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Camelus grattardi

Mammalia - Ungulata - Camelidae

Taxonomy
Camelus grattardi was named by Geraads (2014). Its type specimen is Omo 75S-70–956, a maxilla (maxilla fragment with heavily worn P4–M2), and it is a 3D body fossil. Its type locality is Omo - O.75s, which is in the Gelasian of sandstone the Shungura Formation Ethiopia.

Synonymy list
YearName and author
2014Camelus grattardi Geraads p. 1481 figs. 1, 3

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Synapsida()
Therapsida()
RankNameAuthor
infraorderCynodontia()
Epicynodontia
infraorderEucynodontia
Probainognathia
Mammaliamorpha
Mammaliaformes
classMammalia
subclassTribosphenida()
infraclassEutheria()
orderUngulata(Linnaeus 1766)
orderArtiodactyla()
suborderTylopoda()
superfamilyCameloidea
familyCamelidae
subfamilyCamelinaeGray 1821
tribeCameliniGray 1821
genusCamelus
speciesgrattardi

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
D. Geraads 2014A large member of Camelus, upper molars only slightly longer than broad, with rounded styles, P4 much nar- rower than M1, lingual wall of m3 hypoconulid not very oblique, mesial border of the mandibular ramus is strongly inclined back- wards, distal humeral articulation almost perpendicular to the long axis of the bone, and with a cylindrical medial part. Differs from species of Paracamelus in the loss of p3. Differs from mod- ern C. bactrianus and C. dromedarius in its much broader molars, both relative to their length and relative to P4, less acute styles in the upper teeth, less oblique m3 hypoconulid, more obli- que anterior border of mandibular ramus, and more cylindrical and more transverse distal humeral articulation. Differs from the Pleistocene North African C. thomasi in its much broader molars, both relative to their length and relative to P4, and lack of mandibular pachyostosis.