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Clareocercaria

Insecta

Taxonomy
Clareocercaria was named by Prokop et al. (2017). It is extant.

It was assigned to Neoptera by Prokop et al. (2017).

Sister taxa
Acercaria

Synonymy list
YearName and author
2017Clareocercaria Prokop et al. p. 2

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
RankNameAuthor
classInsecta
Dicondylia
Paranotalia
subclassPterygota()
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
Clareocercaria
Clareocercaria

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Clareocercaria Prokop et al. 2017
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Unr. Acercaria Börner 1904
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Superor. Pancondylognatha Yoshizawa and Leinhard 2016
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Unr. Condylognatha Börner 1904
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Or. Hemiptera Linnaeus 1758 [true bug]
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Subor. Auchenorrhyncha Duméril 1805 [true bug]
G. Belostomum Latreille 1807 [giant water bug]
G. †Cercopyllis Scudder 1890
Fm. †Cicadopsyllidae Martynov 1931
Subor. Coleorrhyncha Myers and China 1929
Unr. Hemelytrata Fallen 1829
Subor. Heteroptera Latreille 1810
Subor. Homoptera Leach 1815
Fm. Hydrocorisae Latreille 1802
G. †Hypocixius Cockerell 1926
G. †Ludibrium Becker-Migdisova 1962
G. †Meshemipteron Cockerell 1915
G. †Mesococcus Becker-Migdisova 1959
Subor. †Paleorrhyncha Carpenter 1931
G. †Planophlebia Scudder 1879
G. †Shaandongia Hong 1985
Subor. Sternorrhyncha Amyot and Audinet-Serville 1843
Invalid names: Aphidii Latreille 1802 [empty], Auchenorinques Duméril 1805 [empty], Biturritidae Evans 1953 [empty], Cimicides Latreille 1802 [empty], Delphax rhenana Statz 1950 [nomen dubium], Delphax senilis Scudder 1877 [nomen dubium], Eofulgoridium chanmaense Hong 1982 [nomen dubium], Gallinsecta Latreille 1802 [empty], Heteropterodea Zrzavy 1992 [empty], Limnochares antiquus von Heyden 1862 [nomen dubium], Lithopsis lineatus Piton 1940 [nomen dubium], Lithopsis punctinervis Piton 1940 [nomen dubium], Lystra vollenhoveni Weyenbergh 1869 [nomen dubium], Palaeohemiptera Handlirsch 1906 [empty], Peloridiina Popov 1982 [empty], Pincombella Chapman 1932 [nomen dubium], Pincombella belmontensis Chapman 1932 [nomen dubium], Proboscidea Scopoli 1763 [synonym], Protocoris ovalis Heer 1864 [nomen nudum], Ryngota Fabricius 1775 [synonym]
Superor. Thripida Fallen 1814
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Or. †Lophioneurida Tillyard 1921
Unr. †Panthysanoptera Nel et al. 2012
Or. Thysanoptera Haliday 1836 [thrips]
Invalid names: Cyphoneuridae Carpenter 1932 [empty]
Or. †Permopsocida Tillyard 1926
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Fm. †Archipsyllidae Handlirsch 1906
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G. †Archipsylla Handlirsch 1906
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Archipsylla lata Vishnyakova 1976
Archipsylla primitiva Handlirsch 1906
Archipsylla similis Vishnyakova 1976
Archipsylla sinica Huang et al. 2008
Archipsylla turanica Martynov 1926
G. †Archipsyllodes Vishnyakova 1976
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Archipsyllodes speciosus Vishnyakova 1976
G. †Archipsyllopsis Vishnyakova 1976
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Archipsyllopsis baissica Vishnyakova 1976
G. †Bittacopsocus Beutel et al. 2019
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Bittacopsocus megacephalus Beutel et al. 2019
G. †Burmopsylla Liang et al. 2016
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Burmopsylla maculata Liang et al. 2016
G. †Dinmopsylla Lambkin 2018
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Dinmopsylla semota Lambkin 2018
G. †Eopsylla Vishnyakova 1976
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Eopsylla sojanense Becker-Migdisova 1962
G. †Mydiognathus Yoshizawa and Leinhard 2016
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Mydiognathus eviohlhoffae Yoshizawa and Leinhard 2016
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Invalid names: Psocorrhyncha burmitica Huang et al. 2016 [synonym]
Invalid names: Psocorrhyncha Huang et al. 2016 [synonym]
Fm. †Permopsocidae Tillyard 1926
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G. †Lithopsocidium Carpenter 1932
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Lithopsocidium permianum Carpenter 1932
G. †Orthopsocus Carpenter 1932
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Orthopsocus singularis Carpenter 1932
G. †Permopsocus Tillyard 1926
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Permopsocus latipennis Tillyard 1926
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Invalid names: Ancylopsocus insolitus Tillyard 1926 [synonym], Permopsocus congener Tillyard 1926 [synonym], Permopsocus enderleini Tillyard 1926 [synonym]
Permopsocus ovatus Carpenter 1939
Invalid names: Ancylopsocus Tillyard 1926 [synonym]
G. †Progonopsocus Tillyard 1926
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Progonopsocus permianus Tillyard 1926
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Invalid names: Progonopsocus pusillus Tillyard 1926 [synonym]
Fm. †Psocidiidae Tillyard 1926
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G. †Austropsocidium Tillyard 1935
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Austropsocidium pincombei Tillyard 1935
Austropsocidium stigmaticum Tillyard 1935
G. †Dichentomum Tillyard 1926
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Dichentomum arroyo Rasnitsyn 2004
Dichentomum complexum Carpenter 1926
Dichentomum grande Carpenter 1933
Dichentomum latum Carpenter 1932
Dichentomum minimum Tillyard 1926
Dichentomum parvulum Carpenter 1932
Dichentomum tinctum Tillyard 1926
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Invalid names: Chaetopsocidium sellardsi Tillyard 1926 [synonym], Metapsocidium loxoneurum Tillyard 1926 [synonym], Pentapsocidium indistinctum Tillyard 1926 [synonym], Permentomum tenuiforme Tillyard 1926 [synonym], Psocidium abnormale Tillyard 1926 [synonym], Psocidium kansasense Tillyard 1926 [synonym], Psocidium permianum Tillyard 1926 [synonym], Psocidium robustum Tillyard 1926 [synonym]
Invalid names: Chaetopsocidium Tillyard 1926 [synonym], Metapsocidium Tillyard 1926 [synonym], Pentapsocidium Tillyard 1926 [synonym], Permentomum Tillyard 1926 [synonym], Psocidium Tillyard 1926 [synonym]
G. †Hypopsylla Prokop et al. 2016
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Hypopsylla belmontensis Prokop et al. 2016
G. †Liassopsocus Ansorge 1996
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Liassopsocus lanceolatus Ansorge 1996
G. †Megapsocidium Tillyard 1935
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Megapsocidium australe Tillyard 1935
G. †Parapsocidium Zalessky 1937
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Parapsocidium uralicum Zalessky 1937
G. †Stenopsocidium Tillyard 1935
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Stenopsocidium elongatum Tillyard 1935
Invalid names: Dichentomidae Carpenter 1932 [synonym]
Fm. †Westphalopsocidae Azar et al. 2013
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G. †Westphalopsocus Azar et al. 2013
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Westphalopsocus pumilio Azar et al. 2013
Or. †Hypoperlida Martynov 1928
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Fm. †Hypoperlidae Martynov 1928
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G. †Boreopsocus Shcherbakov 1994
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Boreopsocus danksae Shcherbakov 1994
Boreopsocus ficticius Novokshonov 1995
Invalid names: Psocivena Novokshonov 1995 [synonym]
G. †Hypoperla Martynov 1928
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Hypoperla elegans Martynov 1928
Hypoperla grata Novokshonova 1998
Hypoperla nobilis Novokshonov 1995
Hypoperla vaulevi Novokshonov 2001
G. †Idelopsocus Zalessky 1929
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Idelopsocus arcuatus Martynov 1928
Idelopsocus diradiatus Rasnitsyn 1996
Idelopsocus galinae Novokshonov 2001
Idelopsocus incommendatus Novokshonov et al. 2002
Idelopsocus levis Novokshonov 1995
Idelopsocus mutovinus Rasnitsyn and Aristov 2013
Idelopsocus splendens Zalessky 1948
Idelopsocus tataricus Zalessky 1929
Invalid names: Hypoperlopsis Zalessky 1948 [synonym], Martynopsocus Karny 1930 [synonym]
G. †Kaltanelmoa Rohdendorf 1961
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Kaltanelmoa sibirica Rohdendorf 1961
G. †Onthomastax Rasnitsyn and Aristov 2013
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Onthomastax coprinus Rasnitsyn and Aristov 2013
Invalid names: Farianopteridae Martynova 1961 [synonym], Kaltanelmoidae Rohdendorf 1961 [synonym], Martynopsocidae Karny 1930 [synonym]
Fm. †Sojanoperidae Novokshonov 2002
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G. †Sojanopus Novokshonov 2002
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Sojanopus festivum Novokshonov 2002
Invalid names: Eukulojina Sinitshenkova 1981 [empty], Hypoperlina [invalid subgroup], Strephocladina [invalid subgroup]
Or. †Miomoptera Martynov 1927
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G. †Miomina Martins-Neto and Gallego 1999
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Miomina mendozina Martins-Neto and Gallego 1999
Miomina riojana Martins-Neto and Gallego 2006
Fm. †Palaeomanteidae Handlirsch 1906
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G. †Archisialis Martynov 1932
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Archisialis praecox Martynov 1932
G. †Balymotikha Rasnitsyn and Aristov 2013
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Balymotikha deterior Rasnitsyn and Aristov 2013
G. †Belmomantis Prokop et al. 2017
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Belmomantis azari Prokop et al. 2017
G. †Delopsocus Tillyard 1928
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Delopsocus elongatus Sellards 1909
Delopsocus fasciatus Tillyard 1928
Delopsocus furcatulus Martynov 1930
Delopsocus kamensis Martynov 1938
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Invalid names: Delopterum kamense Carpenter 1939 [objective synonym]
Delopsocus kansanum Carpenter 1939
Delopsocus latus Sellards 1909
Delopsocus lepidus Kukalová 1963
Delopsocus sinuosus Kukalová 1963
Delopsocus stenopterus Rasnitsyn 2004
G. †Delopteriella Zalessky 1956
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Delopteriella graciosa Zalessky 1956
G. †Delopterinus Rasnitsyn 2004
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Delopterinus bicolor Rasnitsyn 2004
Delopterinus zebra Rasnitsyn 2004
G. †Delopterum Sellards 1909
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Delopterum anale Martynov 1928
Delopterum candidum Zhuzhgova 2002
Delopterum commune Rasnitsyn 2004
Delopterum hangardi Guthorl 1950
Delopterum iljinskiense Martynova 1961
Delopterum incertum Martynov 1928
Delopterum insigne Martynov 1928
Delopterum kaltanicum Martynova 1961
Delopterum minutum Sellards 1909
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Invalid names: Delopterum dunbari Tillyard 1928 [synonym]
Delopterum pantherinum Rasnitsyn 2004
Delopterum radtshenkoi Martynova 1961
Delopterum rasnitsyni Novokshonov 2000
Delopterum truncatum Kukalová 1963
Delopterum zonatum Rasnitsyn 2004
Invalid names: Leptoneurula Martynov 1928 [synonym], Pseudomantis Martynov 1928 [synonym]
G. †Elmomantis Prokop et al. 2017
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Elmomantis engeli Prokop et al. 2017
G. †Mazonopsocus Prokop et al. 2017
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Mazonopsocus testai Prokop et al. 2017
G. †Miomatoneura Martynov 1927
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Miomatoneura angusta Kukalová 1963
Miomatoneura candida Kukalová 1963
Miomatoneura frigida Martynov 1927
Miomatoneura intermedia Martynov 1928
Miomatoneura palaeozoica Kukalová 1963
Miomatoneura parvula Martynov 1930
Miomatoneura permica Kukalová 1963
Miomatoneura rotundata Martynov 1928
G. †Miomatoneurella Martynova 1958
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Miomatoneurella americana Rasnitsyn 2004
Miomatoneurella reducta Martynova 1958
Miomatoneurella rossoshana Rasnitsyn and Aristov 2013
G. †Miomatoneurites Zalessky 1956
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Miomatoneurites sylvaensis Zalessky 1956
G. †Neodelopterum Rasnitsyn 2004
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Neodelopterum anomalum Rasnitsyn 2004
G. †Palaeomantina Rasnitsyn 1977
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Palaeomantina pentamera Rasnitsyn 1977
G. †Palaeomantis Handlirsch 1904
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Palaeomantis aestiva Novokshonov 2000
Palaeomantis apicalis Rasnitsyn 2004
Palaeomantis hangardi Guthörl 1962
Palaeomantis laeta Novokshonov and Zhuzhgova 2002
Palaeomantis ostertali Guthörl 1962
Palaeomantis schmidti Handlirsch 1904
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Invalid names: Palaeomantis aequalis Martynov 1928 [synonym]
Palaeomantis sylvaensis Martynov 1940
Palaeomantis tshekardensis Novokshonov 2000
Invalid names: Stefanomioptera Guthörl 1962 [synonym]
G. †Permodelopterum Kukalová 1963
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Permodelopterum obscurum Kukalová 1963
G. †Permonia Kukalová 1963
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Permonia arta Kukalová 1963
Permonia insolida Kukalová 1963
Permonia permoni Kukalová 1963
G. †Permonikia Kukalová 1963
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Permonikia permoniki Kukalová 1963
G. †Perunopterum Kukalová 1963
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Perunopterum corium Kukalová 1963
Perunopterum lepidum Kukalová 1963
Perunopterum lumbiforme Kukalová 1963
Perunopterum macrum Kukalová 1963
Perunopterum mirum Kukalová 1963
Perunopterum peruni Kukalová 1963
Perunopterum saxeum Kukalová 1963
G. †Saaromioptera Guthörl 1963
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Saaromioptera jordani Guthörl 1963
G. †Sellardsiopsis Zalessky 1939
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Sellardsiopsis conspicua Zalessky 1939
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Invalid names: Miomantisca clara Zalessky 1956 [synonym], Miomoptilon pusillus Zalessky 1956 [synonym], Palaeomantisca lata Martynov 1940 [synonym], Paramioma palida Zalessky 1956 [synonym]
Invalid names: Miomantisca Zalessky 1956 [synonym], Miomoptilon Zalessky 1956 [synonym], Palaeomantisca Martynov 1940 [synonym], Paramioma Zalessky 1956 [synonym]
G. †Stigmodelopterum Rasnitsyn 2004
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G. †Tridelopterum Rasnitsyn 2004
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Tridelopterum indebitum Rasnitsyn and Aristov 2013
Tridelopterum variegatum Rasnitsyn 2004
Invalid names: Archisialidae Martynov 1932 [synonym], Delopteridae Sellards 1909 [synonym], Palaeomantiscidae Rasnitsyn 1977 [synonym], Stefanomiopterinae Guthörl 1962 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Marsdenian to the top of the Holocene or 321.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 320.0 Ma

Collections (1403 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Marsdenian321.3 - 320.0Belgium Protoprosbole straeleni (164479)
Westphalian C - Westphalian D314.6 - 307.5France Aviorrhyncha magnifica, Westphalothripides oudardi, Westphalopsocus pumilio (122504)
Westphalian D309.8 - 307.5USA (Illinois) Mazonopsocus testai (188251)
Stephanian A305.1 - 301.5Germany Saaromioptera jordani, Stefanomioptera ostertali, Stefanomioptera hangardi (123835)
Gzhelian303.7 - 298.9Russian Federation Archeglyphis crassinervis (124589) Blattoprosbole tomiensis (124585)
Asselian298.9 - 295.5USA (Colorado) Cercopyllis delicatula, Cercopyllis justiciae, Cercopyllis adolescens (106345)
Asselian298.9 - 295.5Czech Republic Miomatoneura palaeozoica (114187) Permodelopterum obscurum, Delopterum truncatum, Delopterum sp., Permonikia permoniki, Perunopterum peruni, Perunopterum macrum, Perunopterum corium, Perunopterum saxeum, Perunopterum mirum, Permodelopterum lepidum, Permodelopterum lumbiformis, Perunopterum sp., Permonia arta, Permonia insolida, Permonia permoni, Miomatoneurella boskovicensis, Miomatoneura angusta, Miomatoneura candida, Miomatoneura permica, Delopterum lepidum, Delopterum sinuosum (114186) Perunopterum peruni (114188)
Asselian298.9 - 295.5Germany (Rhineland-Palatinate) Permoridium fresenaci (156248)
Asselian298.9 - 295.5USA (New Mexico) Delopterum commune, Delopterum sp., Neodelopterum anomalum (124385) Dichentomum arroyo, Arroyoscyta novaemexicana, Delopterum pantherinum, Delopterum commune, Delopterum zonatum, Tridelopterum variegatum, Miomatoneurella americana, Palaeomantis apicalis, Stigmodelopterum pterostigmalis, Delopterinus zebra, Delopterinus bicolor, Delopsocus stenopterus (124386) Dichentomum sp., Delopterinus bicolor, Palaeomanteidae indet. (124387)
Permian298.9 - 251.902China (Nei Mongol) Palaeocicadopsis chinensis (156333)
Artinskian290.1 - 279.3USA (Kansas) Dichentomum complexum (151412) Dichentomum tinctum, Delopterum minutum, Delopterum latum, Delopterum elongatum, Urba punctata (21979) Elmomantis engeli (114450) Lithopsocidium permianum, Permopsocus latipennis, Permopsocus ovatus, Dichentomum tinctum, Permopsylla minuta, Permopsylla americana, Permopsylla permiana, Permopsylla grandis, Archescytina permiana, Permoscytina kansasensis, Delopterum minutum, Delopterum elongatum, Delopterum kansanum (114151) Orthopsocus singularis, Progonopsocus permianus, Lithopsocidium permianum, Permopsocus latipennis, Dichentomum tinctum, Dichentomum minimum, Dichentomum latum, Dichentomum parvulum, Permopsylla minuta, Permopsylla americana, Permopsylla permiana, Archescytina permiana, Permoscytina kansasensis, Permoscytina muiri, Paleoscytina brevistigma, Cyphoneura permiana, Cyphoneura reducta (111217) Permopsocus latipennis, Permopsylla minuta, Permopsylla americana, Delopterum latum (114152) Progonopsocus permianus, Lithopsocidium permianum, Permopsocus latipennis, Dichentomum tinctum, Dichentomum grande, Permopsylla americana, Permopsylla anomala, Permopsylla grandis, Permopsylla sp., Lithoscytina cubitalis, Delopterum minutum, Delopterum elongatum (111319) Progonopsocus permianus, Permopsocus latipennis, Dichentomum tinctum, Permopsylla americana, Delopterum minutum, Delopterum latum, Delopterum elongatum (111320) Progonopsocus pusillus, Progonopsocus permianus, Permopsocus congener, Permopsocus enderleini, Ancylopsocus insolitus, Permopsocus latipennis, Psocidium permianum, Psocidium abnormale, Psocidium kansasense, Chaetopsocidium sellardsi, Permentomum tenuiforme, Pentapsocidium indistinctum, Metapsocidium loxoneurum, Psocidium robustum, Dichentomum tinctum, Psocidium sp., Psocidium minimum, Permopsylla americana, Archescytina permiana, Permoscytina kansasensis, Permoscytina muiri, Delopterum dunbari, Delopterum minutum, Delopsocus latus, Delopsocus fasciatus, Delopsocus elongatus, Delopteridae indet. (111219)
Artinskian290.1 - 279.3USA (Oklahoma) Cyphoneurodes patriciae (104818) Delopterum latum (104816) Dichentomum tinctum, Permopsylla americana, Parapermopsylla tricubita, Delopterum minutum, Delopterum latum, Delopterum elongatum (104826)
Artinskian290.1 - 279.3Uruguay Paracicadopsis mendezalzolai (124521)
Artinskian290.1 - 279.3France (Languedoc) Lutevanaphis permiana, Palaeomantis sp. (105044) Permopsyllidium lesclansis (122883)
Kungurian279.3 - 272.3France (Languedoc) Pervestigia veteris, Thripida indet. (105045) Prosbole dio (122920)
Kungurian279.3 - 272.3Russian Federation (Perm) Delopteriella graciosa (111218) Idelopsocus incommendatus (117254) Scytoneurella sp. (173727)
Kungurian279.3 - 272.3Russian Federation Cicadopsis sp. (115214) Delopterum candidum, Palaeomantis tshekardensis, Palaeomantis aestiva, Hypoperla vaulevi, Idelopsocus splendens (107787) Idelopsocus galinae (124284) Maueria pusillus (196229) Maueria rhynchota (155821) Palaeomantis aestiva (114207) Parapsocidium uralicum, Permopsyllopsis maueriaeformis, Sellardsiopsis conspicua (196228) Parapsocidium uralicum, Scytoneurella major, Scytoneurella minor, Maueria sylvensis, Permoscytinopsis maueriaeformis, Maueria intermedia, Maueria rhynchota, Uraloscytina prosbolioides, Maueria pusillus, Permopsyllopsis rossica, Maripsocus ambiguus, Miomatoneurites sylvaensis, Miomantisca clara, Paramioma palida, Miomoptilon pusillus, Sellardsiopsis conspicua, Hypoperlopsis splendens (102504) Parapsocidium uralicum, Uraloscytina similis (155819) Uraloscytina multinervosa (152060)
Kungurian279.3 - 272.3Russian Federation (Suksun) Archescytina tshekardaensis, Archescytinidae indet., Palaeomantisca lata (116641) Dichentomum sp., Dinoscyta microcephala, Tschekardus hispidus, Delopterum rasnitsyni, Palaeomantis laeta, Palaeomantis aestiva, Palaeomantina pentamera, Hypoperla grata, Idelopsocus diradiatus (108206) Palaeomantis sylvaensis (113379) Tshekardaella tshekardaensis (124241)
Kungurian279.3 - 272.3South Africa (Gauteng) Protopsyllidiidae indet. (122863)
Kungurian279.3 - 272.3France Archescytina sp. A, Euroscytina lutevanorum (103637) Prosbole garrici (174415)