PBDB Taxon

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Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Gadiformes

Actinopteri - Gadiformes

Taxonomy

Gadiformes was named by Goodrich (1909). It is extant.

It was assigned to Osteichthyes by Purdy et al. (2001) and Sepkoski (2002); to Teleostei by Fierstine et al. (2012); to Gadariae by Betancur et al. (2013); to Paracanthopterygii by Nelson (2006), Long (2011) and Nelson et al. (2016); and to Teleosti by Carnevale and Godfrey (2018).

Synonymy list

Year	Name and author
1909	Gadiformes Goodrich
2001	Gadiformes Purdy et al. p. 166
2002	Gadiformes Sepkoski
2006	Gadiformes Nelson
2011	Gadiformes Long p. 243
2012	Gadiformes Fierstine et al. p. 60
2013	Gadiformes Betancur et al.
2016	Gadiformes Nelson et al. p. 293
2018	Gadiformes Carnevale and Godfrey p. 169

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
superclass	Actinopterygii	()
class	Actinopteri	()
subclass	Neopterygii	(Regan 1923)
—	Teleosteomorpha
—	Teleostei	(Müller 1846)

Rank	Name	Author
—	Osteoglossocephalai
—	Clupeocephala	Patterson and Rosen 1977
—	Euteleosteomorpha
—	Neoteleostei	Rosen 1973
—	Eurypterygia	(Rosen 1973)
—	Ctenosquamata
—	Acanthomorphata	(Rosen 1973)
—	Paracanthomorphacea
—	Zeiogadaria
—	Gadariae
order	Gadiformes	Goodrich 1909
order	Gadiformes	Goodrich 1909

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Gadiformes Goodrich 1909

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Fm. Bathygadidae Jordan and Evermann 1898

G. Gadomus Regan 1903

†Gadomus tejkali Brzobohaty and Schultz 1978

G. †Paleobathygadus David 1956

†Paleobathygadus careagrus David 1956

†Paleobathygadus oligocenicus David 1956

†Paleobathygadus wilmingtonensis David 1956

†Paleobathygadus yaguinensis David 1956

†Paleobathygadus zemorrianus David 1956

G. †Probathygadus David 1956

†Probathygadus keaseyensis David 1956

G. Brosme Rafinesque 1815

Fm. †Euclichthyidae Cohen 1984

G. †Archaemacruroides Stinton 1965

†Archaemacruroides ornatus Stinton 1965

G. Euclichthys McCulloch 1926

†Euclichthys eocenicus Schwarzhans 1980

†Euclichthys novaezeelandiae Frost 1924

Subor. Gadoidei

Fm. †Bregmacerotidae

G. †Bregmacerina Danil'chenko 1957

G. Bregmaceros Thompson 1840

†Bregmaceros albyi Sauvage 1880

†Bregmaceros antiquus Schwarzhans 1980

Bregmaceros cantori Milliken and Houde 1984 [striped codlet]

†Bregmaceros deklaszi Schwarzhans 2013

†Bregmaceros felkeri Schwarzhans 2007

†Bregmaceros filamentosus Priem 1908

†Bregmaceros hybridus Schwarzhans 2013

†Bregmaceros kreyenhagus David 1946

†Bregmaceros minutus Stinton 1957

†Bregmaceros prosoponus Grenfell 1984

†Bregmaceros troelli Frizzell and Dante 1965

Fm. Gadidae Rafinesque 1810

G. †Arnoldina Jordan and Gilbert 1919

†Arnoldina iniistia Jordan and Gilbert 1919

G. †Arnoldites Jordan 1923

G. Brosmius Cuvier 1829

†Brosmius andrussovi Bogachev 1933

G. Gadiculus Guichenot 1850

Subg. †Gadiculus (Archaeogadiculus) Schwarzhans 1978

†Gadiculus (Archaeogadiculus) labiatus Schubert 1905

Subg. Gadiculus (Gadiculus) Guichenot 1850

†Gadiculus (Gadiculus) antipodus Schwarzhans 1980

†Gadiculus (Gadiculus) argenteus communis Prochazka 1893

Subg. †Gadiculus (Microgadiculus) Schwarzhans 1978

†Gadiculus (Microgadiculus) rumanus Weiler 1943

Subg. Gadiculus (Paratrisopterus) Fedotov 1971

†Gadiculus (Paratrisopterus) avus Fedotov 1971

†Gadiculus adversus Schwarzhans 2019

Gadiculus argenteus Guichenot 1850 [silvery pout]

Gadiculus argenteus thori Schmidt 1913

†Gadiculus pereawa Schwarzhans 2019

G. †Gadidarum Koken 1885

†Gadidarum amsianus Schwarzhans 2007

Gadidarum ensiformis Steurbaut and Herman 1978

†Gadidarum insectus Weiler 1943

Subfm. †Gadinae

G. †Semeniolum Gaemers 1984

†Semeniolum repelense Gaemers 1984

G. Gadus Linnaeus 1758 [cod]

†Gadus benedeni Leriche 1926

†Gadus elegans Koken 1884

†Gadus elegans scuplta Koken 1891

†Gadus friedbergi Chaine and Duvergier 1928

†Gadus minisculoides Schubert 1912

†Gadus planatus Bassoli and Schubert 1906

†Gadus simplex Koken 1891

†Gadus spectabilis Koken 1891

†Gadus venustus Koken 1891

G. Gaidropsarus Rafinesque 1810

†Gaidropsarus murdjadjensis Arambourg 1927

†Gaidropsarus pilleri Carnevale and Harhauzer 2013

G. Melanogrammus Gill 1862

G. †Merlangiogadus Gaemers 1973

G. Merlangius Geoffroy 1767

†Merlangus cognatus Koken 1891

†Merlangus pseudaeglifinus Newton 1891

†Merlangus spatulus Koken 1891

†Merlangus spaulatus Koken 1942

†Merlangus spaulatus miocenicus Heinrich 1969

†Merlangus tenuis Heinrich 1969

Merlangus vulgaris Fleming 1828

†Merlangus vulgaris suffolkensis Koken 1891

G. Microgadus Gill 1865

Microgadus proximus Girard 1845

G. Micromesistius Gill 1863

†Micromesistius australis Hutton 1872

†Micromesistius boscheinei Schwarzhans 1978

Micromesistius poutassou Risso 1827 [blue whiting]

†Micromesistius schwarzhansi Holec 1975

G. Morrhua Oken 1817

†Morrhua faba Koken 1884

†Morrhua latus Koken 1891

†Morrhua sollingenensis Koken 1891

G. †Palaeomolva Daniltshenko 1947

†Palaeomolva tarchanica Daniltshenko 1947

G. †Palimphemus Kner 1862

†Palimphemus anceps Kner 1862

†Palimphemus minusculoides Schubert 1912

Invalid names: Micromesistius arcuatus Radwanska 1992 [synonym], Otolithus (Gadidarum) angustus Suzin 1968 [synonym], Otolithus (Gadidarum) minusculus Schubert 1906 [synonym]

†Palimphemus seymourensis Schwarzhans et al. 2017

Invalid names: Colliolus Gaemers and Schwarzhans 1973 [synonym], Doyenneichthys Gaemers 1983 [synonym], Gadichthys Gaemers and Schwarzhans 1973 [synonym], Gadus elegans sculptus Koken 1891 [synonym], Pseudocolliolus Gaemers 1978 [synonym]

G. †Progadus David 1956

†Progadus miocenicus David 1956

†Progadus pliocenicus David 1956

G. †Protocolliolus Gaemers 1976

†Protocolliolus amorphus Schwarzhans 2003

†Protocolliolus eocenicus Frost 1931

Subfm. Ranicipinae

G. Palaeoraniceps Gaemers 1976

G. Theragra Lucas 1898

Theragra chalcogramma Pallus 1814

G. Trisopterus Rafinesque 1814

Trisopterus luscus Linnaeus 1758

G. Urophycis Gill 1864

Invalid names: Phycinae [empty]

Fm. Lotidae Bonaparte 1842

G. †Gadomorpholithus Schwarzhans 2003

†Gadomorpholithus ponderosus Koken 1885

G. Molva Lesueur 1819

†Molva dubia Casier 1966

†Molva palaeomorpha Schwarzhans 2003

G. Phycis Walbaum 1792

†Phycis bartonensis Schubert 1916

†Phycis simplex Koken 1942

†Phycis simplex miocenica Weiler 1942

†Phycis simplex elongatus Posthumus 1958

†Phycis tenuis Koken 1891

Fm. Merlucciidae Gill 1884

G. †Macrurulus Schwarzhans 1980

†Macrurulus depressus Schwarzhans 2019

†Macrurulus fordycei Schwarzhans 2019

†Macrurulus immanis Schwarzhans 1980

G. †Merlucciidarum Gill 1884

†Merlucciidarum papillosus Casier 1966

G. Merluccius Rafinesque 1810

†Merluccius abreviatus Nolf and Brzobohaty 2002

†Merluccius athanasiui Pauca 1929

†Merluccius attenuatus Koken 1891

†Merluccius balticus Koken 1885

†Merluccius bilinearis Mitchill 1814

†Merluccius emarginatus Koken 1884

†Merluccius fimbriatus Stinton 1957

†Merluccius inferus Daniltshenko 1947

†Merluccius lednevi Bogatshov 1933

†Merluccius macroactus Kramberger 1879

Merluccius merluccius Linnaeus 1758 [European hake]

†Merluccius miocenicus Koken 1891

†Merluccius obtusus Koken 1891

†Merluccius ovalis Ciobanu 1977

Merluccius productus Ayres 1855

†Merluccius pukeuriensis Frost 1924

†Merluccius rattazzii Lin et al. 2017

†Merluccius romanicus Pauca 1929

†Merluccius vulgaris Fleming 1828

G. †Palaeogadus Von Rath 1859

Subg. †Palaeogadus (Lotimorpha) Danil'chenko 1950

†Palaeogadus (Lotimorpha) simionescui Daniltshenko 1950

†Palaeogadus abbreviatus Bogatshev 1933

†Palaeogadus atropatanus Bogachev 1933

†Palaeogadus compactus Gaemers and van Hinsbergh 1978

†Palaeogadus intergerinus Daniltshenko 1947

†Palaeogadus latebrosus Danil'chenko 1960

†Palaeogadus nodosus Stinton 1977

†Palaeogadus ruisbroekensis Gaemers 1984

†Palaeogadus serratus Casier 1966

†Palaeogadus sinangulatus Schwarzhans 2003

Invalid names: Nemopteryx Agassiz 1842 [synonym]

G. †Promerluccius David 1956

†Promerluccius venturaensis David 1956

G. †Rhinocephalus Casier 1966

Rhinocephalus planiceps Casier 1966

Invalid names: Merlucciinae Rafinesque 1815 [empty]

Fm. Moridae

G. †Actuariolum Karrer 1971

†Actuariolum otekaikensis Schwarzhans 1980

G. Auchenoceros Günther 1889

†Auchenoceros bicaudatus Frost 1924

Auchenoceros punctatum Hutton 1873

†Auchenoceros simplex Schwarzhans 2019

G. Austrophycis Ogilby 1897

†Austrophycis pseudomegalops Schwarzhans 1980

G. †Eophycis Jerzmanska 1968

†Eophycis jamnensis Jerzmanska 1968

G. †Fanteichthys Carnevale 2007

†Fanteichthys torricellensis Carnevale 2007

G. †Gadella Lowe 1843

G. Laemonema Guenther 1862

†Laemonema yarelli Lowe 1841

G. Lepidion Swainson 1838

Lepidion lepidion Risso 1810

G. †Lotella Kaup

†Lotella latidorsalis Schwarzhans et al. 2017

†Lotella matata Schwarzhans 2019

†Lotella pfeili Schwarzhans 2019

†Lotella rhacina Lotella rhacina 1801

†Lotella smirnovi Danilshenko 1953

†Lotella spicata Schwarzhans 2019

G. Mora Risso 1826

G. †Onobrosmius Bogatshov 1938

†Onobrosmius oligocaenicus Bogatshov 1933

G. Paralaemonema Trunov 1990

G. †Physiculinorum Unknown

†Physiculinorum vandenboschi Steurbaut 1984

G. Physiculus Kaup 1858

†Physiculus beui Schwarzhans 2019

†Physiculus huloti Poll 1953

Physiculus rastrelliger Gilbert 1890

†Physiculus terakohensis Frost 1934

G. †Pseudophycis Guenther 1862

Pseudophycis bachus Forster 1801

Pseudophycis barbata Günther 1863

Pseudophycis breviuscula Richardson 1846

Pseudophycis marginata Günther 1878

†Pseudophycis muringa Schwarzhans 2019

G. Tripterophycis Boulenger 1902

†Tripterophycis elongatissimus Schwarzhans 1985

†Tripterophycis immutatus Schwarzhans 1980

Fm. Phycidae

G. †Palaeomorrhua Gaemers and Schwarzhans 1973

†Palaeomorrhua bulbus Nolf 1978

†Palaeomorrhua thulei Schwarzhans 2004

G. †Protobrotula Danil'chenko 1960

†Protobrotula sobijevi Daniltshenko 1953

Invalid names: Ensigadus Gaemers 1978 [synonym]

G. †Symmetrosulcus Schwarzhans 1981

Fm. Gaidropsaridae Jordan and Evermann 1898

G. †Enchelyopus Agassiz 1835

Enchelyopus dorsoplicatus Gaemers 1984

†Enchelyopus tigrinus Agassiz 1835

G. †Onogadus Buen 1934

G. Lota Oken 1817

Subor. Macrouroidei

Fm. Macrouridae Jordan and Evermann 1898 [grenadiers]

G. †Amblygoniolepidus David 1956

†Amblygoniolepidus polystichus David 1956

G. Bathygadus Guenther 1878

†Bathygadus mandorovensis Schwarzhans 2013

†Bathygadus novus Bossoli 1906

†Bathygadus waiohaensis Schwarzhans 2019

G. †Bolbocara Jordan 1927

†Bolbocara gyrinus Jordan 1927

G. †Calilepidus David 1956

†Calilepidus tauxei David 1956

G. Coelorhynchus Agassiz 1843

†Coelorhynchus arthaberi Schubert 1905

Coelorhynchus australis Richardson 1839

†Coelorhynchus buanaiutoi Schwarzhans 1985

†Coelorhynchus burtinii Le Hon 1871

Coelorhynchus coelorhynchus Risso 1810

†Coelorhynchus cretaceus Dixon 1850

†Coelorhynchus cristasus Bassolini 1906

Coelorhynchus demonstratus Schwarzhans 1980

†Coelorhynchus elevatus Stinton 1956

†Coelorhynchus libanicus Woodward 1942

†Coelorhynchus pretoulai Schwarzhans 1985

†Coelorhynchus robustus Robba 1970

†Coelorhynchus toulai Schubert 1906

†Coelorhynchus yakujiensis Ohe and Araki 1973

G. Coelorinchus Giorna 1809

†Coelorinchus asperocephalus Waite 1911

†Coelorinchus balushkini Schwarzhans et al. 2017

Coelorinchus biclinozonalis Arai and McMillan 1982

Coelorinchus carminatus Goode 1880

Coelorinchus coelorhincus Risso 1810

†Coelorinchus divulgatus Schwarzhans 2019

Coelorinchus fasciatus Günther 1878

†Coelorinchus nordenskjoeldi Schwarzhans et al. 2017

Coelorinchus oliverianus Phillipps 1927

†Coelorinchus pakaurangiensis Schwarzhans 2019

†Coelorinchus preaustralis Schwarzhans 2019

Coelorinchus regularis Stinton 1957

†Coelorinchus robustus Robba 1970

†Coelorinchus stellaris Nolf and Girone 2000

G. Coryphaenoides Gunnerus 1765

†Coryphaenoides amager Schwarzhans 2003

†Coryphaenoides biobtusus Lin et al. 2017

†Coryphaenoides contortus Bassoli 1906

Coryphaenoides guentheri Vaillant 1888

†Coryphaenoides maximus Bassoi 1906

†Coryphaenoides rupestris Gunnerus 1765

G. †Homeocoryphaenoides David 1956

†Homeocoryphaenoides chiasmacanthus David 1956

G. †Homeomacrurus David 1956

†Homeomacrurus fernandensis David 1956

G. †Homeonezumia David 1956

†Homeonezumia multispinosus David 1956

G. Hymenocephalus Giglioli 1884

†Hymenocephalus globosus Posthumus 1923

†Hymenocephalus italicus Giglioli 1884

†Hymenocephalus medius Posthumus 1923

†Hymenocephalus rosenkrantzi Schwarzhans 2003

G. Lepidorhynchus Richardson 1846

Lepidorhynchus denticulatus Richardson 1846

†Lepidorhynchus frosti Schwarzhans 2019

G. †Leptacantholepidus David 1956

†Leptacantholepidus delmontianus David 1956

†Leptacantholepidus rotundus David 1956

Subfm. †Macrourinae Bonaparte 1832

G. †Morigadus Schwarzhans 1980

†Morigadus anarchicus Schwarzhans 1980

†Morigadus delicatulus Schwarzhans 1980

G. Ventrifossa Gilbert and Hubbs 1920

G. †Macrourus Bloch 1786

†Macrourus communis Prochazka 1893

†Macrourus debilis Posthumus 1923

†Macrurus altus Weiler 1943

†Macrurus dorsolobatus Weiler 1943

†Macrurus ellipticus Schubert 1905

†Macrurus gracilis Schubert 1905

†Macrurus obliquus Weiler 1950

†Macrurus praecursor Koken 1891

†Macrurus rhombicus Weiler 1943

†Macrurus rotundus Weiler 1943

†Macrurus toulai Schubert 1906

†Macrurus toulai cristasa Basolini 1906

†Macrurus trolli Bassoli 1906

G. Nezumia Jordan and Starks 1904

†Nezumia contemplata Schwarzhans 1978

†Nezumia contortus Bassoli 1906

†Nezumia lidsayi Schwarzhans 1985

†Nezumia mangapariensis Schwarzhans 1980

†Nezumia ornata Bassoli 1906

†Nezumia sagittiformis Frost 1934

Nezumia sclerorynchus Valenciennes 1838

†Nezumia teoulerensis Lin et al. 2017

†Otolithus (Macrurus) ornatus apicata Bassoli 1906

G. †Oxygoniolepidus David 1956

†Oxygoniolepidus washingtonensis David 1956

G. †Promacrurus David 1956

†Promacrurus alferitziensis David 1956

†Promacrurus alseanus David 1956

†Promacrurus asymmetricus David 1956

†Promacrurus mimus David 1956

†Promacrurus newmani David 1956

†Promacrurus oregonensis David 1956

†Promacrurus saucesianus David 1956

†Promacrurus zemorrianus David 1956

G. †Pyknolepidus David 1956

†Pyknolepidus macrospinosus David 1956

G. †Razelainia Carnevale 2007

†Razelainia paradoxa Carnevale 2007

G. Trachyrincus Giorna 1809

†Trachyrincus aulax Schwarzhans 2019

†Trachyrincus densus Stinton 1957

†Trachyrincus iocosus Lin et al. 2017

†Trachyrincus tewaewae Schwarzhans et al. 2017

Invalid names: Trachyrincinae Günther 1870 [empty]

Invalid names: Steindachneriidae [empty]

Fm. †Macruroididae

G. †Macruroides Smith and Radcliffe 1912

Fm. †Macruronidae Regan 1903

G. Macruronus Guenther 1873

†Macruronus eastmani Schwarzhans et al. 2017

†Macruronus merlucciiformis Schwarzhans 1980

Macruronus novaezeelandiae Hector 1871

†Macruronus pukeuriensis Frost 1924

G. †Maorigadus Schwarzhans 1980

†Maorigadus anarchicus Schwarzhans 1980

Invalid names: Maorigadus delicatulus Schwarzhans 1980 [synonym]

†Maorigadus ukrainicus Schwarzhans and Bratischko 2011

Fm. †Melanonidae

G. †Karrerichthys Schwarzhans 1980

†Karrerichthys admirabilis Schwarzhans 1980

†Karrerichthys latisulcatus Frost 1934

†Karrerichthys tenuis Stinton 1957

G. †Melanonidarum Günther

†Melanonidarum venheuckelomae Nolf and Steurbaut 1983

G. Melanonus Guenther 1878

Invalid names: Trachyrhynchinae [empty]

G. †Paratichthys

G. †Pollachius Nilsson 1832

G. †Pseudoraniceps Fedotov 1974

Fm. Ranicipitidae

G. Raniceps Oken 1817

†Raniceps hermani Nolf 1978

†Raniceps latidens Casier 1966

†Raniceps latisulcatus Koken 1884

†Raniceps parcus Danil'chenko 1960

†Raniceps planus Koken 1906

Raniceps raninus Linnaeus 1758

†Raniceps tuberculosus Koken 1884

G. †Rankinian David 1946

G. Squalogadus Gilbert and Hubbs 1916

G. Trachyrhynchus Bonaparte 1841

†Trachyrhynchus trachyrhynchus Risso 1810

G. †Trichurides

Invalid names: Merluccidae [empty], Muraenolepididae [empty]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

phosphaticsubp

Environment:

marinesubp

Locomotion:

actively mobileo

Life habit:

nektonico

Diet:

carnivoreo

Created:

2009-07-20 21:19:31

Modified:

2009-07-20 23:19:31

Source: o = order, subp = subphylum

References: Carroll 1988, Hendy et al. 2009

Age range

Maximum range based only on fossils: base of the Late/Upper Cenomanian to the top of the Holocene or 99.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 93.5 Ma

Collections (401 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Late/Upper Cenomanian	99.6 - 93.5	Lebanon	Coelorhynchus libanicus (221080 221233)
Danian	66.0 - 61.6	Greenland	Protocolliolus amorphus, Gadomorpholithus ponderosus, Archaemacruroides ornatus, Coelorinchus balticus, Palaeomorrhua thulei (22943)
Teurian	66.0 - 55.8	New Zealand (South Island)	Macrurulus sp. (219516)
Selandian	61.6 - 59.2	Denmark	Gadiformes indet., Palaeogadus sinangulatus, Coelorhynchus balticus, Hymenocephalus rosenkrantzi, Gadomorpholithus ponderosus (159794) Gadiformes indet., Palaeogadus sinangulatus, Molva palaeomorpha, Protocolliolus amorphus, Coelorhynchus balticus, Coryphaenoides amager, Gadomorpholithus ponderosus (159793) Gadiformes indet., Palaeogadus sinangulatus, Protocolliolus amorphus, Coelorhynchus balticus, Gadomorpholithus ponderosus (159795) Gadiformes indet., Raniceps hermani, Palaeogadus sinangulatus, Molva palaeomorpha, Protocolliolus amorphus, Coelorhynchus balticus, Coryphaenoides amager, Hymenocephalus rosenkrantzi, Gadomorpholithus ponderosus (159792)
Selandian - Thanetian	61.6 - 56.0	Ukraine	Maorigadus ukrainicus, Raniceps hermani, Merlucciidarum antiquus (159509)
Thanetian	59.2 - 56.0	Australia (South Australia)	Nezumia lindsayi (219145)
Ypresian	56.0 - 47.8	Antarctica	Macruronus eastmani, Coelorinchus balushkini, Coelorinchus nordenskjoeldi (100414)
Ypresian	56.0 - 47.8	United Kingdom	Rhinocephalus planiceps (160682)
Ypresian	56.0 - 47.8	United Kingdom (England)	Rhinocephalus planiceps (8050 13299) Rhinocephalus sp. (76601) Trichurides sp. (3260)
Ypresian	56.0 - 47.8	Antarctica (Seymour Island)	Palimphemus seymourensis (176084) Tripterophycis immutatus (176085)
Ypresian	56.0 - 47.8	Belgium	Gadidae indet. (219001 219008)
Waipawan - Mangaorapan	55.8 - 49.5	New Zealand (South Island)	Maorigadus anarchicus, Macrurulus immanis, Macrurulus fordycei (219497)
Early/Lower Eocene	55.8 - 48.6	United Kingdom (England)	Merlucciidae indet. (38949 38950) Microgadus subnotus (3262)
Early/Lower Eocene - Middle Eocene	55.8 - 37.71	United Kingdom (England)	Trichurides sagittidens, Trisopterus dimidiatus (3257) Trisopterus dimidiatus (3258)
Mangaorapan	53.0 - 49.5	New Zealand (South Island)	Maorigadus anarchicus, Macrurulus immanis, Lotella spicata (219499)
Mangaorapan - Heretaungan	53.0 - 46.2	New Zealand (South Island)	Maorigadus anarchicus, Macrurulus immanis, Lotella spicata (219500 219501)
Heretaungan	49.5 - 46.2	New Zealand (South Island)	Maorigadus anarchicus, Macrurulus immanis (219498)
Lutetian	47.8 - 41.3	France	Bregmaceros rappei (188958 188960 188961 188962) Bregmaceros rappei, Trachyrincus iocosus, Nezumia teoulerensis, Bathygadus novus (188957)
Lutetian	47.8 - 41.3	United Kingdom (England)	Hymenocephalus fimbriatus, Trisopterus dimidiatus, Merluccius nodosus (41998) Trisopterus dimidiatus (41888 41989 41992 41994 41996)
Lutetian	47.8 - 41.3	Germany (Lower Saxony,)	Raniceps sp., Palaeogadus nodosus, Merlucciidarum papillosus, Gadidarum amsianus, Bregmaceros felkeri, Hoplobrotuloides bartonensis (188954)