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Waagenella crassus

Gastropoda - Bellerophontida - Bellerophontidae

Taxonomy
Bellerophon crassus was named by Meek and Worthen (1860). It is not a trace fossil.

It was recombined as Bellerophon (Bellerophon) crassus by Yochelson and Saunders (1967), Kues (1984), Batten (1995), Hoare et al. (1997) and Kues and Batten (2001); it was recombined as Waagenella crassus by Wagner (2017).

Synonymy list
YearName and author
1860Bellerophon crassus Meek and Worthen p. 458
1866Bellerophon crassus Meek and Worthen p. 385 figs. pl. 16a-b
1891Bellerophon crassus White p. 26
1909Bellerophon crassus Girty p. 479 figs. Pl 29, fig 16
1967Bellerophon (Bellerophon) crassus Yochelson and Saunders p. 40
1977Bellerophon crassus Termier and Termier p. 62 fig. 11
1984Bellerophon (Bellerophon) crassus Kues figs. f. 3.1-3.2
1995Bellerophon (Bellerophon) crassus Batten p. 7 fig. 2
1997Bellerophon (Bellerophon) crassus Hoare et al. p. 1026
2001Bellerophon (Bellerophon) crassus Kues and Batten pp. 11 - 12 figs. f. 2.31 - 2.33
2017Waagenella crassus Wagner p. S4050

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Spiralia
superphylumLophotrochozoa
phylumMollusca
classGastropodaCuvier 1797
RankNameAuthor
orderBellerophontidaUlrich and Scofield 1897
suborderBellerophontina
superfamilyBellerophontoidea()
familyBellerophontidae
subfamilyBellerophontinae
genusWaagenella
speciescrassus()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
F. B. Meek and A. H. Worthen 1860Shell large, very thick, subglobose, or a little longer than wide; volutions expanding somewhat rapidly, rounded over the dorsuim and sides; umbilical region excavated, but not perforate; mesial band narrow, rather prominent, and margined on each side by a raised line; aperture transverse, reniform, or sublunate; lip strongly thickened near the umbilical excavations on each side, but not covering them, thinner and but slightly prominent on either side of the narrow sinus in its outer margin, and spreading in the form of a moderately thick, smooth, callus, over the inner whorls within the aperture, and between the callosities on each side; surface marked by rather distinct irreegular wrinkles, and lines of growth, the former of which are strongest on each side near the lip. Greatest length 2 20 inches; breadth at the aperture 2 inches; breadth of the mesial band near the aperture 0 12 inch.
F. B. Meek and A. H. Worthen 1866Shell large, very thick, subglobose, or a little longer than wide; volutions expanding moderately, rounded over the dorsum and sides; umbilical region somewhat excavated; mesial band narrow, rather prominent, and subangular; aperture transverse, reniform or sublunate; lip strongly thickened near the umbilical excavations on each side, but not covering them, thinner and moderately prominent in outline on each side of the narrow sinus in the outer margin, and spreading in the form of a rather thick, smooth callous, over the inner whorls, and between the callosites on each side ; surface marked by rather distinct striae of growth, and irregular, transverse wrinkles, which latter are strongest in the umbilical region and near the aperture ; lines of growth and wrinkles making a very gentle forward curve in crossing from the umbilicus to the dorsal band. Greatest length, 2.26 inches; breadth, 2 inches; ditto of aperture, 1.70 inches; breadth of mesial band near the aperture, about 0.12 inch.
R. L. Batten 1995Small, compressed bellerophontiform shells. Whorl expansion rate moderate, forming an evenly developed, nearly equiangular spiral shell. Selenizone flush to slightly raised above dorsum. Lateral lips slightly reflexed in area of umbilici and nearly straight over dorsum to selenizone where they form a rectangular-shaped sinus. Selenizone is flattened to convex and has straight lunulae with little arcing. Selenizone margins are sharply indented, narrow troughs. Growth lines are slightly imbricated and, on some specimens, clustered into broad, flat ribs. Anomphalus.
B. S. Kues and R. L. Batten 2001Shel1 large (up to 48 mm in length), slightly compressed laterally, expanding moderately with growth anteriorly so that maximum width nearly equals length; transverse profile broadly and gently convex across midline before sloping more curved, becoming slightly flattened and gently geniculate near
plane of aperture, owing to a slightly thickened inductural pad;
lateral lips thickened, gently reflexed, meeting whorl side at a 60-
80 degree angle; partly covering slightly depressed umbilical area;
aperture reniform, moderately wide; whorl surface with fine,
closely-spaced, somewhat irregular to lamellate growth lines; selenizone narrow, flat, flush with shell surface, bordered by a fine, sharp lira along each side for most of body whorl circumference, but becoming a relatively broad rounded crest near anterior margin of largest specimen; details of apertural lips and slit unknown.