporites simpliformis Van der Kaars 1983

†Psilabrevitricolporites triangularis Jaramillo and Dilcher 2001

G. †Psilamonocolpites Van der Hammen and Garcia de Mutis 1966

†Psilamonocolpites grandis Van der Hammen 1954

†Psilamonocolpites longicolpatus Mathur 1966

†Psilamonocolpites magnus Du Chene 1978

†Psilamonocolpites medius Van der Hammen 1954

G. †Psilastephanocolpites Leidelmeyer 1966

†Psilastephanocolpites adinos González-Guzmán 1967

†Psilastephanocolpites marginatus González-Guzmán 1967

†Psilastephanocolpites parvus Salard-Cheboldaeff 1978

†Psilastephanocolpites verrucosus González-Guzmán 1967

G. †Psilastephanocolporites Leidelmeyer 1966

†Psilastephanocolporites brevicolpatus Jaramillo and Dilcher 2001

†Psilastephanocolporites fissilis Leidelmeyer 1966

†Psilastephanocolporites globulus van Hoeken-Klinkenberg 1966

†Psilastephanocolporites grandis Salard-Cheboldaeff 1978

†Psilastephanocolporites laevigatus Salard-Cheboldaeff 1978

†Psilastephanocolporites minor Salard-Cheboldaeff 1978

†Psilastephanocolporites perforatus Salard-Cheboldaeff 1978

†Psilastephanocolporites punctatus Salard-Cheboldaeff 1978

†Psilastephanocolporites scabratus Salard-Cheboldaeff 1978

G. †Psilastephanoporites Regali et al. 1974

G. †Psilatricolpites Van der Hammen and Wijmstra 1964

†Psilatricolpites acerbus González-Guzmán 1967

†Psilatricolpites brevis González-Guzmán 1967

†Psilatricolpites colpiconstrictus van Hoeken-Klinkenberg 1966

†Psilatricolpites inargutus Archangelsky 1973

Invalid names: Tricolpites inargutus McIntyre 1968 [replaced]

†Psilatricolpites khargalensis Kedves 1999

†Psilatricolpites minutus González-Guzmán 1967

†Psilatricolpites palaeocenica Van der Hammen and Garcia de Mutis 1966

†Psilatricolpites pannosus Burger 1970

†Psilatricolpites patagonicus Freile 1972

†Psilatricolpites polaroides González-Guzmán 1967

†Psilatricolpites psilascabratus Archangelsky 1973

Invalid names: Tricolpites psilascabratus Norton 1969 [replaced]

†Psilatricolpites psilatus Potonié 1966

†Psilatricolpites pulcherrimum Freile 1972

†Psilatricolpites simplex Jaramillo and Dilcher 2001

†Psilatricolpites tetradus Brenner 1968

Invalid names: Psilatricolpites rubini Van der Hammen and Wymstra 1964 [nomen nudum]

G. †Ranunculacidites Sah 1967

†Ranunculacidites communis Sah 1967

†Ranunculacidites operculatus Jaramillo and Dilcher 2001

†Psilatricolporites operculatus medius González-Guzmán 1967

Invalid names: Psilatricolporites operculatus Van der Hammen and Wijmstra 1964 [synonym]

G. †Retibrevitricolpites van Hoeken-Klinkenberg 1966

†Retibrevitricolpites aegypticus Kedves 1999

†Retibrevitricolpites catatumbus González-Guzmán 1967

†Retibrevitricolpites distinctus van Hoeken-Klinkenberg 1966

†Retibrevitricolpites grandis Jaramillo and Dilcher 2001

†Retibrevitricolpites increatus González-Guzmán 1967

†Retibrevitricolpites retibolus Leidelmeyer 1966

†Retibrevitricolpites triangulatus van Hoeken-Klinkenberg 1966

G. †Retibrevitricolporites Legoux 1978

†Retibrevitricolporites grandis Jaramillo and Dilcher 2001

†Retibrevitricolporites obodoensis Legoux 1978

†Retibrevitricolporites speciosus Jaramillo and Dilcher 2001

G. †Retipollenites González-Guzmán 1967

†Retipollenites confusus González-Guzmán 1967

G. †Retistephanocolpites Leidelmeyer 1966

†Retistephanocolpites angeli Leidelmeyer 1966

†Retistephanocolpites finalis González-Guzmán 1967

†Retistephanocolpites fossulatus Jaramillo and Dilcher 2001

†Retistephanocolpites minutus González-Guzmán 1967

†Retistephanocolpites regularis van Hoeken-Klinkenberg 1966

†Retistephanocolpites williamsii Germeraad et al. 1968

G. †Retistephanoporites González-Guzmán 1967

†Retistephanoporites angelicus González-Guzmán 1967

†Retistephanoporites crassiexinatus Jaramillo and Dilcher 2001

†Retistephanoporites minutiporus Jaramillo and Dilcher 2001

G. †Retisyncolporites González-Guzmán 1967

†Retisyncolporites angularis González-Guzmán 1967

†Retisyncolporites aureus González-Guzmán 1967

G. †Retitrescolpites Sah 1967

†Retitrescolpites assamicus Dutta and Sah 1970

†Retitrescolpites baculatus Jaramillo and Dilcher 2001

†Retitrescolpites irregularis Jaramillo and Dilcher 2001

Invalid names: Retitricolporites irregularis Van der Hammen and Wijmstra 1964 [replaced]

†Retitrescolpites magnus Jaramillo and Dilcher 2001

Invalid names: Retitricolpites magnus González-Guzmán 1967 [replaced]

†Retitrescolpites minor Dutta and Sah 1970

†Retitrescolpites peculiaris Jaramillo and Dilcher 2001

†Retitrescolpites saturum Jaramillo and Dilcher 2001

Invalid names: Retitricolpites saturum González-Guzmán 1967 [replaced]

G. †Retitricolpites Van der Hammen 1956

†Retitricolpites absolutus Jaramillo and Dilcher 2001

†Retitricolpites adeptus González-Guzmán 1967

†Retitricolpites adultus González-Guzmán 1967

†Retitricolpites amapaensis Regali et al. 1974

†Retitricolpites americana Wymstra 1966

†Retitricolpites antonii González-Guzmán 1967

†Retitricolpites bonus González-Guzmán 1967

†Retitricolpites clarensis González-Guzmán 1967

†Retitricolpites conciliatus González-Guzmán 1967

†Retitricolpites constrictus González-Guzmán 1967

†Retitricolpites farafraensis Kedves 1999

†Retitricolpites fragosus Hedlund and Norris 1968

†Retitricolpites georgensis Brenner 1963

†Retitricolpites geranioides Benner 1963

†Retitricolpites herrerae Van der Hammen and Garcia de Mutis 1966

†Retitricolpites heteroreticulatus Boltenhagen 1976

†Retitricolpites incisus González-Guzmán 1967

†Retitricolpites jardinei Kedves 1999

†Retitricolpites magloirae Kedves 1999

†Retitricolpites maledictus González-Guzmán 1967

†Retitricolpites marginatus van Hoeken-Klinkenberg 1966

†Retitricolpites maturus González-Guzmán 1967

†Retitricolpites maximus Singh 1971

†Retitricolpites medireticulatus Mathur 1966

†Retitricolpites megareticulatus Mathur 1966

†Retitricolpites microreticulatus Van der Hammen 1954

†Retitricolpites minutus Kedves 1999

†Retitricolpites obtusus van Hoeken-Klinkenberg 1966

†Retitricolpites operculatus Herngreen 1973

†Retitricolpites perditus González-Guzmán 1967

†Retitricolpites polaris Gonzalez 1966

†Retitricolpites prosimilis Norris 1967

†Retitricolpites robustus Sah and Kar 1970

†Retitricolpites salardae Kedves 1999

†Retitricolpites simplex González-Guzmán 1967

†Retitricolpites sitholeyii Ramanujam 1966

†Retitricolpites sowunmiae Kedves 1999

†Retitricolpites variabilis Kedves 1999

†Retitricolpites vermimurus Brenner 1963

†Retitricolpites virgeus Brenner 1962

†Retitricolpites vulgaris Pierce 1961

G. †Scabratriporites Van der Hammen 1956

†Scabratriporites annellus van Hoeken-Klinkenberg 1964

†Scabratriporites moderatus González-Guzmán 1967

†Scabratriporites pororatus Mathur 1966

†Scabratriporites redundans González-Guzmán 1967

†Scabratriporites simpliformis van Hoeken-Klinkenberg 1966

G. †Schrankipollis Doyle et al. 1990

†Schrankipollis kachaikensis Barreda and Archangelsky 2006

†Schrankipollis mawhoubensis Doyle et al. 1990

†Schrankipollis reticulatus Regali and Santos 1996

G. †Senectotetradites Dettmann 1973

†Senectotetradites fistulosus Dettmann 1973

†Senectotetradites varireticulatus Dettmann 1973

G. †Stephanoporopollenites Thomson 1953

†Stephanoporopollenites obscurus Harris 1965

G. †Striatriporites van Hoeken-Klinkenberg 1966

†Striatriporites nigeriensis van Hoeken-Klinkenberg 1966

G. †Translucentipollis Chlonova 1961

†Translucentipollis plicatilis Chlonova 1961

G. †Tricolporoidites Pacltova 1971

†Tricolporoidites pacltovae Kedves 1999

†Tricolporoidites warringtonii Burger 1993

G. †Trifossapollenites Rouse 1957

†Trifossapollenites constatus Dutta and Sah 1970

†Trifossapollenites ivoirensis Jardiné and Magloire 1965

G. †Triporites Van der Hammen 1954

G. †Triporotetradites van Hoeken-Klinkenberg 1964

†Triporotetradites hoekenii Salard-Cheboldaeff 1978

†Triporotetradites letouzeyi Salard-Cheboldaeff 1978

†Triporotetradites scabratus van Hoeken-Klinkenberg 1964

G. †Verrustephanocolporites Salard-Cheboldaeff 1978

†Verrustephanocolporites complanatus Salard-Cheboldaeff 1978

G. †Verrutricolpites Pierce 1961

†Verrutricolpites irregularis González-Guzmán 1967

†Verrutricolpites perverrucatus Ramanujam 1966

†Verrutricolpites triangulus Sah and Kar 1970

†Verrutricolpites unicus González-Guzmán 1967

Ph. †Moresnetiophyta Doweld 2001

Cl. †Moresnetiopsida Doweld 2001

Or. †Pullarithecales Doweld 1998

Invalid names: Calathiopsidaceae Doweld 2001 [empty], Gnetopsidaceae Doweld 2001 [empty], Pullarithecaceae Doweld 1998 [empty]

Invalid names: Moresnetiales Doweld 2001 [empty]

Ph. †Noeggerathiophyta Doweld 2001

G. †Koilosphenus Bohlin 1971

†Koilosphenus cuneifolius Bohlin 1971

Ph. †Peltaspermophyta Doweld 2001

Cl. †Arberiopsida Doweld 2001

Or. †Arberiales Meyen 1982

Invalid names: Arberiaceae Rigby 1972 [empty], Glossopteridales Melville 1960 [replaced]

Or. †Vojnovskyales Neuburg 1965

Fm. †Vojnovskyaceae Neuburg 1963

G. †Kuznetskia Gorelova and Meyen 1982

G. †Scirostrobus Doweld and Naugolnykh 2002

Invalid names: Pholidophyllum Zalessky 1937 [replaced]

G. †Sylvella Zalessky 1937

G. †Vojnovskya Neuburg 1955

Invalid names: Rufloriaceae Meyen 1982 [synonym]

Cl. †Dictyopteridiopsida Doweld 2001

Or. †Dictyopteridiales Doweld 2001

Fm. †Dictyopteridiaceae Rigby 1978

Invalid names: Glossopteridaceae Trapl 1926 [synonym]

Invalid names: Breyteniaceae Doweld 2001 [empty]

Or. †Lidgettoniales Doweld 2001

Invalid names: Denkaniaceae Doweld 2001 [empty], Parthaceae Doweld 2001 [empty]

Invalid names: Glossopteridopsida Plumstead 1956 [synonym], Rigbyales Doweld 2001 [empty]

G. †Hermanophyton Arnold 1962

†Hermanophyton kirkbyorum Arnold 1962

G. †Lanceolatus Plumstead 1952

†Lanceolatus derouxides Plumstead 1952

Cl. †Peltaspermopsida Doweld 2001

G. †Aneimites Dawson 1860

†Aneimites acadica Dawson 1860

†Aneimites obtusilobus Naumann 1849

Or. †Peltaspermales Delevoryas 1979

Fm. †Cardiolepidaceae Meyen 1977

G. †Nucicarpus Neuburg 1965

G. †Phylladoderma Zalessky 1914

Fm. †Leuthardtiaceae Doweld 2001

G. †Arberophyllum Doweld 2000

Invalid names: Glossophyllum Kraeusel 1943 [replaced]

Fm. †Matatiellaceae

G. †Matatiella Anderson and Anderson 2003

G. †Townrovia Retallack 1981

Fm. †Peltaspermaceae Harris 1937

G. †Aipteris Zalessky 1934

G. †Angaridium Zalessky 1950

G. †Angaropteridium Zalessky 1932

G. †Antevsia Harris 1937

G. †Anthicocladus Zalessky 1937

G. †Autunia Krasser 1921

G. †Callipterianthus Roselt 1962

G. †Callipteris Brongniart 1849

G. †Edyndella Mogutcheva 1973

G. †Hissarella Sixtel 1966

G. †Kirjamkenia Prynada 1970

G. †Laskovskia Zalessky 1939

G. †Lepidopteris Schimper 1869

G. †Madygenia Sixtel 1956

G. †Madygenopteris Sixtel 1956

G. †Odontopteris Brongniart 1825

G. †Pachydermophyllum Thomas and Bose 1955

G. †Peltaspermum Harris 1937

G. †Pseudoodontopteris Grand'Eury 1877

G. †Pterispermostrobus Stopes 1914

G. †Reinitsia Walkom 1932

G. †Scytophyllum Bornemann 1856

G. †Stiphorus Meyen [1982]

G. †Tatarina Meyen 1969

G. †Tersiella Radczenko 1960

G. †Uralophyllum Kryshtofovich and Prynada 1933

G. †Vittaephyllum Dobruskina 1975

G. †Permotheca Zalessky 1929

†Permotheca sardykensis Zalessky 1929

G. †Pseudocallipteris Grand'Eury 1877

†Pseudocallipteris discreta Weiss 1870

G. †Syniopteris Zalessky 1929

†Syniopteris nesterenkoi Zalessky 1929

Fm. †Umkomasiaceae Petriella 1981

G. †Harrisiothecium Lundblad 1961

Invalid names: Harrisia Lundblad 1950 [replaced]

G. †Nidia Bose and Srivastava 1973

G. †Nidistrobus Bose and Srivastava 1973

G. †Pachypteris Brongniart 1828

G. †Pteroma Harris 1964

G. †Ptilozamites Nathorst 1878

G. †Rhaphidopteris Barale 1972

Invalid names: Stenopteris Saporta 1872 [replaced]

G. †Rhexoxylon Bancroft 1913

Fm. †Vetlugospermaceae Naugolnykh 2012

G. †Navipelta Karasev 2009

G. †Vetlugospermum Naugolnykh 2012

G. †Xenopteris Weiss 1870

†Xenopteris brardi Brongniart 1828

Invalid names: Autuniaceae Doweld 2001 [empty]

G. †Psaliangium Remy 1953

†Psaliangium odontopteroides Remy 1953

Or. †Sporophyllitales Doweld 2001

Invalid names: Sporophyllitaceae Doweld 2001 [empty]

Or. †Trichopityales Doweld 2001

Fm. †Trichopityaceae

G. †Biarmopteris Zalessky 1937

G. †Diceratosperma Andrews 1941

G. †Dichophyllum Elias 1936

G. †Mauerites Zalessky 1933

G. †Trichopitys Saporta 1875

Or. †Umkomasiales Doweld 2001

Fm. †Angaropeltidaceae Doweld 2001

G. †Angaropeltis Doweld 2001

Invalid names: Cardiolepis Neuburg 1965 [replaced]

G. †Reticulopteris Gothan 1941

†Reticulopteris muensteri Eichwald 1840

Invalid names: Pentoxylopsida Pant 1959 [empty]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

No ecological data are available

Age range

Maximum range based only on fossils: base of the Early/Lower Givetian to the top of the Late/Upper Pleistocene or 388.10000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 387.7 Ma

Collections (2689 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Eifelian	393.3 - 387.7	USA (New York)	Aneurophytales indet. (28538)
Middle Devonian	393.3 - 382.7	USA (New York)	Aneurophyton hallii (14214)
Middle Devonian	393.3 - 382.7	Germany (Wuppertals)	Aneurophyton germanicum (32209)
Early/Lower Givetian	388.1 - 383.7	Czech Republic	Aneurophyton bohemicum (23727 23739) Aneurophyton germanicum (23743)
Early/Lower Givetian	388.1 - 383.7	Czech Republic (Bohemia)	Aneurophyton bohemicum (27157 27159)
Givetian	387.7 - 382.7	USA (New York)	Aneurophytales indet. (20878 28048) Aneurophyton hallii (14246) Aneurophyton sp. (28414 28424) Archaeopteris obtusa, Aneurophyton sp. (14247)
Givetian	387.7 - 382.7	Germany	Aneurophyton germanicum (22759 22761 23660 23703)
Givetian	387.7 - 382.7	Germany (Elberfeld)	Aneurophyton bohemicum (23701)
Givetian - Late/Upper Devonian	387.7 - 358.9	Belgium	Aneurophyton germanicum (23745)
Early/Lower Frasnian	383.7 - 382.4	USA (New York)	Archaeopteris macilenta (20879) Archaeopteris sp. (28179 28287 28392)
Early/Lower Frasnian	383.7 - 382.4	Canada (Quebec)	Archaeopteris halliana, Archaeopteris obtusa (26084)
Early/Lower Frasnian	383.7 - 382.4	Canada (Alberta)	Archaeopteris macilenta, Archaeopteris sp. (21241)
Middle Givetian	383.0 - 380.2	Belgium (Brabant)	Aneurophyton furcatum (32214)
Frasnian	382.7 - 372.2	USA (New York)	Aneimites indet. (28409) Aneurophytales indet. (28206 28431 28551 29136) Aneurophyton germanicum (20730) Aneurophyton hallii (28967) Aneurophyton rachides (28928) Aneurophyton sp. (28957) Archaeopteris macilenta (14269 14297 28946 28947 29006) Archaeopteris macilenta, Archaeopteris halliana, Archaeopteris obtusa, Archaeopteris sphenophyllifolia, Archaeopteris hibernica (14274) Archaeopteris macilenta, Archaeopteris halliana, Archaeopteris obtusa, Archaeopteris sphenophyllifolia, Archaeopteris sp., Archaeopteris hibernica (14270) Archaeopteris macilenta, Archaeopteris sp. (28916) Archaeopteris macilenta, Archaeopteris sphenophyllifolia, Archaeopteris obtusa, Archaeopteris hibernica (14277) Archaeopteris sp. (14278 28192 28388 28412 28499 28516 28518 28521 28532 28537 28750 28767 28840 28841 28917 28951 29244 29246 29249) Archaeopteris sp., Aneimites indet. (28399) Archaeopteris sphenophyllifolia (14240)
Frasnian	382.7 - 372.2	Russian Federation (Arkhangel'skaya Oblast')	Archaeopteris roemeriana (40713)
Frasnian	382.7 - 372.2	USA (Pennsylvania)	Archaeopteris halliana, Archaeopteris macilenta (14230)
Frasnian	382.7 - 372.2	Canada (Quebec)	Archaeopteris halliana, Archaeopteris sp. (179176)
Late/Upper Devonian	382.7 - 358.9	USA (Pennsylvania)	Archaeopteris halliana, Archaeopteris macilenta, Archaeopteris minor (14235) Archaeopteris latifolia (14221) Archaeopteris rogersi, Archaeopteris halliana (14234) Archaeopteris sp. (31175)
Late/Upper Devonian	382.7 - 358.9	USA (New York)	Archaeopteris halliana (14236) Archaeopteris sp. (14226)
Late/Upper Devonian	382.7 - 358.9	Canada (Quebec)	Archaeopteris jacksoni (21131) Archaeopteris obtusa, Archaeopteris jacksoni, Archaeopteris gaspiensis (14218) Archaeopteris sp. (24551) Archaeopteris sp., Archaeopteris gaspiensis, Archaeopteris jacksoni (21231)