PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Trachyceratoidea

Cephalopoda - Ceratitida

Taxonomy

Trachycerataceae was named by Haug (1894).

It was corrected as Trachyceratoidea by Urlichs (2017) and Krystyn et al. (2019).

It was assigned to Trachyceratina by Tatzreiter (1981); and to Ceratitida by Tozer (1981), Fantini Sestini (1994), Urlichs (2017) and Krystyn et al. (2019).

Subtaxa

Arpaditidae, Buchitidae, Distichitidae, Dronovitidae, Noridiscitidae, Trachyceratidae (syn. Nannitidae)

Synonymy list

Year	Name and author
1894	Trachycerataceae Haug
1981	Trachycerataceae Tatzreiter
1981	Trachycerataceae Tozer
1994	Trachycerataceae Fantini Sestini
2017	Trachyceratoidea Urlichs p. 177
2019	Trachyceratoidea Krystyn et al. p. 239

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Spiralia
superphylum	Lophotrochozoa

Rank	Name	Author
phylum	Mollusca
class	Cephalopoda	Cuvier 1797
subclass	Ammonoidea	()
order	Ceratitida	Hyatt 1884
superfamily	Trachyceratoidea	(Haug 1894)
superfamily	Trachyceratoidea	(Haug 1894)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Superfm. †Trachyceratoidea Haug 1894

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Fm. †Buchitidae Hyatt 1900

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G. †Buchites Mojsisovics 1900

†Buchites aldrovandii Mojsisovics 1893

Fm. †Distichitidae Diener 1920

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G. †Distichites Mojsisovics 1893

†Distichites canadensis McLearn 1940

†Distichites celticus Mojsisovics 1893

†Distichites columbianus Tozer 1994

†Distichites falcatus Diener 1923

†Distichites gethingi McLearn 1940

†Distichites harpalos Dittmar 1866

Invalid names: Distichites palliseri McLearn 1947 [synonym]

†Distichites kmetyi Mojsisovics 1893

Invalid names: Distichites younghusbandi Diener 1906 [synonym]

†Distichites leptacanthus Diener 1923

†Distichites megacanthus Mojsisovics 1893

Invalid names: Distichites hypsacanthus Diener 1923 [synonym]

†Distichites mesacanthus Diener 1923

†Distichites minos Mojsisovics 1893

†Distichites sollasii Diener 1906

†Distichites tropicus Diener 1923

G. †Ectolcites Mojsisovics 1893

†Ectolcites childerhosei Tozer 1994

†Ectolcites hochstetteri Mojsisovics 1893

†Ectolcites hollandi Diener 1906

†Ectolcites pseudoaries von Hauer 1850

Invalid names: Ectolcites arietiformis Diener 1906 [synonym], Ectolcites duncani Diener 1906 [synonym]

†Ectolcites sidoniae Diener 1920

G. †Leiodistichites Tozer 1994

†Leiodistichites anacanthus Diener 1923

†Leiodistichites beachi Tozer 1994

†Leiodistichites hacqueti Mojsisovics 1893

†Leiodistichites ursidens Tozer 1994

G. †Mesodistichites Krystyn et al. 2019

†Mesodistichites evolutus Krystyn et al. 2019

†Mesodistichites tozeri Seyed-Emami 1975

G. †Paradistichites Diener 1916

†Paradistichites ectolcitiformis Diener 1906

G. †Pleurodistichites Tozer 1980

†Pleurodistichites hindei von Mojsisovics 1896

†Pleurodistichites lilli Guembel 1861

†Pleurodistichites stotti Tozer 1980

†Pleurodistichites trailli Diener 1906

Invalid names: Dittmarites lilliformis Diener 1908 [synonym]

G. †Trachypleuraspidites Diener 1906

†Trachypleuraspidites griffithi Diener 1906

Invalid names: Distichites tingriensis Wang and He 1976 [synonym]

†Trachypleuraspidites malayicus Welter 1914

Fm. †Dronovitidae Shevyrev 1990

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G. †Dronovites Shevyrev 1990

†Dronovites pamiricus Shevyrev 1990

Fm. †Noridiscitidae Spath 1951

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G. †Nairites Kiparisova and Azarian 1963

†Nairites armenius Kiparisova and Azarian 1963

†Nairites laevis Kiparisova and Azarian 1963

G. †Noridiscites Spath 1951

†Noridiscites nodosus Krystyn et al. 2019

†Noridiscites viator Mojsisovics 1893

Fm. †Trachyceratidae Haug 1894

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Subfm. †Anolcitinae Mietto and Manfrin 2008

G. †Anolcites Mojsisovics 1893

†Anolcites anguinus Tozer 1994

†Anolcites angustus Tozer 1994

†Anolcites dieneri Jaworski 1922

†Anolcites doleriticus von Mojsisovics 1869

†Anolcites gemmatus Tozer 1994

†Anolcites impolitus Tozer 1994

†Anolcites papillatus Tozer 1994

†Anolcites politus Tozer 1994

†Anolcites rasilis Tozer 1994

†Trachyceras (Anolcites) barberi Smith 1914

†Trachyceras (Anolcites) drakei Smith 1914

G. †Asklepioceras Renz 1911

†Asklepioceras altilis Tozer 1994

†Asklepioceras exilis Tozer 1994

†Asklepioceras laurenci McLearn 1940

Invalid names: Asklepioceras mahaffii McLearn 1947 [synonym]

†Asklepioceras segmentatum Mojsisovics 1893

†Asklepioceras squammatum Arthaber 1915

G. †Daxatina Strand 1929

†Daxatina canadensis Whiteaves 1889

†Daxatina laubei Tozer 1994

†Daxatina limpida Tozer 1994

†Daxatina megabrotheus Tozer 1994

Invalid names: Dawsonites Boehm 1903 [replaced]

G. †Falsanolcites Rieber and Brack 2004

†Falsanolcites arminae Mojsisovics 1893

†Falsanolcites cappellinii Tommasi 1899

†Falsanolcites clapsavonum von Mojsisovics 1882

†Falsanolcites doleriticum von Mojsisovics 1869

†Falsanolcites elisabethae Mojsisovics 1893

†Falsanolcites furcosus Mojsisovics 1893

†Falsanolcites gervasuttii Fantini Sestini 1994

†Falsanolcites gortanii Pisa 1966

†Falsanolcites julium von Mojsisovics 1882

†Falsanolcites laczkoi Diener 1899

†Falsanolcites recubariense Mojsisovics 1882

†Falsanolcites richthofeni Mojsisovics 1882

†Falsanolcites rieberi Fantini Sestini 1994

G. †Frankites Tozer 1971

†Frankites apertus Mojsisovics 1893

†Frankites glaber Tozer 1994

†Frankites regoledanus Mojsisovics 1869

Invalid names: Trachyceras mundevillae Mojsisovics 1882 [synonym]

†Frankites sutherlandi McLearn 1947

G. †Maclearnoceras Tozer 1963

†Maclearnoceras ensio Tozer 1994

†Maclearnoceras maclearni Tozer 1963

G. †Zestoceras Tozer 1994

†Zestoceras barwicki Johnson 1941

Invalid names: Zestoceras cerastes Tozer 1994 [synonym]

†Zestoceras enode Tozer 1972

†Zestoceras lorigae Mietto and Manfrin 2008

†Zestoceras nitidum Tozer 1994

Subfm. †Arpaditinae Hyatt 1900

G. †Arctoarpadites Tozer 1994

†Arctoarpadites costatus Tozer 1961

G. †Arctosirenites Tozer 1961

†Arctosirenites canadensis Tozer 1961

Invalid names: Striatosirenites seimkanensis Bychkov 1973 [synonym]

†Arctosirenites columbianus Tozer 1994

†Arctosirenites southeri Tozer 1994

†Arctosirenites sverdrupi Kittl 1907

G. †Argolites Renz 1939

†Argolites arietiformis De Toni 1913

†Argolites arpaditoides von Mojsisovics 1882

†Argolites celtitoides Airaghi 1902

†Argolites fortis Fantini Sestini 1994

†Argolites mojsisovicsi De Lorenzo 1896

†Argolites paronai Mariani 1902

†Argolites trinodosus Balini et al. 2006

G. †Arpadites Mojsisovics 1879

†Arpadites arpadis Mojsisovics 1870

†Arpadites bracki Fantini Sestini 1994

†Arpadites busiris Munster 1834

Invalid names: Ammonites bidenticulatus Klipstein 1843 [synonym], Ammonites subdenticulatus Klipstein 1843 [synonym], Ceratites basileus Münster 1841 [synonym]

†Arpadites cinensis von Mojsisovics 1882

†Arpadites dichotomus Fantini Sestini 1994

†Arpadites helenae von Mojsisovics 1882

†Arpadites kingi Smith 1927

†Arpadites lissarensis von Mojsisovics 1896

†Arpadites manzonii Benecke 1876

†Arpadites orientalis von Mojsisovics 1879

†Arpadites pilari von Mojsisovics 1882

†Arpadites pulcher Fantini Sestini 1994

†Arpadites rimosus Münster 1841

†Arpadites rueppeli Klipstein 1843

†Arpadites sakawanus von Mojsisovics 1888

†Arpadites sesostris Laube 1869

†Arpadites stracheyi von Mojsisovics 1896

†Arpadites szaboi Boeckh 1873

†Arpadites telleri von Mojsisovics 1882

†Arpadites toldyi von Mojsisovics 1882

†Arpadites trettensis von Mojsisovics 1882

†Arpadites vaceki von Mojsisovics 1882

G. †Dittmarites Mojsisovics 1893

G. †Edmundites Diener 1916

†Edmundites rimkinensis von Mojsisovics 1896

G. †Hisnitites Tozer 1994

†Hisnitites janmulleri Tozer 1994

G. †Klipsteinia von Mojsisovics 1882

†Klipsteinia achelous Munster 1834

Invalid names: Ammonites hoplophorus Giebel 1852 [synonym], Goniatites armatus Münster 1841 [synonym], Goniatites friesei Münster 1841 [synonym]

†Klipsteinia boetus Münster 1841

†Klipsteinia disciformis Lukeneder and Lukeneder 2014

†Klipsteinia irregularis Münster 1841

†Klipsteinia nataliae von Mojsisovics 1882

†Klipsteinia radiata Klipstein 1843

Invalid names: Dinarites eduardi von Mojsisovics 1882 [synonym]

G. †Liardites Tozer 1963

†Liardites whiteavesi Tozer 1963

G. †Meginoceras McLearn 1930

†Meginoceras aylardi McLearn 1947

†Meginoceras caurinum McLearn 1947

†Meginoceras effervescens Tozer 1994

†Meginoceras meginae McLearn 1930

†Meginoceras tetsa McLearn 1947

†Meginoceras triviale Tozer 1994

G. †Otoarpadites Tozer 1994

†Otoarpadites auritus Tozer 1994

G. †Silenticeras McLearn 1930

†Silenticeras bamberi Tozer 1994

†Silenticeras gibsoni Tozer 1994

†Silenticeras hatae McLearn 1930

†Silenticeras involutum Tozer 1994

†Silenticeras liardense Tozer 1994

G. †Trachystenoceras Johnston 1941

†Trachystenoceras gabbi Hyatt and Smith 1905

G. †Yakutosirenites Tozer 1994

†Yakutosirenites pentastichus Vozin 1964

G. †Boreotrachyceras Konstantinov 2012

†Boreotrachyceras omkutchanicum Bychkov 1973

G. †Brotheotrachyceras Urlichs 1994

†Brotheotrachyceras brotheus Munster 1834

†Brotheotrachyceras difforme Munster 1834

G. †Hannaoceras Tomlin 1931

†Hannaoceras nasturtium Dittmar 1866

Invalid names: Polycyclus Mojsisovics 1889 [replaced], Polysphinctoceras Spath 1934 [objective synonym], Smithoceras Hanna 1924 [replaced]

Subfm. †Haoceratinae Balini and Zou 2015

G. †Haoceras Balini and Zou 2015

†Haoceras xingyiense Balini and Zou 2015

G. †Sinomeginoceras Balini and Zou 2015

†Sinomeginoceras wangi Balini and Zou 2015

†Sinomeginoceras xingyiense Balini and Zou 2015

G. †Langdaiceras Wang 1983

†Langdaiceras pateriformis Wang 1983

G. †Nannites Mojsisovics 1881

†Nannites bittneri von Mojsisovics 1881

†Nannites fugax von Mojsisovics 1882

†Nannites spurius Münster 1841

Invalid names: Goniatites blumii Klipstein 1843 [synonym]

G. †Okhototrachyceras Konstantinov 2012

†Okhototrachyceras seimkanense Bychkov 1973

Subfm. †Protrachyceratinae Tozer 1971

G. †Eoprotrachyceras Tozer 1980

†Eoprotrachyceras americanum Mojsisovics 1882

†Eoprotrachyceras curionii Mojsisovics 1882

†Eoprotrachyceras curionii ramonensis Parnes 1962

†Eoprotrachyceras dunni Smith 1914

†Eoprotrachyceras gibsoni Tozer 1994

†Eoprotrachyceras gredleri Mojsisovics 1882

†Eoprotrachyceras lahontanum Smith 1914

†Eoprotrachyceras matutinum Tozer 1980

†Eoprotrachyceras meeki Mojsisovics 1882

†Eoprotrachyceras subasperum Meek 1877

G. †Paratrachyceras Arthaber 1915

†Paratrachyceras hofmanni Boekh 1873

†Paratrachyceras ulynense Bychkov 1973

G. †Protrachyceras Mojsisovics 1893

†Protrachyceras anatolicum Toula 1898

†Protrachyceras archelaus Laube 1869

†Protrachyceras cautleyi Diener 1907

†Protrachyceras costulatum Mansuy 1912

†Protrachyceras deprati Mansuy 1912

†Protrachyceras guizhouense Wang 1983

†Protrachyceras hispanicum von Mojsisovics 1881

†Protrachyceras ibericum Mojsisovics 1882

†Protrachyceras irregulare Fantini Sestini 1994

†Protrachyceras ladinum Mojsisovics 1882

Invalid names: Ammonites aequinodosus Klipstein 1843 [synonym], Trachyceras altum von Mojsisovics 1878 [synonym]

†Protrachyceras lermani Parnes 1986

†Protrachyceras longobardicum Mojsisovics 1882

†Protrachyceras margaritosum von Mojsisovics 1882

†Protrachyceras mutabile Wang 1983

†Protrachyceras negevense Parnes 1986

†Protrachyceras parinaense Fantini Sestini 1994

†Protrachyceras pseudoarchelaus Boeckh 1873

†Protrachyceras sikanianum McLearn 1930

Invalid names: Protrachyceras zauwae McLearn 1930 [synonym]

†Protrachyceras sirenitiforme Parnes 1962

†Protrachyceras spitiense Diener 1908

†Protrachyceras steinmanni von Mojsisovics 1882

†Protrachyceras wahrmani Parnes 1962

†Trachyceras (Protrachyceras) ralphuanum von Mojsisovics 1896

†Trachyceras (Protrachyceras) springeri Smith 1914

†Trachyceras (Protrachyceras) storrsi Smith 1927

Invalid names: Protrachyceras ladinum parana Parnes 1962 [replaced]

G. †Spirogmoceras Silberling 1956

†Spirogmoceras shastense Smith 1904

Invalid names: Trachyceras (Protrachyceras) beckeri Smith 1927 [synonym], Trachyceras (Protrachyceras) californicum Smith 1927 [synonym], Trachyceras (Protrachyceras) lecontei Hyatt and Smith 1905 [synonym], Trachyceras (Protrachyceras) lindgreni Smith 1927 [synonym], Trachyceras (Protrachyceras) madisonense Smith 1927 [synonym]

Subfm. †Sirenitinae Tozer 1971

G. †Anasirenites Mojsisovics 1893

†Anasirenites briseis Mojsisovics 1893

†Anasirenites crassicrenulatus Lukeneder and Lukeneder 2014

†Anasirenites ekkehardi Mojsisovics 1893

G. †Diplosirenites Mojsisovics 1893

†Diplosirenites raineri Mojsisovics 1893

G. †Neosirenites Popov 1961

†Neosirenites pseudopentastichus Bychkov 1973

G. †Norosirenites Tozer 1994

†Norosirenites krystyni Tozer 1994

G. †Orientosirenites Konstantinov 2018

†Orientosirenites bytschkovi Konstantinov 2018

†Orientosirenites yakutensis Kiparisova 1937

G. †Pamphagosirenites Popov 1961

†Pamphagosirenites pacificus Tozer 1994

†Pamphagosirenites pamphagus Dittmar 1866

G. †Pseudosirenites Arthaber 1911

†Pseudosirenites bullatus Tozer 1994

†Pseudosirenites elegans von Mojsisovics 1896

Invalid names: Sirenites elegantiformis Diener 1906 [synonym]

†Pseudosirenites evae Mojsisovics 1893

Invalid names: Sirenites richteri von Mojsisovics 1896 [synonym]

†Pseudosirenites falcatus Tozer 1994

†Pseudosirenites pardoneti McLearn 1940

Invalid names: Sirenites inaequiornatus McLearn 1947 [synonym]

†Pseudosirenites pressus McLearn 1947

†Pseudosirenites stachei Mojsisovics 1893

G. †Pseudotibetites Tozer 1980

G. †Pterosirenites Tozer 1980

†Pterosirenites auritus Tozer 1980

†Pterosirenites nelgehensis Arkhipov 1974

†Pterosirenites obrucevi Bajarunas 1932

†Pterosirenites seimkanensis Bychkov 1974

G. †Seimkanites Konstantinov 1999

†Seimkanites aculeatus Bychkov et al. 1976

G. †Sirenites Mojsisovics 1893

†Sirenites betulinus Dittmar 1886

†Sirenites hayesi Smith 1927

†Sirenites homfrayi Gabb 1864

†Sirenites irregularis Kiparisova 1937

†Sirenites lawsoni Hyatt and Smith 1905

†Sirenites malayicus Welter 1914

†Sirenites nanseni Tozer 1961

†Sirenites ovinus Tozer 1994

†Sirenites senticosus Dittmar 1866

†Sirenites serotinus Tozer 1994

†Sirenites smithi Burckhardt and Scalia 1905

G. †Sirenotrachyceras Krystyn 1978

†Sirenotrachyceras furcatum Münster 1841

Invalid names: Ammonites nodocostatus Klipstein 1843 [synonym], Ceratites agassizii Klipstein 1843 [synonym]

†Sirenotrachyceras hadwigae Mojsisovics 1893

†Sirenotrachyceras rudolphi von Mojsisovics 1882

†Sirenotrachyceras thusneldae Mojsisovics 1893

Invalid names: Anolcites (Trachyceras) carnerii Mojsisovics 1893 [synonym], Anolcites (Trachyceras) lenaui Mojsisovics 1893 [synonym]

G. †Striatosirenites Popov 1961

†Striatosirenites buralkitensis Popov 1961

†Striatosirenites kedonensis Bychkov 1973

†Striatosirenites kinasovi Bychkov et al. 1976

†Striatosirenites repini Bychkov et al. 1976

†Striatosirenites striatofalcatus Hauer 1847

†Striatosirenites ulynensis Bychkov 1973

G. †Vredenburgites Diener 1916

†Vredenburgites vredenburgi Diener 1906

Invalid names: Sirenites nabeschi McLearn 1960 [nomen dubium]

Subfm. †Trachyceratinae Haug 1894

G. †Austrotrachyceras Krystyn 1978

†Austrotrachyceras obesum Tozer 1967

G. †Muensterites Mojsisovics 1893

†Muensterites delicatulus McLearn 1947

†Muensterites ectodus Mojsisovics 1893

†Muensterites glaciensis McLearn 1940

†Muensterites helenae Renz 1911

†Muensterites hirschi Laube 1869

†Muensterites karreri von Mojsisovics 1882

†Muensterites okeani Münster 1841

Invalid names: Ceratites jaegeri Klipstein 1843 [synonym]

G. †Neoprotrachyceras Krystyn 1978

†Neoprotrachyceras attila Mojsisovics 1870

†Neoprotrachyceras baconicum Mojsisovics 1870

G. †Trachyceras Laube 1869

†Trachyceras acutocostatum Klipstein 1843

†Trachyceras amicum von Mojsisovics 1882

†Trachyceras aon Munster 1834

†Ammonites aon bipunctatus Munster 1834

Invalid names: Ammonites armatocingulatus Klipstein 1843 [synonym], Ammonites credneri Klipstein 1843 [synonym], Ammonites dechenii Klipstein 1843 [synonym], Ammonites larva Klipstein 1843 [synonym], Ammonites mirabilis Klipstein 1843 [synonym], Ammonites spinulocostatus Klipstein 1843 [synonym], Ammonites velthemii Klipstein 1843 [synonym], Ceratites brevicostatus Klipstein 1843 [synonym], Ceratites meriani Klipstein 1843 [synonym], Trachyceras sulciferum Münster 1841 [synonym]

†Trachyceras aonoides Mojsisovics 1882

†Trachyceras austriacum von Mojsisovics 1871

†Trachyceras avisianum von Mojsisovics 1882

†Trachyceras bipunctatum Munster 1834

†Trachyceras bouei Klipstein 1843

†Trachyceras cordevolicum von Mojsisovics 1878

†Trachyceras desatoyense Johnston 1941

†Trachyceras dichotomum Münster 1841

†Trachyceras douvillei Mansuy 1912

†Trachyceras epolense Mojsisovics 1878

†Trachyceras fedaiae Salomon 1895

†Trachyceras hacqueti von Mojsisovics 1882

†Trachyceras humboldti Klipstein 1843

†Trachyceras ibericum von Mojsisovics 1881

†Trachyceras infundibuliforme Klipstein 1843

†Trachyceras judicaricum von Mojsisovics 1869

†Trachyceras judicaricum subasperum Meek 1877

†Trachyceras laricum Mojsisovics 1882

†Trachyceras liepoldti von Mojsisovics 1878

†Trachyceras mandelslohi Klipstein 1843

†Trachyceras medusae Mojsisovics 1882

†Trachyceras muensteri Wissmann 1841

Invalid names: Ammonites nodulosocostatus Klipstein 1843 [synonym], Ceratites zeuschneri Klipstein 1843 [synonym], Goniatites ornatus Klipstein 1843 [synonym], Trachyceras candaules Laube 1869 [synonym]

†Trachyceras multituberculatum Hsu 1940

†Trachyceras neumayri von Mojsisovics 1869

†Trachyceras oenanum von Mojsisovics 1882

†Trachyceras paronai Salomon 1895

†Trachyceras pescolense von Mojsisovics 1882

†Trachyceras pontius Laube 1869

†Trachyceras pustericum Mojsisovics 1878

†Trachyceras roderici von Mojsisovics 1882

†Trachyceras rutoranum Mojsisovics 1882

†Trachyceras silberlingi Balini et al. 2012

†Trachyceras stuerzenbaumi von Mojsisovics 1882

†Trachyceras symmetricum Salomon 1895

†Trachyceras tibeticum von Mojsisovics 1896

†Trachyceras trinodosum von Mojsisovics 1878

†Trachyceras villanovae D'Archiac 1860

Invalid names: Nannitidae Diener 1897 [synonym]

Invalid names: Arpaditidae Hyatt 1900 [empty]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

aragoniteo

Environment:

marinec

Locomotion:

actively mobileo

Life habit:

nektonico

Diet:

carnivoreo

Vision:

well-developedc

Created:

2005-01-28 02:33:06

Modified:

2005-10-06 16:36:47

Source: o = order, c = class

References: Kiessling 2003, Kiessling 2004

Age range: base of the Smithian to the top of the Norian or 251.30000 to 208.50000 Ma

Collections (703 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Smithian	251.3 - 247.2	Russian Federation	Nannites simplex (212291)
Anisian	247.2 - 242.0	Thailand (Lampang)	Trachyceras regoledanum (211615)
Anisian	247.2 - 242.0	USA (Nevada)	Trachyceras meeki (33207)
Anisian	247.2 - 242.0	Austria (Innsbruck)	Protrachyceras archelaus (202275)
Anisian	247.2 - 242.0	Japan (Shikoku)	Arpadites sakawanus (215820)
Anisian	247.2 - 242.0	India (Himachal Pradesh)	Protrachyceras longobardicum, Protrachyceras cautleyi (206205)
Anisian	247.2 - 242.0	Italy	Arpadites arpadis (128184) Trachyceras curionii, Arpadites arpadis (135878)
Anisian	247.2 - 242.0	Germany	Ceratites muensteri (215587)
Anisian	247.2 - 242.0	Germany (Niedersachsen)	Ceratites muensteri (215586 215633)
Anisian	247.2 - 242.0	Japan (Miyagi)	Arpadites sp. (215818)
Youngest occurrences
Time interval	Ma	Country or state	PBDB collection number
Lacian	221.5 - 215.56	Svalbard and Jan Mayen	Argosirenites nelgehensis, Argosirenites obrucevi (151217 151218) Argosirenites sp. (151223)
Lacian - Alaunian	221.5 - 212.0	China (Sichuan)	Dittmarites sp. (25402)
Lacian - Alaunian	221.5 - 212.0	Russian Federation	Pseudosirenites sp. (136158)
Alaunian	215.56 - 212.0	Iran	Mesodistichites sp., Mesodistichites tozeri, Mesodistichites evolutus (200273) Mesodistichites tozeri, Mesodistichites evolutus, Noridiscites nodosus (200277)
Alaunian	215.56 - 212.0	Canada (British Columbia)	Distichites canadensis (137100) Distichites gethingi (141706) Distichites gethingi, Leiodistichites ursidens (141680) Ectolcites childerhosei (141661) Leiodistichites ursidens (141682) Pleurodistichites hindei (137102) Pleurodistichites sp., Pseudosirenites pardoneti, Distichites gethingi (141696) Pseudosirenites bullatus (137039 139035 139036 139120 141748) Pseudosirenites falcatus (137988) Pseudosirenites pardoneti (46596 46598 46629 141699 141712 141725) Pseudosirenites pardoneti, Distichites canadensis, Leiodistichites ursidens (141713) Pseudosirenites pardoneti, Distichites columbianus, Distichites canadensis, Pleurodistichites stotti (141707) Pseudosirenites pardoneti, Distichites columbianus, Distichites gethingi (139137) Pseudosirenites pardoneti, Distichites gethingi (137037 141705 141729) Pseudosirenites pardoneti, Distichites gethingi, Distichites canadensis, Leiodistichites ursidens (46631) Pseudosirenites pardoneti, Distichites gethingi, Pleurodistichites stotti (138026) Pseudosirenites pardoneti, Ectolcites childerhosei (141733) Pseudosirenites pardoneti, Leiodistichites beachi (141724) Pseudosirenites pardoneti, Leiodistichites ursidens (141722) Pseudosirenites pardoneti, Pseudosirenites pressus, Distichites canadensis, Leiodistichites ursidens (141711) Pseudosirenites pardoneti, Pseudosirenites pressus, Distichites gethingi, Distichites canadensis (46630) Pseudosirenites pardoneti, Sirenites pressus (137030) Pseudosirenites sp. (47377 47378)
Alaunian	215.56 - 212.0	Nepal	Distichites harpalos, Trachypleuraspidites griffithi, Pleurodistichites trailli (92142)
Alaunian	215.56 - 212.0	Indonesia	Distichites falcatus (170397) Distichites falcatus, Ectolcites hollandi (220204) Distichites hacqueti, Distichites harpalos, Distichites megacanthus (220207) Distichites harpalos, Distichites megacanthus, Pseudosirenites evae, Pseudosirenites elegans, Distichites kmetyi, Distichites mesacanthus, Distichites leptacanthus (139412) Distichites harpalos, Pseudosirenites evae, Distichites kmetyi, Distichites sollasii, Paradistichites ectolcitiformis (170396) Distichites harpalos, Pseudosirenites evae, Distichites kmetyi, Trachypleuraspidites malayicus (220211) Distichites megacanthus (220214) Distichites megacanthus, Distichites tropicus, Paradistichites ectolcitiformis (151058) Distichites megacanthus, Pseudosirenites evae, Distichites kmetyi, Trachypleuraspidites malayicus (220210) Distichites tropicus (151066) Ectolcites pseudoaries, Ectolcites hochstetteri (151065) Ectolcites pseudoaries, Ectolcites hollandi (151064) Ectolcites pseudoaries, Pseudosirenites evae, Distichites falcatus, Ectolcites hollandi (151063) Paradistichites ectolcitiformis (151067) Pseudosirenites evae (220212) Pseudosirenites evae, Distichites kmetyi, Paradistichites ectolcitiformis, Ectolcites hollandi (220205) Pseudosirenites evae, Paradistichites ectolcitiformis (170398) Pseudosirenites sp. (220213)
Alaunian	215.56 - 212.0	Russian Federation	Pseudosirenites pardoneti (136160)
Alaunian	215.56 - 212.0	Armenia	Nairites armenius, Nairites laevis (144213)
Sevatian	212.0 - 205.6	Tajikistan	Dronovites pamiricus (151033)