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Eucnemesaurus entaxonis

Reptilia

Taxonomy
Eucnemesaurus entaxonis was named by McPhee et al. (2015). Its type specimen is BP/1/6234, a set of postcrania (Posteriormost dorsal vertebrae, sacrum, anterior caudal vertebrae, pelvis and hindlimb), and it is a 3D body fossil. Its type locality is Cannon Rock Farm, Aliwal North, which is in a Norian/Rhaetian floodplain mudstone in the Elliot Formation of South Africa.

Synonymy list
YearName and author
2015Eucnemesaurus entaxonis McPhee et al.
2016Eucnemesaurus entaxonis Peyre de Fabrègues and Allain p. 18
2021Eucnemesaurus entaxoni Müller and Garcia p. 4
2023Eucnemesaurus entaxonis Sciscio et al. p. 3

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
RankNameAuthor
Romeriida
Diapsida()
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Saurischia()
Eusaurischia
Sauropodomorpha(Huene 1932)
Massopoda
Riojasauridae
genusEucnemesaurus
speciesentaxonis

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Eucnemesaurus entaxonis McPhee et al. 2015
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Diagnosis
ReferenceDiagnosis
B. W. McPhee et al. 2015Two potential autapomorphies diagnose E. entaxonis: a deep brevis fossa with relatively thin lateral and medial walls on the ventral surface of the postacetabular process of the ilium (a pronounced brevis fossa is also present in the holotype of Riojasaurus, but this can be distinguished from E. entaxonis with respect to its expansive mediolateral width and considerably thicker medial and lateral margins); and a sharp ventral keel on the centra of the proximal caudal vertebrae (although this region is poorly preserved).
In addition to these features, E. entaxonis can be further diagnosed with respect to a unique suite of local autapomorphies (given its current position within our phylogeny): a small, circular pit (‘non-articulating gap’) that excavates the sacral rib of the first primordial sacral at midheight (present also in Melanorosaurus); femoral shaft beneath the fourth trochanter transversely elliptical in cross-section (present in taxa from Melanorosaurus crownwards); posterior descending process of the distal tibia does not extend as far laterally as the anterior ascending process, rendering the latter visible in posterior view (present in Aardonyx crownwards; convergently acquired in Anchisaurus); and a semi-stout pes in which the maximum proximal breadth of the first metatarsal is approximately 0.6 times its proximodistal length (present in several derived basal sauropodiform taxa).
E. entaxonis differs from E. fortis in the following features: (1) E. fortis presents a femur that is subcircular in cross-section rather than elliptical, and (2) the holotype of E. fortis (TM 119) displays a distal tibia in which the mediolaterally extensive posterior surface extends as far laterally as the anterior ascending process (for articulation with the astragalus). It is possible that the distal embayment present on the fourth trochanter in E. fortis is absent in E. entaxonis, but poor preservation in the latter taxon precludes confirmation of this. Furthermore, a dorsal neural arch of E. fortis possesses a unique accessory lamina within the centrodiapophyseal fossa—a feature that cannot be confirmed in the preserved material of E. entaxonis. This feature therefore remains a valid diagnostic character of E. fortis for the time being.