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Rukwatitan bisepultus

Reptilia

Taxonomy
Rukwatitan bisepultus was named by Gorscak et al. (2014). Its type specimen is (RRBP) 07409, a partial skeleton (partial skeleton including three posterior cervical vertebrae, one partial anterior dorsal vertebral neural arch, three anterior caudal vertebrae, six middle ca), and it is a 3D body fossil. Its type locality is Namba 2, which is in a Cenomanian/Campanian fluvial sandstone in the Galula Formation of Tanzania. It is the type species of Rukwatitan.

Synonymy list
YearName and author
2014Rukwatitan bisepultus Gorscak et al. p. 1134 figs. 2-12
2017Rukwatitan bisepultus Gorscak et al. p. 2

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
RankNameAuthor
Diapsida()
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Saurischia()
Sauropoda()
Gravisauria
Eusauropoda
Neosauropoda
Macronaria
Titanosauriformes
Titanosauria
genusRukwatitan
speciesbisepultus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Rukwatitan bisepultus Gorscak et al. 2014
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Diagnosis
ReferenceDiagnosis
E. Gorscak et al. 2014Titanosaurian sauropod diagnosed by the following
suite of autapomorphies: ventral surface of posterior cervical
vertebrae with (1) weakly developed carotid processes posteriorly
and (2) paired shallow fossae anteriorly that are not separated
by a thin keel; (3) posterior cervical vertebrae with
anteroposteriorly elongate and undivided fossa posterior to the
anterior centrodiapophyseal lamina that deeply invades the ventral
surface of the diapophysis; (4) accessory tubercle on the ventral
surface of the cervical rib capitulum, lateral to the
capituloparapophyseal suture; (5) deep coracobrachialis fossa on
proximal humerus bounded by strong anteriorly projecting deltopectoral
crest and strong anteromedial ridge, extending to
approximately the humeral midshaft; (6) flattened anterior,
medial, lateral, and posterior surfaces at the humeral midshaft,
resulting in a subquadrangular cross-section; and (7) pubic
peduncle on the ilium that is teardrop-shaped, with the
‘tail’ curving posteriorly, slightly longer transversely than
anteroposteriorly.
Additionally, the following combination of phylogenetically
informative characters assist with the placement of Rukwatitan
bisepultus as a basal titanosaurian: undivided pleurocoels on cervical
vertebrae; strongly procoelous anterior caudal vertebrae,
with the apex of the posterior condyle positioned on the dorsal
half of the centrum; pedicles of the neural arch attached to the
anterior half of caudal vertebral centrum; unforked chevrons
with a deep hemal canal; and squared proximolateral corner of
the humerus. Notably, Rukwatitan lacks the ventral longitudinal
hollow on the caudal vertebral series typical of many other
titanosaurians.
Measurements
No measurements are available
Composition: hydroxyapatiteinfrao
Entire body: yesinfrao
Adult length: 10 to < 100infrao
Adult width: 10 to < 100infrao
Adult height: 10 to < 100infrao
Thickness: thickinfrao
Architecture: compact or denseinfrao
Form: sphericalinfrao
Ontogeny: accretion, modification of partsinfrao
Grouping: gregariousinfrao
Environment: terrestrialinfrao
Locomotion: actively mobileinfrao
Life habit: ground dwellinginfrao
Diet: herbivoreinfrao
Reproduction: oviparousinfrao
Dispersal: direct/internalinfrao
Dispersal 2: mobileinfrao
Created: 2004-12-14 12:49:10
Modified: 2004-12-14 14:49:10
Source: infrao = infraorder
Reference: Marsh 1875

Age range: base of the Cenomanian to the top of the Campanian or 100.50000 to 72.10000 Ma

Collections: one only


Time interval Ma Country or state Original ID and collection number
Cenomanian - Campanian100.5 - 72.1Tanzania (Mbeya) Rukwatitan bisepultus (type locality: 156028)