Basic info Taxonomic history Classification Included Taxa
Morphology Ecology and taphonomy External Literature Search Age range and collections

Sclerophorata

Taxonomy
Sclerophorata was named by Kamenz et al. (2011). It is extant.

It was assigned to Dekatriata by Lamsdell (2013).

Subtaxa
Arachnida

Synonymy list
YearName and author
2011Sclerophorata Kamenz et al.
2013Sclerophorata Lamsdell p. 20

Is something missing? Join the Paleobiology Database and enter the data

RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—EcdysozoaAguinaldo et al. 1997
—Panarthropoda
phylumArthropodaGravenhorst 1843
—Deuteropoda
RankNameAuthor
subphylumChelicerataHeymons 1901
—EuchelicerataWeygoldt and Paulus 1979
—Prosomapoda
—Planaterga
—Dekatriata
—SclerophorataKamenz et al. 2011
—SclerophorataKamenz et al. 2011

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Sclerophorata Kamenz et al. 2011
show all | hide all
Cl. Arachnida Lamarck 1801
hide
G. Aulonia Koch 1847
Subcl. Caulogastra Gall 1971
+
Invalid names: Labellata Petrunkevitch 1949 [empty]
G. †Cretopilio Fritsch 1910
+
†Cretopilio granulatus Fritsch 1910
G. †Dinopilio Fritsch 1904
+
†Dinopilio gigas Fritsch 1904
Unr. Haplocnemata Borner 1904
+
Or. Pseudoscorpiones Latreille 1817 [pseudoscorpion]
+
G. Heterolophus Tömösváry 1884
Subor. Heterosphyronida Chamberlin 1929
+
Superfm. Chthonioidea Daday 1889
Unr. Homosphyronida Chamberlin 1929
+
Subor. Atoposphyronida Harvey 2019
Subor. Iocheirata Harvey 1992
G. Obisium Leach 1814
Subor. †Palaeosphyronida Harvey 2019
+
Superfm. †Dracocheloidea Schawaller et al. 1991
Invalid names: Chelifer ehrenbergii Koch and Berendt 1854 [nomen dubium], Chelonethi Thorell 1883 [invalid subgroup], Epiochierata Harvey 1992 [invalid subgroup]
G. †Schneidarachne Dunlop and Rössler 2003
+
†Schneidarachne saganii Dunlop and Rössler 2003
Or. Palpigradi Thorell 1888 [microwhip scorpion]
+
Fm. Eukoeneniidae Petrunkevitch 1955
+
G. †Electrokoenenia Engel and Huang 2016
+
†Electrokoenenia yaksha Engel and Huang 2016
G. †Paleokoenenia Rowland and Sissom 1980
+
†Paleokoenenia mordax Rowland and Sissom 1980
Unr. Pantetrapulmonata Shultz 2007
+
G. †Douglassarachne Selden and Dunlop 2024
+
†Douglassarachne acanthopoda Selden and Dunlop 2024
G. †Ecchosis Selden and Shear 1991
+
†Ecchosis pulchribothrium Selden and Shear 1991
Subcl. Tetrapulmonata Schultz 1990
+
G. †Eocteniza Pocock 1911
+
†Eocteniza silvicola Pocock 1911
G. †Protocteniza Petrunkevitch 1949
+
†Protocteniza britannica Petrunkevitch 1949
G. †Rakovnicia Kušta 1885
+
†Rakovnicia antiqua Kušta 1885
Unr. Schizotarsata Shultz 2007
+
Or. †Haptopoda Pocock 1911
Unr. Pedipalpi Borner 1904
Unr. Serikodiastida Garwood and Dunlop 2014
+
Or. Araneae Clerck 1757 [spider]
Or. Araneida Clerck 1757
G. †Idmonarachne Garwood et al. 2016
Or. †Uraraneida Selden and Shear 2008 [spider]
G. †Xenarachne Dunlop and Poschmann 1997
+
†Xenarachne willwerathensis Dunlop and Poschmann 1997
Or. †Trigonotarbida Petrunkevitch 1949
+
G. †Aenigmatarbus Poschmann et al. 2016
+
†Aenigmatarbus rasteli Poschmann et al. 2016
Fm. †Anthracomartidae Haase 1890
+
G. †Anthracomartus Karsch 1882
+
Invalid names: Brachylycosa Fritsch 1904 [synonym], Cleptomartus Petrunkevitch 1949 [synonym], Coryphomartus Petrunkevitch 1945 [synonym], Cryptomartus Petrunkevitch 1945 [synonym], Oomartus Petrunkevitch 1953 [synonym], Perneria Fritsch 1904 [synonym], Pleomartus Petrunkevitch 1945 [synonym], Promygale Fritsch 1899 [synonym]
G. †Brachypyge Woodward 1878
G. †Maiocercus Pocock 1911
Invalid names: Brachypygidae Pocock 1911 [synonym], Coryphomartidae Petrunkevitch 1945 [synonym], Eurymartus Matthew 1895 [nomen dubium], Eurymartus latus Matthew 1895 [nomen dubium], Pleomartidae Petrunkevitch 1945 [synonym], Promygalidae Fritsch 1904 [synonym]
G. †Anthracophrynus Andrée 1913
+
Fm. †Anthracosironidae Pocock 1903
+
G. †Anthracosiro Pocock 1903
G. †Arianrhoda Dunlop and Selden 2004
G. †Vratislavia Fritsch 1904
Fm. †Aphantomartidae Petrunkevitch 1945
+
G. †Alkenia Størmer 1970
G. †Aphantomartus Pocock 1911
+
Invalid names: Phrynomartus Petrunkevitch 1945 [synonym], Trigonomartus Petrunkevitch 1913 [synonym]
G. †Doubravatarbus Hradská et al. 2022
Invalid names: Trigonomartidae Petrunkevitch 1945 [synonym]
Fm. †Archaeomartidae Poschmann and Dunlop 2010
+
G. †Archaeomartus Størmer 1970
G. †Areomartus Petrunkevitch 1913
+
†Areomartus ovatus Petrunkevitch 1913
Fm. †Eophrynidae Karsch 1882
+
G. †Eophrynus Woodward 1871
G. †Nyranytarbus Harvey and Selden 1995
+
Invalid names: Hemiphrynus Fritsch 1899 [replaced]
G. †Petrovicia Fritsch 1904
G. †Planomartus Petrunkevitch 1953
G. †Pleophrynus Petrunkevitch 1945
+
Invalid names: Acrokreischeria Petrunkevitch 1953 [synonym]
G. †Pocononia Petrunkevitch 1953
G. †Somaspidion Jux 1982
G. †Stenotrogulus Fritsch 1904
+
Invalid names: Cyclotrogulus Fritsch 1904 [synonym], Pseudoeophrynus Pribyl 1958 [synonym]
Invalid names: Hemiphrynidae Fritsch 1904 [synonym], Pleophrynidae Petrunkevitch 1945 [synonym]
Fm. †Kreischeriidae Haase 1890
+
G. †Anzinia Petrunkevitch 1953
G. †Gondwanarachne Pinto and Hunicken 1980
G. †Hemikreischeria Fritsch 1904
G. †Kreischeria Geinitz 1882
G. †Pseudokreischeria Petrunkevitch 1953
Fm. †Lissomartidae Dunlop 1995
+
G. †Lissomartus Petrunkevitch 1949
G. †Namurotarbus Poschmann and Dunlop 2010
+
†Namurotarbus roessleri Dunlop and Brauckmann 2006
Fm. †Palaeocharinidae Hirst 1923
+
G. †Aculeatarbus Shear et al. 1987
G. †Gelasinotarbus Shear et al. 1987
G. †Gigantocharinus Shear 2000
G. †Gilboarachne Shear et al. 1987
G. †Palaeocharinus Hirst 1923
+
Invalid names: Palaeocharinoides Hirst 1923 [synonym]
G. †Spinocharinus Poschmann and Dunlop 2011
G. †Palaeocteniza Hirst 1923
+
G. †Palaeotarbus Dunlop 1999
+
†Palaeotarbus jerami Dunlop 1996
Invalid names: Eotarbus Dunlop 1996 [replaced]
G. †Permotarbus Dunlop and Rößler 2013
+
†Permotarbus schuberti Dunlop and Rößler 2013
Fm. †Trigonotarbidae Petrunkevitch 1949
+
G. †Trigonotarbus Pocock 1911
G. †Tynecotarbus Hradská and Dunlop 2013
+
†Tynecotarbus tichaveki Hradská and Dunlop 2013
Invalid names: Anthracomarti Karsch 1882 [replaced], Anthracomartus hageni Goldenberg 1873 [nomen dubium], Elaverimartus Petrunkevitch 1953 [nomen dubium], Elaverimartus pococki Petrunkevitch 1953 [nomen dubium], Eophrynus scharfi Scharf 1924 [nomen dubium], Eurymarti Matthew 1895 [synonym], Eurymartidae Matthew 1895 [invalid subgroup], Eurymartus spinulosus Matthew 1895 [nomen dubium], Termes buchi Goldenberg 1873 [nomen dubium], Trigonomartus woodruffi Scudder 1893 [nomen dubium]
Or. Parasitiformes Reuter 1909
+
Subor. Ixodida Leach 1815 [tick]
+
Fm. †Deinocrotonidae Peñalver et al. 2017
+
G. †Deinocroton Peñalver et al. 2017
Superfm. Ixodoidea Banks 1907
+
Fm. Argasidae Koch 1844 [soft tick]
Fm. Ixodidae Banks 1907 [hard tick]
Subor. Mesostigmata Canestrini 1891
+
Infraor. Monogynaspida Camin and Gorirossi 1955
+
Unr. Gamasina Kramer 1881
Unr. Microgyniina Tragardh 1942
Unr. Uropodina Kramer 1881 [tortoise mite]
Infraor. Sejida Kramer 1885
+
Superfm. Sejoidea Berlese 1885
Infraor. Trigynaspida Camin and Gorirossi 1955
+
Unr. Antennophorina Berlese 1882
Invalid names: Gamasida Leach 1815 [synonym]
Subor. Opilioacarida Zachvatkin 1952
+
Superfm. Opilioacaroidea Vitzthum 1931
+
Fm. Opilioacaridae Vitzthum 1932
Or. †Phalangiotarbida Haase 1890 [phalangiotarbid]
+
Fm. †Anthracotarbidae Kjellesvig-Waering 1969
+
G. †Anthracotarbus Kjellesvig-Waering 1969
+
†Anthracotarbus hintoni Kjellesvig-Waering 1969
Fm. †Architarbidae Karsch 1882
+
G. †Architarbus Scudder 1868
+
†Architarbus hoffmanni Guthörl 1934
+
Invalid names: Goniotarbus sarana Guthorl 1965 [objective synonym], Opiliotarbus kliveri Waterlot 1934 [objective synonym]
†Architarbus minor Petrunkevitch 1913
†Architarbus rotundatus Scudder 1868
+
Invalid names: Geraphrynus carbonarius Scudder 1884 [synonym]
Invalid names: Geraphrynus Scudder 1884 [synonym]
G. †Bornatarbus Rossler and Schneider 1997
+
†Bornatarbus mayasii Haupt 1955
G. †Discotarbus Petrunkevitch 1913
+
†Discotarbus deplanatus Petrunkevitch 1913
G. †Geratarbus Scudder 1890
+
†Geratarbus bohemicus Petrunkevitch 1953
†Geratarbus lacoei Scudder 1890
G. †Goniotarbus Petrunkevitch 1949
+
†Goniotarbus angulatus Pocock 1911
†Goniotarbus tuberculatus Pocock 1911
+
Invalid names: Goniotarbus tuberculatus Petrunkevitch 1949 [synonym]
G. †Hadrachne Melander 1903
+
†Hadrachne horribilis Melander 1903
G. †Leptotarbus Petrunkevitch 1945
+
†Leptotarbus torpedo Pocock 1911
G. †Mesotarbus Petrunkevitch 1949
+
†Mesotarbus angustus Pocock 1911
†Mesotarbus eggintoni Pocock 1911
†Mesotarbus hindi Pocock 1911
†Mesotarbus intermedius Petrunkevitch 1949
†Mesotarbus peteri Dunlop and Horrocks 1997
G. †Metatarbus Petrunkevitch 1913
+
†Metatarbus triangularis Petrunkevitch 1913
G. †Ootarbus Petrunkevitch 1945
+
†Ootarbus ovatus Petrunkevitch 1945
†Ootarbus pulcher Petrunkevitch 1945
G. †Orthotarbus Petrunkevitch 1945
+
†Orthotarbus longipes Simon 1971
†Orthotarbus minutus Petrunkevitch 1913
†Orthotarbus nyranensis Petrunkevitch 1953
†Orthotarbus robustus Petrunkevitch 1945
G. †Paratarbus Petrunkevitch 1945
+
†Paratarbus carbonarius Petrunkevitch 1945
G. †Phalangiotarbus Haase 1890
+
†Phalangiotarbus subovalis Woodward 1872
G. †Pycnotarbus Daber 1990
+
†Pycnotarbus verrucosus Daber 1990
G. †Triangulotarbus Patrick 1989
+
Invalid names: Phalangiotarbidae Haase 1890 [synonym]
Fm. †Devonotarbidae Poschmann and Dunlop 2012
+
G. †Devonotarbus Poschmann et al. 2005
+
†Devonotarbus hombachensis Poschmann et al. 2005
Fm. †Heterotarbidae Petrunkevitch 1913
+
G. †Heterotarbus Petrunkevitch 1913
+
†Heterotarbus ovatus Petrunkevitch 1913
Fm. †Opiliotarbidae Petrunkevitch 1945
+
G. †Opiliotarbus Pocock 1911
+
†Opiliotarbus elongatus Scudder 1890
Invalid names: Architarbida Petrunkevitch 1945 [synonym], Eotarbus Kušta 1889 [nomen dubium], Eotarbus litoralis Kušta 1889 [nomen dubium], Protopilio depressus Petrunkevitch 1913 [nomen dubium]
Unr. Poecilophysidea Pepato et al. 2010
+
Or. Acariformes Zakhvatkin 1952 [mite]
+
Subor. Sarcoptiformes Reuter 1909
+
Infraor. Endeostigmata
Infraor. Oribatida Dugès 1834 [oribatid mite]
Subor. Trombidiformes Reuter 1909
+
Infraor. Prostigmata
Unr. Tetrapodili Oudemans 1923
Or. Solifugae Sundevall 1833 [camel spider]
+
Fm. Ammotrechidae Roewer 1934
+
G. †Happlodontus Poinar and Santiago-Blay 1989
Fm. Ceromidae Roewer 1933
+
G. †Cratosolpuga Selden 1996
G. †Cushingia Dunlop et al. 2015
+
†Cushingia ellenbergeri Dunlop et al. 2015
Fm. Daesiidae Kraepelin 1899
+
Subfm. Gnosippinae Roewer 1933
G. †Palaeoblossia Dunlop et al. 2004
G. Galeodes Olivier 1791
+
Subg. Galeodes (Galeodes) Olivier 1791
G. †Protosolpuga Petrunkevitch 1913
+
†Protosolpuga carbonaria Petrunkevitch 1913
Or. Ricinulei Thorell 1876 [hooded tickspider]
+
Subor. Posteriorricinulei Wunderlich 2015
+
Fm. †Curculioididae Cockerell 1916
+
G. †Amarixys Selden 1992
G. †Curculioides Buckland 1837
+
Invalid names: Hevila Giebel 1856 [objective synonym]
Fm. †Poliocheridae Scudder 1884
+
G. †Poliochera Scudder 1884
G. †Terpsicroton Selden 1992
Subor. †Primoricinulei Wunderlich 2015
+
Fm. †Hirsutisomidae Wunderlich 2017
+
G. †Hirsutisoma Wunderlich 2017
Fm. †Primoricinuleidae Wunderlich 2015
+
G. †Primoricinuleus Wunderlich 2015
Fm. †Sigillaricinuleidae Wunderlich 2022
+
G. †Sigillaricinuleus Wunderlich 2022
Invalid names: Palaeoricinulei Selden 1992 [empty]
G. †Saccogulus Dunlop et al. 2006
+
†Saccogulus seldeni Dunlop et al. 2006
Unr. Stomothecata Shultz 2007
+
Or. Opiliones Sundevall 1833 [harvestman]
+
G. †Arachnometa Petrunkevitch 1949
+
†Arachnometa tuberculata Petrunkevitch 1949
Subor. Cyphophthalmi Simon 1879
+
G. †Foveacorpus Bartel et al. 2023
Fm. Sironidae Simon 1879
Fm. Stylocellidae Hansen and Sorensen 1904
G. †Hasseltides Weyenburgh 1869
Subor. Laniatores Thorell 1876
+
Fm. Beloniscidae Kury et al. 2019
Infraor. Grassatores Kury 2002
Infraor. Insidiatores Loman 1900
Fm. †Mesokanidae Bartel et al. 2020
G. Philacarus Sorensen 1932
Fm. Pyramidopidae Sharma et al. 2011
Fm. Tithaeidae Sharma and Giribet 2011
Fm. †Monooculricinuleidae Wunderlich 2017
+
G. †Monooculricinuleus Wunderlich 2017
Unr. Phalangida Bristowe 1949
+
Fm. †Archaeometidae Petrunkevitch 1949
Subor. Dyspnoi Hansen and Sorensen 1904
Subor. Eupnoi Hansen and Sorensen 1904
Subor. †Tetrophthalmi Garwood et al. 2014
G. Phalangium Linnaeus 1758
G. Platybunus Koch 1839
Invalid names: Cheiromachus Menge 1854 [nomen dubium], Cheiromachus coriaceus Menge 1854 [nomen dubium], Phalangidae Roewer 1923 [empty], Phalangium succineum Presl 1822 [nomen dubium], Phalangodidae Simon 1879 [empty]
Or. Scorpiones Latreille 1817 [scorpion]
+
Subor. †Bilobosternina Kjellesvig-Waering 1986
+
Superfm. †Branchioscorpionoidea Kjellesvig-Waering 1986
G. †Brontoscorpio Kjellesvig-Waering 1972
+
†Brontoscorpio anglicus Kjellesvig-Waering 1972
G. †Eramoscorpius Waddington et al. 2015
+
†Eramoscorpius brucensis Waddington et al. 2015
G. †Gondwanascorpio Gess 2013
+
G. †Gymnoscorpius Jeram 1994
+
Subor. †Holosternina Kjellesvig-Waering 1986
+
Superfm. †Acanthoscorpionoidea Kjellesvig-Waering 1986
Superfm. †Eoctonoidea Kjellesvig-Waering 1986
Superfm. †Gigantoscorpionoidea Kjellesvig-Waering 1986
Superfm. †Mesophonoidea Wills 1910
Superfm. †Palaeoscorpoidea Lehmann 1944
Superfm. †Spongiophonoidea Kjellesvig-Waering 1986
Invalid names: Stoermeroscorpionoidea Kjellesvig-Waering 1986 [invalid subgroup]
G. †Hubeiscorpio Walossek et al. 1990
+
†Hubeiscorpio gracilitarsus Walossek et al. 1990
G. †Liassoscorpionides Bode 1951
+
Subor. †Lobosternina Pocock 1911
+
Superfm. †Isobuthoidea Petrunkevitch 1913
Superfm. †Loboarchaeoctonoidea Kjellesvig-Waering 1986
Superfm. †Paraisobuthoidea Kjellesvig-Waering 1986
Superfm. †Pseudobuthiscorpioidea Kjellesvig-Waering 1986
Invalid names: Anthracochaeriloidea Kjellesvig-Waering 1986 [empty], Palaeophonoidea Thorell and Lindstroem 1884 [empty]
Subor. †Meristosternina Kjellesvig-Waering 1986
+
Superfm. †Cyclophthalmoidea Thorell and Lindstroem 1885
Superfm. †Palaeobuthoidea Kjellesvig-Waering 1986
Subor. Orthosternina Pocock 1911
+
Unr. Buthida Soleglad and Fet 2003
Superfm. Chactoidea Pocock 1893
Superfm. Chaeriloidea Pocock 1893
G. †Corniops Jeram 1994
Unr. Pseudochactida Soleglad and Fet 2003
Superfm. Scorpionoidea Latreille 1802
G. †Suraju Martine et al. 2020
Invalid names: Chaerilida Soleglad and Fet 2003 [empty]
G. †Palaeomachus Pocock 1911
+
†Palaeomachus anglicus Woodward 1876
Fm. †Palaeotrilineatidae Lourenço 2012
+
G. †Palaeotrilineatus Lourenço 2012
G. †Permomatveevia Dammann 2017
+
†Permomatveevia perneri Dammann 2017
Fm. †Proscorpiidae Scudder 1885
+
G. †Archaeoctonus Pocock 1911
G. †Hydroscorpius Kjellesvig-Waering 1986
+
Invalid names: Archaeophonus Kjellesvig-Waering 1966 [synonym], Stoermeroscorpio Kjellesvig-Waering 1986 [synonym]
G. †Pseudoarchaeoctonus Kjellesvig-Waering 1986
G. †Waeringoscorpio Størmer 1970
Invalid names: Archaeoctonidae Petrunkevitch 1949 [synonym], Hydroscorpionidae Kjellesvig-Waering 1986 [synonym], Labriscorpionidae Kjellesvig-Waering 1986 [synonym], Stoermeroscorpioniidae Kjellesvig-Waering 1986 [synonym], Waeringoscorpionidae Størmer 1970 [synonym]
G. †Titanoscorpio Kjellesvig-Waering 1986
+
†Titanoscorpio douglassi Kjellesvig-Waering 1986
G. †Wattisonia Wills 1960
+
Invalid names: Allopalaeophonoidea Kjellesvig-Waering 1986 [invalid subgroup], Archaeoctonoidea Petrunkevitch 1949 [invalid subgroup], Branchioscorpionina Kjellesvig-Waering 1986 [invalid subgroup], Liassoscorpionidae Kjellesvig-Waering 1986 [invalid subgroup], Mesoscorpionina Stockwell 1989 [empty], Palaeophonus lightbodyi Kjellesvig-Waering 1954 [nomen dubium]
Invalid names: Acari Leach 1817 [invalid subgroup], Acaromorpha Dubinin 1957 [invalid subgroup], Arthrolycosa tarda Frič 1912 [nomen dubium], Podurites Matthew 1895 [nomen dubium], Podurites saltator Matthew 1895 [nomen dubium], Pseudoscorpionida Latreille 1817 [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Locomotion: actively mobilep
Created: 2004-02-29 09:38:49
Modified: 2004-02-29 11:38:49
Source: p = phylum
Reference: Kiessling 2004

Age range

Maximum range based only on fossils: base of the Telychian to the top of the Holocene or 438.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 432.9 Ma

Collections (729 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Telychian438.6 - 432.9United Kingdom (Scotland) Palaeophonus loudonensis (153581)
Telychian - Sheinwoodian438.6 - 430.6United Kingdom (South Lanarkshire) Palaeophonus caledonicus (153592)
Sheinwoodian432.9 - 430.6Sweden Palaeophonus nuncius (153548)
Wenlock432.9 - 426.7Canada (Ontario) Eramoscorpius brucensis (165189 165190)
Ludfordian425.0 - 422.7United Kingdom (Wales) Scorpiones indet. (153611)
Pridoli422.7 - 419.62United Kingdom (England) Trigonotarbida indet. (11128)
Pridoli422.7 - 419.62United Kingdom (Shropshire) Palaeophonus lightbodyi (153549)
Pridoli422.7 - 419.62USA (New York) Palaeophonus osborni (153533) Proscorpius osborni (491 153530 153531 238279) Proscorpius osborni, Archaeophonus eurypteroides (153532) Proscorpius osborni, Archaeophonus eurypteroides, Stoermeroscorpio delicatus (153529)
Lochkovian419.62 - 413.02United Kingdom (England) Praearcturus gigas (153544)
Lochkovian419.62 - 413.02USA (Wyoming) Hydroscorpius denisoni, Acanthoscorpio mucronatus, Praearcturus sp., Branchioscorpio richardsoni (153534)
Lochkovian419.62 - 413.02United Kingdom Arianrhoda bennetti (153478)
Lochkovian419.62 - 413.02United Kingdom (Worcestershire) Brontoscorpio anglicus (153545)
Lochkovian419.62 - 413.02United Kingdom (Herefordshire) Praearcturus gigas (201736)
Pragian413.02 - 410.62Germany Devonotarbus hombachensis (162728 162729) Devonotarbus hombachensis, Trigonotarbida indet. (152107) Waeringoscorpio westerwaldensis (155761) Waeringoscorpio westerwaldensis, Archaeomartus levis, Spinocharinus steinmeyeri (155760)
Pragian413.02 - 410.62United Kingdom (Scotland) Palaeocharinus kidstoni (10721) Palaeocharinus tuberculatus (153492) Palaeocteniza crassipes, Palaeocharinus sp., Protacarus crani (172778) Protacarus crani, Palaeotydeus devonicus, Paraprotacarus hirsti, Protospeleorchestes pseudoprotacarus, Palaeocharinus sp., Palaeocharinoides hornei, Palaeocharinus calmani, Palaeocharinus scourfieldi, Palaeocharinus rhyniensis (172777) Saccogulus seldeni, Eophalangium sheari, Palaeocharinus sp. (153498)
Early/Lower Emsian410.62 - 399.5Germany Alkenia mirabilis (155774) Devonotarbus hombachensis (162730) Devonotarbus hombachensis, Xenarachne willwerathensis (155775) Palaeoscorpius devonicus (26904) Trigonotarbus stoermeri (172343) Waeringoscorpio hefteri (155758 155759) Waeringoscorpio hefteri, Devonotarbus hombachensis, Archaeomartus levis, Alkenia mirabilis (155757)
Early/Lower Emsian410.62 - 399.5Germany (Alken) Archaeometa devonica, Waeringoscorpio hefteri, Archaeomartus tuberculatus, Alkenia mirabilis (27049)
Eifelian393.47 - 387.95USA (New York) Oribatida indet., Devonacarus sellnicki, Protochthonius gilboa (106001)
Early/Lower Givetian387.95 - 385.9USA (New York) Arachnida indet., Attercopus fimbriungus (28295) Gelasinotarbus bonamoae, Gelasinotarbus fimbriunguis, Gelasinotarbus reticulatus, Gilboarachne griersoni (153480)
Givetian387.95 - 382.31USA (New York) Arachnida indet. A, Arachnida indet. B, Archaeacarus dubinini, Dracochela deprehendor, Attercopus fimbriunguis, Ecchosis pulchribothrium, Gelasinotarbus bonamoae, Gelasinotarbus heptops, Gelasinotarbus bifidus, Gelasinotarbus reticulatus, Gilboarachne griersoni, Aculeatarbus depressus (28246) Arachnida indet. B, Gelasinotarbus bonamoae, Gelasinotarbus fimbriunguis, Gelasinotarbus reticulatus, Gilboarachne griersoni (21172)