Basic info Taxonomic history Classification Included Taxa
Morphology Ecology and taphonomy External Literature Search Age range and collections

Dytiscidae (predaceous diving beetle)

Insecta - Coleoptera - Dytiscidae

Synonyms
Synonymy list
YearName and author
1856Necticus palustris Aymard p. 234
1861Dytiscidae LeConte p. 38
1981Dytiscidae Crowson p. 694
1992Dytiscidae Carpenter
1995Dytiscidae Lawrence and Newton, Jr.
2003Dytiscidae Wappler
2011Dytiscidae Bouchard et al. p. 162
2019Dytiscidae Yang et al. p. 2
2022Dytiscidae Cai et al.

Is something missing? Join the Paleobiology Database and enter the data

RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Protostomia
—Ecdysozoa
—Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
—Dicondylia
—Paranotalia
subclassPterygota()
RankNameAuthor
—NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
—Holometabola
—Aparaglossata
—Neuropteriformia
—Coleopterida
orderColeoptera
—Mesocoleoptera
—Metacoleoptera
suborderAdephaga
familyDytiscidae()
familyDytiscidae()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Dytiscidae Leach 1815 [predaceous diving beetle]
show all | hide all
Subfm. Agabinae Thomson 1867
+
Tr. Agabini Thomson 1867
+
G. Hydrotrupes Sharp 1882
+
†Hydrotrupes prometheus Gómez and Damgaard 2014
G. Platambus Thomson 1859
+
Platambus optatus Sharp 1884
Platambus stygius Régimbart 1899
G. Agabus Leach 1817
+
Subg. Agabus (Agabus) Leach 1817
+
†Agabus charon Wickham 1912
†Agabus congeneroides Lomnicki 1894
†Agabus corticeus Lomnicki 1894
†Agabus florissantensis Wickham 1913
†Agabus hochi Statz 1939
†Agabus latissimus Ponomarenko 2019
†Agabus lithax Říha 1960
†Agabus niedzwiedzkii Lomnicki 1894
†Agabus parvulus Lomnicki 1894
†Agabus perditus Scudder 1900
†Agabus praelugens Wickham 1917
†Agabus rathbuni Scudder 1900
†Agabus reductus von Heyden and von Heyden 1866
†Agabus reitteri Lomnicki 1894
†Agabus rottensis Nilsson 2001
+
Invalid names: Agabus latus Statz 1939 [replaced]
†Agabus savagei Wickham 1917
†Agabus ungeri Heer 1847
Invalid names: Necticus Hope 1838 [synonym]
G. Ilybius Erichson 1832
+
Ilybius apicalis Sharp 1873
†Ilybius boryslavicus Lomnicki 1894
Ilybius fenestratus Fabricius 1781
†Ilybius gratshevi Ponomarenko 2019
Ilybius subaeneus Erichson 1837
Ilybius vittiger Gyllenhal 1827
G. †Angaragabus Ponomarenko 1963
+
†Angaragabus jurassicus Ponomarenko 1963
Subfm. Colymbetinae Erichson 1837
+
Tr. Colymbetini Erichson 1837
+
G. Colymbetes Clairville 1806
+
†Colymbetes aemulus Heer 1862
Colymbetes fuscus Linnaeus 1758
†Colymbetes miocaenicus Říha 1974
Colymbetes paykulli Erichson 1837
Invalid names: Cymatopterus Dejean 1833 [synonym]
G. Rhantus Dejean 1833
+
Rhantus exsoletus Forster 1771
†Rhantus praesuturellus Lomnicki 1894
Subfm. Copelatinae Van den Branden 1885
+
G. Copelatus Erichson 1832
+
†Copelatus aphroditae Balke 2003
†Copelatus convexus Förster 1891
†Copelatus fossilis Říha 1974
†Copelatus ponomarenkoi Říha 1974
†Copelatus predaveterus Miller 2003
Invalid names: Copelatini Van den Branden 1885 [empty]
Subfm. Coptotominae Van den Branden 1885
+
Tr. Coptotomini Van den Branden 1885
+
G. Coptotomus Say 1830
+
†Coptotomus balticus Hendrich and Balke 2020
G. †Cretodytes Ponomarenko 1977
+
†Cretodytes incertus Prokin et al. 2013
†Cretodytes latipes Ponomarenko 1977
Subfm. Cybisterinae Sharp 1880
+
G. Cybister Curtis 1827
+
†Cybister agassizi Heer 1862
†Cybister atavus Heer 1862
†Cybister fractus Říha 1974
†Cybister imperfectus Říha 1974
†Cybister mancus Říha 1974
†Cybister rotundatus Říha 1974
G. Megadytes Sharp 1882
+
Subg. Megadytes (Paramegadytes) Trémouilles and Bachmann 1980
+
†Megadytes nicoleti Heer 1862
Subfm. Dytiscinae Leach 1815
+
Tr. Aciliini Thomson 1867
+
G. Acilius Leach 1817
+
†Acilius florissantensis Wickham 1909
†Acilius oeningensis Heer 1847
†Acilius praesulcatus Lomnicki 1894
G. Graphoderus Dejean 1833
+
†Graphoderus heeri Nilsson 2001
+
Invalid names: Dytiscus areolatus Heer 1862 [replaced]
†Graphoderus mirabilis Říha 1974
Tr. †Ambarticini Yang et al. 2019
+
G. †Ambarticus Yang et al. 2019
+
†Ambarticus myanmaricus Yang et al. 2019
Tr. Dytiscini Leach 1815
+
G. Dytiscus Linnaeus 1758
+
†Dytiscus avunculus von Heyden 1862
Dytiscus lapponicus Gyllenhal 1808
†Dytiscus latahensis Wickham 1931
†Dytiscus lavateri Heer 1847
Dytiscus marginalis Linnaeus 1758
†Dytiscus miocenicus Lewis and Gundersen 1987
†Dytiscus zersii Sordelli 1882
G. †Miodytiscus Wickham 1911
+
†Miodytiscus hirtipes Wickham 1911
Tr. Eretini Crotch 1873
+
G. Eretes Laporte 1833
+
†Eretes antiquus Oustalet 1870
Invalid names: Eunectes Erichson 1832 [replaced]
Tr. Hydaticini Sharp 1880
+
G. Hydaticus Leach 1817
+
Subg. Hydaticus (Hydaticus) Leach 1817
+
+
Invalid names: Hydaticus laevipennis Thomson 1867 [synonym]
Subfm. Hydroporinae Aubé 1836
+
Tr. Bidessini Sharp 1880
+
G. Bidessus Sharp 1882
Invalid names: Bidessus laminarum Wickham 1914 [nomen dubium], Hydroporus sedimentorum Wickham 1914 [nomen dubium]
G. †Glessaria Koch and Berendt 1854
+
†Glessaria rostrata Koch and Berendt 1854
Tr. Hydroporini Aubé 1836
+
Subtr. Deronectina Galewski 1994
+
G. Boreonectes Angus 2010
+
G. Deronectes Sharp 1882
G. Nectoporus Guignot 1950
+
Nectoporus sanmarkii Sahlberg 1826
G. Oreodytes Seidlitz 1887
+
Oreodytes alpinus Paykull 1798
Oreodytes borealis Gyllenhal 1826
G. Hydrocolus Roughley and Larson 2000
+
Hydrocolus oblitus Aubé 1838
G. Hydroporus Clairville 1806
+
†Hydroporus carstengroehni Balke et al. 2010
†Hydroporus clessini Lomnicki 1894
Hydroporus discretus Fairmaire and Brisout 1859
†Hydroporus edenianus Lesne 1926
†Hydroporus inanimatus Scudder 1900
†Hydroporus inundatus Scudder 1900
†Hydroporus lapideus Říha 1974
Hydroporus lapponum Gyllenhal 1808
Hydroporus morio Aubé 1838
Hydroporus notabilis LeConte 1850
+
Invalid names: Hydroporus arcticus Thomson 1856 [synonym]
Hydroporus palustris Linnaeus 1761
Hydroporus planus Fabricius 1781
†Hydroporus pleistocenicus Lomnicki 1894
†Hydroporus praedorsalis Lomnicki 1894
†Hydroporus praenigrita Lomnicki 1894
†Hydroporus praenivalis Lomnicki 1894
†Hydroporus sandbergeri Lomnicki 1894
†Hydroporus sectus Scudder 1900
†Hydroporus statzi Nilsson 2001
+
Invalid names: Hydroporus macularis Statz 1939 [replaced]
Hydroporus striola Gyllenhal 1826
†Hydroporus subarcticus Lomnicki 1894
†Hydroporus trimaculatus Theobald 1937
G. Nebrioporus Régimbart 1906
+
†Nebrioporus benzeli Heer 1862
Invalid names: Potamonectes Zimmerman 1921 [synonym]
G. †Pteroporus Guignot 1933
+
†Pteroporus antiquus Heer 1862
Tr. Hygrotini Portevin 1929
+
G. Clemnius Villastrigo et al. 2017
+
Subg. Clemnius (Clemnius) Villastrigo et al. 2017
+
G. Hygrotus Stevens 1828
+
Subg. Hygrotus (Coelambus) Thomson 1860
Subg. Hygrotus (Hygrotus) Stephens 1828
†Hygrotus cribrarius Scudder 1900
†Hygrotus derelictus Scudder 1900
†Hygrotus disjectus Scudder 1900
†Hygrotus dzieduszyckii Lomnicki 1894
†Hygrotus ganglbaueri Lomnicki 1894
†Hygrotus infernalis Scudder 1900
†Hygrotus latefasciatus Lomnicki 1894
†Hygrotus miocenus Wickham 1912
†Hygrotus niedzwiedzkii Lomnicki 1894
†Hygrotus ozokeriticus Lomnicki 1894
†Hygrotus picipoides Lomnicki 1894
Tr. Hyphydrini Gistel 1848
+
G. Hyphydrus Illiger 1802
Tr. Methlini Van den Branden 1885
+
G. Methles Sharp 1882
+
†Methles caucasicus Říha 1974
G. †Procoelambus Theobald 1937
+
†Procoelambus macrocephalus Theobald 1937
Tr. †Schistomerini Palmer 1957
+
G. †Schistomerus Palmer 1957
+
Tr. Vatellini Sharp 1880
+
G. †Calicovatellus Miller and Lubkin 2001
+
†Calicovatellus petrodytes Miller and Lubkin 2001
G. Derovatellus Sharp 1882
Subfm. Laccophilinae Gistel 1848
+
G. Japanolaccophilus Sato 1972
+
†Japanolaccophilus beatificus Balke and Hendrich 2019
Tr. Laccophilini Gistel 1848
+
G. †Electruphilus Balke et al. 2019
+
†Electruphilus wendeli Balke et al. 2019
G. Laccophilus Leach 1817
+
†Laccophilus corystes Zhang 1989
†Laccophilus pusulatus Zhang 1989
†Laccophilus strigatus von Schlechtendal 1894
Invalid names: Palaeogyrinus von Schlechtendal 1894 [synonym]
Invalid names: Palaeogyrininae von Schlechtendal 1894 [synonym]
Subfm. †Liadytiscinae Prokin and Ren 2010
+
G. †Liadroporus Prokin and Ren 2010
+
†Liadroporus elegans Prokin and Ren 2010
Tr. †Liadytiscini Prokin and Ren 2010
+
G. †Liadytiscus Prokin and Ren 2010
+
†Liadytiscus cretaceus Prokin and Ren 2010
†Liadytiscus latus Prokin and Ren 2010
†Liadytiscus longitibialis Prokin and Ren 2010
G. †Liadyxianus Prokin et al. 2013
+
†Liadyxianus kirejtshuki Prokin et al. 2013
Tr. †Mesoderini Prokin et al. 2013
+
G. †Mesoderus Prokin and Ren 2010
+
†Mesoderus magnus Prokin and Ren 2010
†Mesoderus ovatus Prokin et al. 2013
†Mesoderus punctatus Prokin et al. 2013
†Mesoderus ventralis Prokin and Ren 2010
G. †Mesodytes Prokin et al. 2013
+
†Mesodytes rhantoides Prokin et al. 2013
G. †Palaeodytes Ponomarenko 1987
+
†Palaeodytes baissiensis Prokin et al. 2013
†Palaeodytes gutta Ponomarenko 1987
†Palaeodytes incompleta Ponomarenko et al. 2005
†Palaeodytes sibiricus Ponomarenko 1987
G. †Sinoporus Prokin and Ren 2010
+
†Sinoporus lineatus Prokin and Ren 2010
Invalid names: Aubehydrinae Guignot 1942 [empty], Necticus palustris Aymard 1856 [nomen nudum]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Toarcian to the top of the Rancholabrean or 182.70000 to 0.01200 Ma
Minimum age of oldest fossil (stem group age): 174.1 Ma

Collections (175 total)


Time interval Ma Country or state Original ID and collection number
Toarcian182.7 - 174.1Russian Federation (Irkutsk) Angaragabus jurassicus (122950 152595)
Callovian - Oxfordian166.1 - 157.3Kazakhstan (Chimkent) Palaeodytes gutta (105971)
Late/Upper Berriasian145.0 - 140.2United Kingdom (England) Palaeodytes incompleta (103833)
Late/Upper Barremian130.0 - 125.45China (Nei Mongol) Mesoderus punctatus, Mesoderus ovatus, Mesodytes rhantoides (125655)
Late/Upper Barremian130.0 - 125.45China (Liaoning) Liadroporus elegans (111716) Liadytiscus cretaceus, Liadytiscus longitibialis, Liadytiscus latus, Liadroporus elegans, Mesoderus magnus, Mesoderus ventralis, Sinoporus lineatus (111712) Liadyxianus kirejtshuki (122554)
Barremian129.4 - 125.0Russian Federation (Chita) Cretodytes incertus (105968)
Aptian125.0 - 113.0Russian Federation (Buryatia) Palaeodytes baissiensis (118216)
Late/Upper Aptian122.46 - 113.0Australia (Victoria) Dytiscidae indet. 2, Dytiscidae indet. 3, Dytiscidae indet. 1 (77192)
Early/Lower Cenomanian99.6 - 93.5Myanmar Ambarticus myanmaricus (198811)
Turonian93.9 - 89.8Kazakhstan Cretodytes latipes (103274)
Tiffanian61.7 - 56.8Canada (Alberta) Dytiscidae indet. (119816)
Bridgerian50.3 - 46.2USA (Utah) Laccophilus sp. (121147)
MP 1348.6 - 40.4Germany (Eifel) Dytiscidae indet. (135641 138912)
Priabonian38.0 - 33.9France Dytiscus sp. (113709)
Priabonian38.0 - 33.9Germany Hydroporini indet. (123769)
Priabonian38.0 - 33.9Germany (Halle) Electruphilus wendeli (207248)
Priabonian38.0 - 33.9Russian Federation (Kaliningrad) Copelatus aphroditae (123770) Derovatellus rostratus, Hydroporus carstengroehni, Hydroporinae indet. 2, Coptotomus balticus (110735) Hydroporinae indet. 1 (120816) Japanolaccophilus beatificus, Coptotomus balticus (145846)
Priabonian38.0 - 33.9Poland Glessaria rostrata (109625)
Priabonian38.0 - 33.9United Kingdom (England) Hydroporinae indet. (21976) Hygrotus sp., Dytiscidae indet., Hydroporinae indet., Agabus latissimus, Ilybius gratshevi (123959) Laccophilus sp., Ilybius gratshevi (123960)
Priabonian38.0 - 33.9Denmark Hydrotrupes prometheus (123762)
Chadronian37.2 - 33.9USA (Colorado) Agabus florissantensis (124624) Coelambus miocenus, Agabus charon, Acilius florissantensis, Miodytiscus hirtipes (106381) Coelambus miocenus, Agabus florissantensis (113749) Coelambus miocenus, Bidessus laminarum, Hydroporus sedimentorum, Agabus rathbuni (106384)
Early/Lower Oligocene33.9 - 28.4France (Auvergne Region) Necticus palustris (31767)
Early/Lower Oligocene33.9 - 28.4France Dytiscus lavateri (122408) Escheria convexa (121473)
MP 2333.9 - 28.4France Procoelambus macrocephalus, Hydroporus trimaculatus (122535)
Rupelian33.9 - 28.1France (Auvergne) Eunectes antiquus (127630)
MP 3028.4 - 23.03Germany (Nordrhein-Westfalen) Palaeogyrinus strigatus (142560)
MP 3028.4 - 23.03Germany Agabus reductus (123945) Dytiscidae indet. (115802)
Chattian28.1 - 23.03Germany (North Rhine-Westphalia) Oreodytes cryptolineatus, Hydroporus macularis, Hydroporus multipunctatus, Agabus hochi, Agabus latus, Palaeogyrinus strigatus, Dytiscidae indet. (122010)
Early/Lower Miocene23.03 - 15.97Mexico Copelatus sp. (21977)
Burdigalian20.44 - 15.97China (Shandong) Laccophilus pusulatus, Laccophilus corystes (128552)
Burdigalian - Langhian20.44 - 13.82Dominican Republic Copelatus predaveterus (122982)
Hemingfordian20.43 - 15.97USA (California) Calicovatellus petrodytes (152044) Schistomerus californense (106431 123022 123023 123025 123026 123027 123028 123029 123030 123031 123032 123033 123034 123035 123036 123037 123039 123040 123041 123042 123043)
MN 320.0 - 16.9Czech Republic Cybister sp., Cybister rotundatus (105508)
MN 416.9 - 15.97Czech Republic Agabus lithax (105501)
Langhian15.97 - 13.82USA (Idaho) Dytiscus miocenicus (130391)
Langhian15.97 - 13.82USA (Washington) Dytiscus latahensis (140652)
MN 515.97 - 13.65Germany (Baden-Württemberg) Dytiscidae indet. (109058)
Middle Miocene15.97 - 11.608Russian Federation Colymbetes sp., Copelatus spp., Hydroporus lapideus, Methles caucasicus, Copelatus stavropolitanus, Copelatus ponomarenkoi, Copelatus fossilis, Colymbetes miocaenicus, Graphoderus mirabilis, Cybister fractus (105571) Copelatus spp. (138756) Cybister rotundatus, Cybister imperfectus, Cybister mancus (152089)
Middle Miocene15.97 - 11.608Australia (New South Wales) Dytiscidae indet. (224044)
Sarmatian12.7 - 11.608Germany (Baden-Württemberg) Dytiscus lavateri, Cybister agassizi, Cybister atavus, Cybister nicoleti, Hydaticus areolatus, Colymbetes aemulus, Hydroporus benzeli (117252) Dytiscus lavateri, Dytiscus oeningensis (115304) Dytiscus zschokkeanus (122210) Hydroporus antiquus (122868)
Sarmatian12.7 - 11.608Croatia Colymbetes ungeri (117246)
Sarmatian12.7 - 11.608Switzerland Cybister nicoleti (153996)
Late/Upper Miocene11.608 - 5.333Poland Hydroporus sp., Acilius sp., Hydaticus laevipennis (167589)
Late/Upper Miocene11.608 - 5.333Russian Federation Agabus sp. (153434) Agabus sp., Hydroporus sp. (113713)
Late/Upper Miocene11.608 - 5.333Germany (Baden-Württemberg) Dytiscus avunculus (152952)
Pliocene5.333 - 2.588Japan Dytiscidae indet. (165745) Ilybius sp., Dytiscidae indet. (165743)
Piacenzian3.6 - 2.588Japan Agabus sp., Ilybius sp., Dytiscidae indet. (165738) Ilybius sp. (165742)
Gelasian2.588 - 1.806Greenland Agabini indet. (160146) Agabus bifarius (160163) Agabus spp., Agabus affinis, Dytiscidae indet. (160108) Agabus spp., Colymbetes sp., Hydroporus spp., Agabinae indet., Oreodytes spp., Ilybius sp., Hygrotus sp., Agabini indet., Hydroporus morio, Hydroporus acutangulus, Hydroporus striola, Hydroporus oblitus, Oreodytes alpinus, Agabus affinis, Agabus clavicornis, Agabus bifarius, Agabus anthracinus, Ilybius vittiger, Dytiscidae indet. (160109) Agabus spp., Hydroporus spp., Agabus bifarius (160145) Agabus spp., Hydroporus spp., Ilybius sp., Hydroporus morio, Oreodytes alpinus, Agabus bifarius, Dytiscidae indet. (160153) Agabus spp., Hydroporus spp., Ilybius sp., Oreodytes alpinus, Agabus affinis, Agabus clavicornis, Hydroporinae indet. (160143) Hydroporus morio (160162) Hydroporus sp., Oreodytes alpinus (160147) Hydroporus spp., Ilybius sp., Agabini indet., Hydroporus morio, Oreodytes sanmarkii, Agabus affinis, Agabus anthracinus, Dytiscidae indet., Hydroporinae indet. (160144) Ilybius sp. (160151)
Early/Lower Pleistocene2.588 - 0.781Russian Federation Agabus sp., Hydroporus sp. (163089) Agabus spp., Colymbetes paykulli (163069)
Early/Lower Pleistocene2.588 - 0.781Japan Agabus sp. (165711)
Pleistocene2.588 - 0.0117Italy Dytiscus zersii (224200)
Calabrian1.8 - 0.774Japan Dytiscus sp., Agabus sp., Ilybius sp., Rhantus sp., Agabus optatus (160539)
Calabrian - Ionian1.8 - 0.129United Kingdom (Durham) Hydroporus edenianus (140489)
Chibanian0.774 - 0.129Japan (Kumamoto) Dytiscidae indet. (164901)
Chibanian0.774 - 0.129Russian Federation Agabus sp. (163100) Hydroporus sp. (163097 163099)
Chibanian0.774 - 0.129Japan Agabus sp. (165706) Dytiscus sp., Ilybius sp., Dytiscidae indet. (165700) Platambus stygius (165704 165705)
Middle Pleistocene0.774 - 0.129United Kingdom (Warwickshire) Agabus sp. (169388) Graphoderus sp. (169390) Ilybius sp. (169391 169396)
Middle Pleistocene0.774 - 0.129Japan Agabus sp. (165709) Ilybius sp., Dytiscidae indet. (165708)
Middle Pleistocene0.774 - 0.129Japan (Kumamoto) Agabus sp. (165715 165716)
Middle Pleistocene0.774 - 0.129United Kingdom Colymbetes sp. (168264 168268) Colymbetes sp., Agabus bipustulatus, Hygrotus decoratus, Potamonectes depressus (168270) Colymbetes sp., Ilybius sp. (168269) Hydroporus sp., Ilybius sp. (168265) Hygrotus decoratus (168267) Ilybius sp., Agabus bipustulatus, Potamonectes depressus, Cybister lateralimarginalis (168266) Potamonectes depressus (168272)
Rancholabrean0.3 - 0.012USA (California) Dytiscus marginicollis (150336)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (England) Agabus sp., Colymbetes sp. (164920) Colymbetes sp. (164913 164917) Hydroporus sp., Rhantus sp. (164905)
Late/Upper Pleistocene0.129 - 0.0117Canada (British Columbia) Dytiscidae indet. (167673)
Late/Upper Pleistocene0.129 - 0.0117Ukraine Coelambus picipoides, Coelambus dzieduszyckii, Coelambus niedzwiedzkii, Coelambus ganglbaueri, Coelambus latefasciatus, Coelambus ozokeriticus, Hydroporus borealis, Hydroporus praedorsalis, Hydroporus lapponum, Hydroporus subarcticus, Hydroporus praenivalis, Hydroporus praenigrita, Hydroporus pleistocenicus, Hydroporus sandbergeri, Hydroporus clessini, Agabus congeneroides, Agabus corticeus, Agabus reitteri, Agabus parvulus, Agabus niedzwiedzkii, Ilybius boryslavicus, Rantus praesuturellus, Cymatopterus striatus, Cymatopterus dolabratus, Acilius praesulcatus, Dytiscus lapponicus (153476)
Late/Upper Pleistocene0.129 - 0.0117USA (Pennsylvania) Colymbetes sp. (137208)
Late/Upper Pleistocene0.129 - 0.0117United Kingdom (Worcestershire) Hydroporus spp., Ilybius sp., Rantus sp., Colymbetes paykulli, Hygrotus novemlineatus, Hygrotus impressopunctatus, Deronectes sp., Deronectes griseostriatus, Hydroporus arcticus, Hydroporus palustris, Hydroporus planus, Agabus congener, Agabus nebulosus, Agabus arcticus, Agabus sturmii, Agabus bipustulatus, Ilybius subaeneus, Rantus exsoletus, Colymbetes fuscus, Dytiscus marginalis (163070)
Late/Upper Pleistocene0.129 - 0.0117Canada (Ontario) Coelambus cribrarius, Hydroporus inundatus (152576) Coelambus derelictus, Coelambus disjectus, Hydroporus inanimatus, Hydroporus sectus, Agabus perditus (152577) Coelambus infernalis, Hydroporus sectus (152575) Potamonectes sp. (174229)
Late/Upper Pleistocene0.129 - 0.0117USA (Massachusetts) Dytiscidae indet. (152284)
Late/Upper Pleistocene0.129 - 0.0117USA (Illinois) Agabus savagei, Agabus praelugens (164922)
Late/Upper Pleistocene0.129 - 0.0117USA (Colorado) Agabus infuscatus (169347 169350 169353)
Late/Upper Pleistocene0.129 - 0.0117Peru Dytiscidae indet. (71346)
Late/Upper Pleistocene0.129 - 0.0117USA (Washington) Colymbetes sp., Ilybius sp. (157092) Dytiscidae indet. (169366 169371 169372 169373 169375 169376 169377 169378) Dytiscus sp. (169379) Dytiscus sp., Agabus spp., Colymbetes sp., Oreodytes sp., Rhantus sp., Hygrotus sp., Ilybius quadrimaculatus (167662)
Late/Upper Pleistocene0.129 - 0.0117Japan Dytiscus sp., Agabus sp., Ilybius sp., Ilybius apicalis, Dytiscidae indet. (166082) Ilybius sp. (165701)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation Agabus sp. (153443) Agabus sp., Hydroporus sp. (153446 173208 173216) Hydroporus sp. (173207)
Late/Upper Pleistocene0.129 - 0.0117Russian Federation (Astrakhan) Agabus sp., Ilybius fenestratus, Graphoderus austriacus, Coelambus parallelogrammus, Hydroporinae indet. (125794)
Late/Upper Pleistocene0.129 - 0.0117Argentina (Buenos Aires) Megadytes glaucus (184124)
Late/Upper Pleistocene0.129 - 0.0117USA (Wisconsin) Hydroporini indet. (169406)
Tarantian0.129 - 0.0117USA (Pennsylvania) Colymbetes sp. (137204) Dytiscidae indet. (137201) Hydroporus sp., Dytiscidae indet. (137205 137206)