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Araucaria hastiensis

Equisetopsida - Araucariaceae

Taxonomy
Araucaria hastiensis was named by Hill and Bigwood (1987). Its type specimen is H-014, stored in the Queen Victoria Museum, Launceston, Tasmania., a leaf (Cuticle), and it is a compression fossil. Its type locality is Hasties, which is in an Eocene/Eocene fluvial mudstone/gravel in Australia.

Sister species lacking formal opinion data

Synonymy list
YearName and author
1987Araucaria hastiensis Hill and Bigwood p. 328 figs. 3A, C-F
1990Araucaria hastiensis Hill p. 218 figs. 36-41
1992Araucaria hastiensis Pole

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RankNameAuthor
kingdomPlantae
superphylumTracheophyta
Pteridobiotina
superphylumPteridophytanae
phylumEquisetophytaScott 1900
RankNameAuthor
classEquisetopsidaAgardh 1825
subclassPinidae
familyAraucariaceaeHenckel and Hochstetter 1865
genusAraucariade Jussieu 1789
specieshastiensis

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Araucaria hastiensis Hill and Bigwood 1987
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Diagnosis
ReferenceDiagnosis
R. S. Hill and A. J. Bigwood 1987Leaves bilateral, triangular, with margins slightly convex. Leaf length 19 mm, width 9 mm. Leaf base almost as broad as the widest point, no petiole present, apex acute. Margins entire, not appreciably thickened. Veins numerous, unbranching, parallel and inconspicuous. Leaf amphistomatous. Stomatal arrangement predominantly parallel to the long axis of the leaf, in sporadic long rows, one stomate wide between veins. Stomata less frequent on upper surface of leaf. Florin rings present but not conspicuous. Stomatal apparatus 55 µm long (36-74 µm), 34 µm wide (24-41 µm). Mean number of subsidiary cells 4.5 (4-6). Three levels of cuticular thickening present over stomates outer surface. Cuticle on subsidiary cells depressed, surface granular. Flange of cuticle between guard cells and subsidiary cells thickened, moderately irregular in outline. A ridge of cuticular tissue occurs in the aperture between guard cells, not extended at poles.
R. S. Hill 1990Leaf about 19 mm long, 9 mm wide, arrangement on axis unknown. Numerous parallel veins present. Leaf amphistomatous. Stomata on adaxial surface usually oriented parallel to leaf axis. Stomatal plugs with smooth surface, probably spongy interior. Guard cells level with hypodermis, subsidiary cells usually four, sometimes five or six. Cuticle on subsidiary cells very thick, granular, outer flange rim uniformly thickened, oval. Flange between guard and subsidiary cells well developed, irregular. Small polar extensions sometimes present. Flange between guard cells thick, smooth or granular, polar extensions present. Cuticle on guard cells granular, grooved for almost entire length on most stomata. Epidermal cell walls not sinuous, buttressing absent. Cuticle very thick and partially enclosing inner epidermal cell wall surface.