Basic info Taxonomic history Classification Included Taxa
Morphology Ecology and taphonomy External Literature Search Age range and collections

Daphoenodon (Borocyon) robustum

Mammalia - Carnivora - Amphicyonidae

Taxonomy
Borocyon robustum was named by Peterson (1910) [genotype]. Its type locality is Whistle Creek (Upper), which is in a Harrisonian terrestrial horizon in the Marsland Formation of Nebraska.

It was recombined as Daphoenodon robustum by Hunt (1971), Woodburne et al. (1974) and Frailey (1979); it was considered a nomen dubium by Matthew (1918); it was recombined as Daphoenodon (Borocyon) robustum by Hunt (2009).

Synonymy list
YearName and author
1910Borocyon robustum Peterson
1971Daphoenodon robustum Hunt
1974Daphoenodon robustum Woodburne et al.
1979Daphoenodon robustum Frailey
2009Daphoenodon (Borocyon) robustum Hunt, Jr. p. 14 figs. 3–8, 15, 21, 22, 24–28, 31, 32, 37–39

Is something missing? Join the Paleobiology Database and enter the data

RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
Mammaliaformes
classMammalia
RankNameAuthor
Theriamorpha(Rowe 1993)
Theriiformes()
Trechnotheria
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Ferae()
CarnivoramorphaWyss and Flynn 1993
CarnivoraformesFlynn et al.
orderCarnivora
suborderCaniformiaKretzoi 1943
familyAmphicyonidaeHaeckel 1886
subfamilyDaphoeninae
genusDaphoenodon
subgenusBorocyon()
speciesrobustum()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

show all | hide all
Diagnosis
ReferenceDiagnosis
R. M. Hunt 2009Largest known daphoenine amphicyonid, and the largest species of Borocyon, with basilar skull length of 285– 294 mm, differing from B. niobrarensis by larger average body size and dentition (c–m2 length, ,110–120 mm), consistent loss of M3, more elongate forelimbs, and greater modification of proximal tarsals (figs. 33, 34, astragalus-calcaneum) for fore–aft motion (see text for detailed discussion). Distin- guished from B. neomexicanus by much larger body size, dentition (figs. 17, 18; tables 2, 3), and more elongate yet anatom- ically similar distal forelimbs.
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: carnivoref
Reproduction: viviparoussubc
Created: 2005-08-30 14:52:56
Modified: 2005-08-30 16:52:56
Source: f = family, subc = subclass, c = class, subp = subphylum
References: Lillegraven 1979, Hendy et al. 2009, Carroll 1988, Ji et al. 2002, Hunt 1998

Age range: base of the Early/Lower Hemingfordian to the top of the Harrisonian or 20.43000 to 18.50000 Ma

Collections (12 total)


Time interval Ma Country or state Original ID and collection number
Harrisonian24.8 - 18.5USA (Oregon) Daphoeninae indet. (19037)
Harrisonian24.8 - 18.5USA (Nebraska) Borocyon robustum (type locality: 17858)
Early/Lower Hemingfordian20.43 - 15.97USA (New Mexico) Daphoenodon robustum (19188)
Early/Lower Hemingfordian20.43 - 15.97USA (Oregon) Daphoenodon robustum (167866)
Early/Lower Hemingfordian20.43 - 15.97USA (Nebraska) Daphoenodon robustum (17881 17944) Daphoenodon sp. (17898 17933 17934 17942 17945 17963)