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Clareocercaria
Taxonomy
Clareocercaria was named by Prokop et al. (2017). It is extant.
It was assigned to Neoptera by Prokop et al. (2017).
It was assigned to Neoptera by Prokop et al. (2017).
Subtaxa
Synonymy list
| Year | Name and author |
|---|---|
| 2017 | Clareocercaria Prokop et al. p. 2 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. Clareocercaria Prokop et al. 2017
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Unr. Acercaria Börner 1904
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Superor. Holothysanoptera Engel et al. 2025
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Unr. †Betathysanoptera Engel 2025
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Fm. †Zoropsocidae Tillyard 1935
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G. †Mogsonus Vishnyakova 1981
G. †Tschekardus Vishnyakova 1981
G. †Zoropsocoides Vishnyakova 1981
G. †Zoropsocus Tillyard 1935
Unr. †Lophioneuroptera Engel et al. 2025
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Fm. †Iotacyphidae Shcherbakov et al. 2024
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G. †Burmacypha Zherikhin 2000
G. †Iotacypha Shcherbakov et al. 2024
G. †Retiptera Cumming et al. 2024
Subfm. †Triassocyphinae Boderau et al. 2025
Fm. †Lophioneuridae Tillyard 1921
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G. †Austrocypha Tillyard 1935
G. †Cyphoneura Carpenter 1932
G. †Cyphoneurodes Becker-Migdisova 1953
G. †Jantardachus Vishnyakova 1981
G. †Karataocypha Vishnyakova 1981
G. †Liophioneura Tillyard 1921
G. †Lophiocypha Tillyard 1935
G. †Lophioneura Tillyard 1921
G. †Lophioneurodes Becker-Migdisova 1953
G. †Lophiosina Nel et al. 2014
G. †Undacypha Vishnyakova 1981
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Invalid names: Beipiaopsocus Hong 1992 [synonym], Edgariekia Jell and Duncan 1986 [synonym], Mesopsocus Hong 1983 [synonym], Undensis Vishnyakova 1981 [synonym]
G. †Vitriala Becker-Migdisova 1961
Invalid names: Edgariekiidae Jell and Duncan 1986 [synonym], Lophioneurinae Tillyard 1921 [empty]
Invalid names: Lophioneurida Tillyard 1921 [replaced]
Fm. †Westphalothripidesidae Nel et al. 2012
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Invalid names: Thripida Nel et al. 2012 [replaced]
Superor. Pancondylognatha Yoshizawa and Leinhard 2016
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Unr. Condylognatha Börner 1904
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Or. Hemiptera Linnaeus 1758 [true bug]
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Subor. Auchenorrhyncha Duméril 1805 [true bug]
G. Belostomum Latreille 1807 [giant water bug]
G. †Cercopidium Westwood 1854
G. †Cercopyllis Scudder 1890
Fm. †Cicadopsyllidae Martynov 1931
Subor. Coleorrhyncha Myers and China 1929
Subor. Heteroptera Latreille 1810
Subor. Homoptera Leach 1815
Fm. Hydrocorisae Latreille 1802
G. †Hypocixius Cockerell 1926
G. †Ludibrium Becker-Migdisova 1962
G. †Meshemipteron Cockerell 1915
G. †Mesococcus Becker-Migdisova 1959
Subor. †Paleorrhyncha Carpenter 1931
G. †Planophlebia Scudder 1879
G. †Shaandongia Hong 1985
Subor. Sternorrhyncha Amyot and Audinet-Serville 1843
Invalid names: Aphidii Latreille 1802 [empty], Auchenorinques Duméril 1805 [empty], Biturritidae Evans 1953 [empty], Cimicides Latreille 1802 [empty], Delphax rhenana Statz 1950 [nomen dubium], Delphax senilis Scudder 1877 [nomen dubium], Eofulgoridium chanmaense Hong 1982 [nomen dubium], Gallinsecta Latreille 1802 [empty], Heteropterodea Zrzavy 1992 [empty], Limnochares antiquus von Heyden 1862 [nomen dubium], Lithopsis lineatus Piton 1940 [nomen dubium], Lithopsis punctinervis Piton 1940 [nomen dubium], Lystra vollenhoveni Weyenbergh 1869 [nomen dubium], Palaeohemiptera Handlirsch 1906 [empty], Peloridiina Popov 1982 [empty], Pincombella Chapman 1932 [nomen dubium], Pincombella belmontensis Chapman 1932 [nomen dubium], Proboscidea Scopoli 1763 [synonym], Protocoris ovalis Heer 1864 [nomen nudum], Ryngota Fabricius 1775 [synonym]
Or. †Permopsocida Tillyard 1926
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Fm. †Archipsyllidae Handlirsch 1906
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G. †Archipsylla Handlirsch 1906
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†Archipsylla lata Vishnyakova 1976
†Archipsylla primitiva Handlirsch 1906
†Archipsylla similis Vishnyakova 1976
†Archipsylla sinica Huang et al. 2008
†Archipsylla turanica Martynov 1926
G. †Mydiognathus Yoshizawa and Leinhard 2016
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†Mydiognathus eviohlhoffae Yoshizawa and Leinhard 2016
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Invalid names: Psocorrhyncha burmitica Huang et al. 2016 [synonym]
Invalid names: Psocorrhyncha Huang et al. 2016 [synonym]
Fm. †Permopsocidae Tillyard 1926
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G. †Permopsocus Tillyard 1926
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†Permopsocus latipennis Tillyard 1926
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Invalid names: Ancylopsocus insolitus Tillyard 1926 [synonym], Permopsocus congener Tillyard 1926 [synonym], Permopsocus enderleini Tillyard 1926 [synonym]
†Permopsocus ovatus Carpenter 1939
Invalid names: Ancylopsocus Tillyard 1926 [synonym]
G. †Progonopsocus Tillyard 1926
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†Progonopsocus permianus Tillyard 1926
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Invalid names: Progonopsocus pusillus Tillyard 1926 [synonym]
Fm. †Psocidiidae Tillyard 1926
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G. †Austropsocidium Tillyard 1935
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†Austropsocidium pincombei Tillyard 1935
†Austropsocidium stigmaticum Tillyard 1935
G. †Dichentomum Tillyard 1926
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†Dichentomum arroyo Rasnitsyn 2004
†Dichentomum complexum Carpenter 1926
†Dichentomum grande Carpenter 1933
†Dichentomum latum Carpenter 1932
†Dichentomum minimum Tillyard 1926
†Dichentomum parvulum Carpenter 1932
†Dichentomum tinctum Tillyard 1926
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Invalid names: Chaetopsocidium sellardsi Tillyard 1926 [synonym], Metapsocidium loxoneurum Tillyard 1926 [synonym], Pentapsocidium indistinctum Tillyard 1926 [synonym], Permentomum tenuiforme Tillyard 1926 [synonym], Psocidium abnormale Tillyard 1926 [synonym], Psocidium kansasense Tillyard 1926 [synonym], Psocidium permianum Tillyard 1926 [synonym], Psocidium robustum Tillyard 1926 [synonym]
Invalid names: Chaetopsocidium Tillyard 1926 [synonym], Metapsocidium Tillyard 1926 [synonym], Pentapsocidium Tillyard 1926 [synonym], Permentomum Tillyard 1926 [synonym], Psocidium Tillyard 1926 [synonym]
Invalid names: Dichentomidae Carpenter 1932 [synonym]
Fm. †Westphalopsocidae Azar et al. 2013
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Or. †Hypoperlida Martynov 1928
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Fm. †Hypoperlidae Martynov 1928
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G. †Boreopsocus Shcherbakov 1994
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†Boreopsocus danksae Shcherbakov 1994
†Boreopsocus ficticius Novokshonov 1995
Invalid names: Psocivena Novokshonov 1995 [synonym]
G. †Hypoperla Martynov 1928
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†Hypoperla elegans Martynov 1928
†Hypoperla grata Novokshonova 1998
†Hypoperla nobilis Novokshonov 1995
†Hypoperla vaulevi Novokshonov 2001
G. †Idelopsocus Zalessky 1929
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†Idelopsocus arcuatus Martynov 1928
†Idelopsocus diradiatus Rasnitsyn 1996
†Idelopsocus galinae Novokshonov 2001
†Idelopsocus incommendatus Novokshonov et al. 2002
†Idelopsocus levis Novokshonov 1995
†Idelopsocus mutovinus Rasnitsyn and Aristov 2013
†Idelopsocus splendens Zalessky 1948
†Idelopsocus tataricus Zalessky 1929
Invalid names: Hypoperlopsis Zalessky 1948 [synonym], Martynopsocus Karny 1930 [synonym]
Invalid names: Farianopteridae Martynova 1961 [synonym], Kaltanelmoidae Rohdendorf 1961 [synonym], Martynopsocidae Karny 1930 [synonym]
Fm. †Sojanoperidae Novokshonov 2002
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Invalid names: Eukulojina Sinitshenkova 1981 [empty], Hypoperlina [invalid subgroup], Strephocladina [invalid subgroup]
Or. †Miomoptera Martynov 1927
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Fm. †Palaeomanteidae Handlirsch 1906
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G. †Delopsocus Tillyard 1928
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†Delopsocus elongatus Sellards 1909
†Delopsocus fasciatus Tillyard 1928
†Delopsocus furcatulus Martynov 1930
†Delopsocus kamensis Martynov 1938
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Invalid names: Delopterum kamense Carpenter 1939 [objective synonym]
†Delopsocus kansanum Carpenter 1939
†Delopsocus latus Sellards 1909
†Delopsocus lepidus Kukalová 1963
†Delopsocus sinuosus Kukalová 1963
†Delopsocus stenopterus Rasnitsyn 2004
G. †Delopterinus Rasnitsyn 2004
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†Delopterinus bicolor Rasnitsyn 2004
†Delopterinus zebra Rasnitsyn 2004
G. †Delopterum Sellards 1909
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†Delopterum anale Martynov 1928
†Delopterum candidum Zhuzhgova 2002
†Delopterum commune Rasnitsyn 2004
†Delopterum hangardi Guthorl 1950
†Delopterum iljinskiense Martynova 1961
†Delopterum incertum Martynov 1928
†Delopterum insigne Martynov 1928
†Delopterum kaltanicum Martynova 1961
†Delopterum pantherinum Rasnitsyn 2004
†Delopterum radtshenkoi Martynova 1961
†Delopterum rasnitsyni Novokshonov 2000
†Delopterum truncatum Kukalová 1963
†Delopterum zonatum Rasnitsyn 2004
Invalid names: Leptoneurula Martynov 1928 [synonym], Pseudomantis Martynov 1928 [synonym]
G. †Miomatoneura Martynov 1927
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†Miomatoneura angusta Kukalová 1963
†Miomatoneura candida Kukalová 1963
†Miomatoneura frigida Martynov 1927
†Miomatoneura intermedia Martynov 1928
†Miomatoneura palaeozoica Kukalová 1963
†Miomatoneura parvula Martynov 1930
†Miomatoneura permica Kukalová 1963
†Miomatoneura rotundata Martynov 1928
G. †Miomatoneurella Martynova 1958
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†Miomatoneurella americana Rasnitsyn 2004
†Miomatoneurella boskovicensis Kukalová 1963
†Miomatoneurella reducta Martynova 1958
†Miomatoneurella rossoshana Rasnitsyn and Aristov 2013
G. †Palaeomantis Handlirsch 1904
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†Palaeomantis aestiva Novokshonov 2000
†Palaeomantis apicalis Rasnitsyn 2004
†Palaeomantis hangardi Guthörl 1962
†Palaeomantis laeta Novokshonov and Zhuzhgova 2002
†Palaeomantis ostertali Guthörl 1962
†Palaeomantis schmidti Handlirsch 1904
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Invalid names: Palaeomantis aequalis Martynov 1928 [synonym]
†Palaeomantis sylvaensis Martynov 1940
†Palaeomantis tshekardensis Novokshonov 2000
Invalid names: Stefanomioptera Guthörl 1962 [synonym]
G. †Permonia Kukalová 1963
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†Permonia arta Kukalová 1963
†Permonia insolida Kukalová 1963
†Permonia permoni Kukalová 1963
G. †Perunopterum Kukalová 1963
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†Perunopterum corium Kukalová 1963
†Perunopterum lepidum Kukalová 1963
†Perunopterum lumbiforme Kukalová 1963
†Perunopterum macrum Kukalová 1963
†Perunopterum mirum Kukalová 1963
†Perunopterum peruni Kukalová 1963
†Perunopterum saxeum Kukalová 1963
G. †Sellardsiopsis Zalessky 1939
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†Sellardsiopsis conspicua Zalessky 1939
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Invalid names: Miomantisca clara Zalessky 1956 [synonym], Miomoptilon pusillus Zalessky 1956 [synonym], Palaeomantisca lata Martynov 1940 [synonym], Paramioma palida Zalessky 1956 [synonym]
Invalid names: Miomantisca Zalessky 1956 [synonym], Miomoptilon Zalessky 1956 [synonym], Palaeomantisca Martynov 1940 [synonym], Paramioma Zalessky 1956 [synonym]
G. †Tridelopterum Rasnitsyn 2004
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†Tridelopterum indebitum Rasnitsyn and Aristov 2013
†Tridelopterum variegatum Rasnitsyn 2004
Invalid names: Archisialidae Martynov 1932 [synonym], Delopteridae Sellards 1909 [synonym], Palaeomantiscidae Rasnitsyn 1977 [synonym], Stefanomiopterinae Guthörl 1962 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Kiessling 2004, Bush and Bambach 2015 | |||||
Age range
Maximum range based only on fossils: base of the Marsdenian to the top of the Holocene or 321.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 320.0 Ma
Minimum age of oldest fossil (stem group age): 320.0 Ma
Collections (1449 total)
Oldest occurrences | |||
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Marsdenian | Belgium | Protoprosbole straeleni (164479) | |
| Westphalian C - Westphalian D | France | Westphalopsocus pumilio, Carbopsyllidium minutum, Aviorrhyncha magnifica, Carbonatura oudardi, Westphalothripides oudardi (122504) | |
| Westphalian D | USA (Illinois) | Mazonopsocus testai (188251) | |
| Westphalian D | Germany (Lower Saxony) | Carbonopsocus mercuryi (224159) | |
| Stephanian A | Germany | Saaromioptera jordani, Stefanomioptera hangardi, Stefanomioptera ostertali (123835) | |
| Gzhelian | Russian Federation | Archeglyphis crassinervis (124589) Blattoprosbole tomiensis (124585) | |
| Asselian | USA (New Mexico) | Arroyoscyta novaemexicana, Dichentomum arroyo, Delopterum commune, Delopterum pantherinum, Delopterum zonatum, Stigmodelopterum pterostigmalis, Delopterinus bicolor, Delopterinus zebra, Miomatoneurella americana, Tridelopterum variegatum, Palaeomantis apicalis, Delopsocus stenopterus (124386) Delopterum sp., Delopterum commune, Neodelopterum anomalum (124385) Dichentomum arroyo (122481) Dichentomum sp., Palaeomanteidae indet., Delopterinus bicolor (124387) | |
| Asselian | Czechia | Delopterum sp., Delopterum truncatum, Miomatoneura permica, Miomatoneura candida, Miomatoneura angusta, Permonia arta, Permonia insolida, Permonia permoni, Permonikia permoniki, Miomatoneurella boskovicensis, Permodelopterum obscurum, Delopterum sinuosum, Delopterum lepidum, Perunopterum sp., Perunopterum peruni, Perunopterum corium, Perunopterum macrum, Perunopterum saxeum, Perunopterum mirum, Permodelopterum lepidum, Permodelopterum lumbiformis (114186) Miomatoneura palaeozoica (114187) Perunopterum peruni (114188) | |
| Asselian | Germany (Rhineland-Palatinate) | Permoridium fresenaci (156248) | |
| Asselian | USA (Colorado) | Cercopyllis justiciae, Cercopyllis adolescens, Cercopyllis delicatula (106345) | |
| Permian | China (Nei Mongol) | Palaeocicadopsis chinensis (156333) | |
| Artinskian | USA (Oklahoma) | Cyphoneurodes patriciae (104818) Delopterum latum (104816) Permopsylla americana, Parapermopsylla tricubita, Dichentomum tinctum, Delopterum minutum, Delopterum latum, Delopterum elongatum (104826) | |
| Artinskian | USA (Kansas) | Dichentomum complexum (151412) Dichentomum tinctum, Delopterum minutum, Urba punctata, Delopterum latum, Delopterum elongatum (21979) Elmomantis engeli (114450) Permopsylla americana, Progonopsocus permianus, Permopsocus latipennis, Dichentomum tinctum, Delopterum minutum, Delopterum latum, Delopterum elongatum (111320) Permopsylla minuta, Permopsylla americana, Permopsocus latipennis, Delopterum latum (114152) Permopsylla sp., Permopsylla grandis, Permopsylla americana, Permopsylla anomala, Lithoscytina cubitalis, Progonopsocus permianus, Permopsocus latipennis, Lithopsocidium permianum, Dichentomum tinctum, Dichentomum grande, Delopterum minutum, Delopterum elongatum (111319) Permoscytina kansasensis, Archescytina permiana, Permopsylla grandis, Permopsylla minuta, Permopsylla permiana, Permopsylla americana, Permopsocus latipennis, Permopsocus ovatus, Lithopsocidium permianum, Dichentomum tinctum, Delopterum minutum, Delopterum kansanum, Delopterum elongatum (114151) Permoscytina muiri, Permoscytina kansasensis, Archescytina permiana, Permopsylla americana, Progonopsocus permianus, Progonopsocus pusillus, Permopsocus latipennis, Permopsocus enderleini, Permopsocus congener, Ancylopsocus insolitus, Psocidium sp., Dichentomum tinctum, Psocidium permianum, Metapsocidium loxoneurum, Psocidium robustum, Chaetopsocidium sellardsi, Pentapsocidium indistinctum, Psocidium kansasense, Permentomum tenuiforme, Psocidium abnormale, Psocidium minimum, Delopterum minutum, Delopterum dunbari, Delopteridae indet., Delopsocus latus, Delopsocus fasciatus, Delopsocus elongatus (111219) Permoscytina muiri, Permoscytina kansasensis, Archescytina permiana, Permopsylla minuta, Permopsylla permiana, Permopsylla americana, Paleoscytina brevistigma, Cyphoneura permiana, Cyphoneura reducta, Progonopsocus permianus, Orthopsocus singularis, Permopsocus latipennis, Lithopsocidium permianum, Dichentomum latum, Dichentomum tinctum, Dichentomum minimum, Dichentomum parvulum (111217) | |
| Artinskian | Uruguay | Paracicadopsis mendezalzolai (124521) | |
| Artinskian | France (Languedoc) | Lutevanaphis permiana, Palaeomantis sp. (105044) Permopsyllidium lesclansis (122883) | |
| Artinskian - Wordian | France (Languedoc) | Kaltanetta gonfaronensis, Tomioscarta dominiquea (192423) | |
| Kungurian | Russian Federation | Cicadopsis sp. (115214) Idelopsocus galinae (124284) Idelopsocus splendens, Hypoperla vaulevi, Delopterum candidum, Palaeomantis tshekardensis, Palaeomantis aestiva (107787) Maueria pusillus (196229) Maueria rhynchota (155821) Palaeomantis aestiva (114207) Permopsyllopsis maueriaeformis, Parapsocidium uralicum, Sellardsiopsis conspicua (196228) Scytoneurella major, Scytoneurella minor, Maripsocus ambiguus, Maueria sylvensis, Maueria pusillus, Maueria intermedia, Maueria rhynchota, Uraloscytina prosbolioides, Permoscytinopsis maueriaeformis, Permopsyllopsis rossica, Parapsocidium uralicum, Hypoperlopsis splendens, Sellardsiopsis conspicua, Paramioma palida, Miomantisca clara, Miomoptilon pusillus, Miomatoneurites sylvaensis (102504) Uraloscytina multinervosa (152060) Uraloscytina similis, Parapsocidium uralicum (155819) | |
| Kungurian | Russian Federation (Kemerovo) | Prosbole sp., Sarbaloscytina angustipennis, Kaltanoscytina latistigmata, Sarbaloptera sarbalensis, Scytinoptera sibirica, Lophioneurodes sarbalensis (112708) Sarbaloptera sarbalensis (104951) | |
| Kungurian | Brazil (Rio Grande do Sul) | Gondwanaptera capsii, Prosbolecicada gondwanica, Prosbole iratiensis, Fulgoringruo kukalovae (116121) | |
| Kungurian | Russian Federation (Suksun) | Archescytinidae indet., Archescytina tshekardaensis, Palaeomantisca lata (116641) Dinoscyta microcephala, Tschekardus hispidus, Dichentomum sp., Idelopsocus diradiatus, Hypoperla grata, Delopterum rasnitsyni, Palaeomantina pentamera, Palaeomantis laeta, Palaeomantis aestiva (108206) Palaeomantis sylvaensis (113379) Tshekardaella tshekardaensis (124241) | |