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Royalella quadrolirata

Gastropoda - Heterostropha - Donaldinidae

Taxonomy
Donaldina quadroliratus was named by Anderson et al. (1990). Its type specimen is OSU 34991, a shell, and it is a 3D body fossil. Its type locality is Ty-3. 60 cm shale zone, immediately above limestone at mall abandoned borrow pi, which is in a Desmoinesian shallow subtidal limestone/shale in the Putnam Hill Formation of Ohio.

It was recombined as Royalella quadrolirata by Mazaev (2015).

Synonymy list
YearName and author
1990Donaldina quadroliratus Anderson et al. pp. 560 - 561 figs. 1.1-1.4
2015Royalella quadrolirata Mazaev p. 891

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Spiralia
superphylumLophotrochozoa
phylumMollusca
RankNameAuthor
classGastropodaCuvier 1797
orderHeterostropha()
superfamilyStreptacidoideaKnight 1931
familyDonaldinidae
genusRoyalella
speciesquadrolirata()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
J. R. Anderson et al. 1990Very high-spired shell, up to 13 whorls, with low spiral angle and low whorl expansion; four revolving lirae on each whorl, two bold lirae in middle of whorl, one faint lira on both upper and lower whorl surfaces.

Small, narrow, low spiral angle, high-spired shell; subrounded whorl profile; anomphalous, sutures shallowly impressed; protoconch of one whorl, smooth, heterostrophic; apical end of protoconch slightly upturned; aperture subcircular in shape; outer lip slightly thickened; inner lip slightly flared; shell composed of as many as 13 whorls, including protoconch.

Teleoconch whorls have thick revolving lirae with deep interareas; first two whorls of teleoconch have faint ornamentation of two revolving lirae; later whorls have two pronounced lirae in middle of each whorl, flanked by two smaller lirae near sutures; growth lines faint.

Whorl cross section circular to subcircular; three shell microstructural layers present; shell thickness ranges from 0.01 to 0.06 mm; slightly arcuate columella. Average measurements and size ranges given in Table 1.