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Acercaria

Insecta

Taxonomy
Acercaria was named by Börner (1904). It is extant.

It was considered an invalid subgroup of Neoptera by Misof et al. (2014); it was reranked as the unranked clade Acercaria by Prokop et al. (2017).

It was assigned to Neoptera by Prokop et al. (2015); and to Clareocercaria by Prokop et al. (2017).

Sister taxa
Pancondylognatha

Synonymy list
YearName and author
1904Acercaria Börner p. 526
2015Acercaria Prokop et al. p. 40
2017Acercaria Prokop et al. p. 2

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
RankNameAuthor
Dicondylia
Paranotalia
subclassPterygota()
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
Clareocercaria
Acercaria()
Acercaria()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. Acercaria Börner 1904
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Superor. Pancondylognatha Yoshizawa and Leinhard 2016
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Unr. Condylognatha Börner 1904
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Or. Hemiptera Linnaeus 1758 [true bug]
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Subor. Auchenorrhyncha Duméril 1805 [true bug]
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Fm. †Blattoprosbolidae Becker-Migdisova 1958
Infraor. Cicadomorpha Evans 1946
Fm. †Permofulgoridae Tillyard 1918
Invalid names: Jassidea Becker-Migdisova 1960 [empty]
G. Belostomum Latreille 1807 [giant water bug]
G. †Cercopyllis Scudder 1890
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Cercopyllis adolescens Scudder 1890
Cercopyllis delicatula Scudder 1890
Cercopyllis justiciae Scudder 1890
Fm. †Cicadopsyllidae Martynov 1931
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G. †Paracicadopsis Pinto and Piñeiro 2000
G. †Scytoneurella Zalessky 1939
Subor. Coleorrhyncha Myers and China 1929
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Infraor. Peloridiomorpha Popov 1982
Fm. †Permoridiidae Burckhardt et al. 2022
Superfm. †Progonocimicoidea Handlirsch 1906
Unr. †Progonocimicomorpha Popov 1982
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Invalid names: Actinoscytinomorpha Popov 1982 [synonym]
Unr. Hemelytrata Fallen 1829
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Subor. Fulgoromorpha Evans 1946 [planthopper]
Unr. Prosorrhyncha Sorensen et al. 1995
Subor. Heteroptera Latreille 1810
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G. †Anacoloptera Handlirsch 1906
G. †Berytopsis Heer 1853
Infraor. Cimicomorpha Leston et al. 1954
Infraor. Dipsocoromorpha Miyamoto 1961
G. †Duncanovelia Jell and Duncan 1986
Unr. Euheteroptera Stys 1985
G. †Galerucites Oppenheim 1888
Infraor. Gerromorpha Popov 1971 [semi-aquatic bug]
Fm. †Hadrocoridae Handlirsch 1920
G. †Hebeicoris Hong 1983
Infraor. Hydrocorisae Latreille 1810
Infraor. Leptopodomorpha Popov 1971 [shore bug]
G. †Liasocoris Wendt 1940
Infraor. Nepomorpha Popov 1971
Infraor. Pentatomomorpha Leston et al. 1954
G. †Sinanabis Zhang 1986
G. †Sinocoris Hong 1983
G. †Sinovelia Yao et al. 2012
G. †Sphongophoriella Becker-Migdisova 1958
Invalid names: Corixa pullus Von Heyden 1859 [nomen dubium], Cryptocerata [empty], Eocimicidae Handlirsch 1906 [empty], Galgulidae Westwood 1840 [empty], Nepa atavina Heer 1853 [nomen dubium], Xishanidae Hong 1981 [empty]
Subor. Homoptera Leach 1815
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G. †Aleuronympha Riek 1974
G. †Homopteron Handlirsch 1907
G. †Knezouria Jell 1993
G. †Mesaleuropsis Martynov 1937
G. †Palaeomyrmex Heer 1864
Fm. †Permaleurodidae Becker-Migdisova 1959
G. †Permocapitus Evans 1943
Invalid names: Biturritiidae Metcalf 1951 [empty], Coccaria Becker-Migdisova 1960 [empty], Lophophidae Stal 1866 [empty], Psyllinea [empty]
Fm. Hydrocorisae Latreille 1802
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Invalid names: Notonectariae Latreille 1802 [empty]
G. †Hypocixius Cockerell 1926
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Hypocixius oblitescens Cockerell 1926
G. †Ludibrium Becker-Migdisova 1962
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Ludibrium ludus Becker-Migdisova 1962
G. †Meshemipteron Cockerell 1915
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Meshemipteron incertum Cockerell 1915
G. †Mesococcus Becker-Migdisova 1959
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Mesococcus asiatica Becker-Migdisova 1959
Subor. †Paleorrhyncha Carpenter 1931
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Superfm. †Archescytinoidea Tillyard 1926
G. †Planophlebia Scudder 1879
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Planophlebia gigantea Scudder 1879
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G. †Protoprosbole Laurentiaux 1952
G. †Shaandongia Hong 1985
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Subor. Sternorrhyncha Amyot and Audinet-Serville 1843
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Unr. Aphidiformes Schlee 1969
Fm. †Archiconiopterygidae Ansorge 1996
Superfm. Coccoidea Fallen 1814 [scale insect]
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Invalid names: Orthezioidea Amyot and Audinet-Serville 1843 [synonym]
Infraor. †Coccomorpha Beier 1938 [scale insect]
Infraor. †Psyllaria Becker-Migdisova 1960
Invalid names: Aphidii Latreille 1802 [empty], Auchenorinques Duméril 1805 [empty], Biturritidae Evans 1953 [empty], Cimicides Latreille 1802 [empty], Delphax rhenana Statz 1950 [nomen dubium], Delphax senilis Scudder 1877 [nomen dubium], Eofulgoridium chanmaense Hong 1982 [nomen dubium], Gallinsecta Latreille 1802 [empty], Heteropterodea Zrzavy 1992 [empty], Limnochares antiquus von Heyden 1862 [nomen dubium], Lithopsis lineatus Piton 1940 [nomen dubium], Lithopsis punctinervis Piton 1940 [nomen dubium], Lystra vollenhoveni Weyenbergh 1869 [nomen dubium], Palaeohemiptera Handlirsch 1906 [empty], Peloridiina Popov 1982 [empty], Pincombella Chapman 1932 [nomen dubium], Pincombella belmontensis Chapman 1932 [nomen dubium], Proboscidea Scopoli 1763 [synonym], Protocoris ovalis Heer 1864 [nomen nudum], Ryngota Fabricius 1775 [synonym]
Superor. Thripida Fallen 1814
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Or. †Lophioneurida Tillyard 1921
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Fm. †Lophioneuridae Tillyard 1921
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Invalid names: Edgariekiidae Jell and Duncan 1986 [synonym]
Fm. †Moundthripidae Nel et al. 2007
G. †Pseudoundacypha Huang 2023
Fm. †Westphalothripidesidae Nel et al. 2012
Fm. †Zoropsocidae Tillyard 1935
Unr. †Panthysanoptera Nel et al. 2012
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Invalid names: Zoropsocinae Tillyard 1935 [empty]
Or. Thysanoptera Haliday 1836 [thrips]
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Subor. Terebrantia Haliday 1836
Subor. Tubulifera Haliday 1836
Invalid names: Lophioneurina Tillyard 1921 [empty], Thrips annosa Heer 1864 [nomen dubium], Thrips annulata Menge 1856 [nomen dubium], Thrips antiqua Heer 1856 [nomen dubium], Thrips oeningensis Heer 1864 [nomen nudum], Thrips sericata Menge 1856 [nomen dubium]
Invalid names: Cyphoneuridae Carpenter 1932 [empty]
Or. †Permopsocida Tillyard 1926
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Fm. †Archipsyllidae Handlirsch 1906
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G. †Archipsylla Handlirsch 1906
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Archipsylla lata Vishnyakova 1976
Archipsylla primitiva Handlirsch 1906
Archipsylla similis Vishnyakova 1976
Archipsylla sinica Huang et al. 2008
Archipsylla turanica Martynov 1926
G. †Archipsyllodes Vishnyakova 1976
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Archipsyllodes speciosus Vishnyakova 1976
G. †Archipsyllopsis Vishnyakova 1976
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Archipsyllopsis baissica Vishnyakova 1976
G. †Bittacopsocus Beutel et al. 2019
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Bittacopsocus megacephalus Beutel et al. 2019
G. †Burmopsylla Liang et al. 2016
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Burmopsylla maculata Liang et al. 2016
G. †Dinmopsylla Lambkin 2018
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Dinmopsylla semota Lambkin 2018
G. †Eopsylla Vishnyakova 1976
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Eopsylla sojanense Becker-Migdisova 1962
G. †Mydiognathus Yoshizawa and Leinhard 2016
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Mydiognathus eviohlhoffae Yoshizawa and Leinhard 2016
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Invalid names: Psocorrhyncha burmitica Huang et al. 2016 [synonym]
Invalid names: Psocorrhyncha Huang et al. 2016 [synonym]
Fm. †Permopsocidae Tillyard 1926
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G. †Lithopsocidium Carpenter 1932
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Lithopsocidium permianum Carpenter 1932
G. †Orthopsocus Carpenter 1932
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Orthopsocus singularis Carpenter 1932
G. †Permopsocus Tillyard 1926
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Permopsocus latipennis Tillyard 1926
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Invalid names: Ancylopsocus insolitus Tillyard 1926 [synonym], Permopsocus congener Tillyard 1926 [synonym], Permopsocus enderleini Tillyard 1926 [synonym]
Permopsocus ovatus Carpenter 1939
Invalid names: Ancylopsocus Tillyard 1926 [synonym]
G. †Progonopsocus Tillyard 1926
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Progonopsocus permianus Tillyard 1926
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Invalid names: Progonopsocus pusillus Tillyard 1926 [synonym]
Fm. †Psocidiidae Tillyard 1926
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G. †Austropsocidium Tillyard 1935
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Austropsocidium pincombei Tillyard 1935
Austropsocidium stigmaticum Tillyard 1935
G. †Dichentomum Tillyard 1926
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Dichentomum arroyo Rasnitsyn 2004
Dichentomum complexum Carpenter 1926
Dichentomum grande Carpenter 1933
Dichentomum latum Carpenter 1932
Dichentomum minimum Tillyard 1926
Dichentomum parvulum Carpenter 1932
Dichentomum tinctum Tillyard 1926
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Invalid names: Chaetopsocidium sellardsi Tillyard 1926 [synonym], Metapsocidium loxoneurum Tillyard 1926 [synonym], Pentapsocidium indistinctum Tillyard 1926 [synonym], Permentomum tenuiforme Tillyard 1926 [synonym], Psocidium abnormale Tillyard 1926 [synonym], Psocidium kansasense Tillyard 1926 [synonym], Psocidium permianum Tillyard 1926 [synonym], Psocidium robustum Tillyard 1926 [synonym]
Invalid names: Chaetopsocidium Tillyard 1926 [synonym], Metapsocidium Tillyard 1926 [synonym], Pentapsocidium Tillyard 1926 [synonym], Permentomum Tillyard 1926 [synonym], Psocidium Tillyard 1926 [synonym]
G. †Hypopsylla Prokop et al. 2016
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Hypopsylla belmontensis Prokop et al. 2016
G. †Liassopsocus Ansorge 1996
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Liassopsocus lanceolatus Ansorge 1996
G. †Megapsocidium Tillyard 1935
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Megapsocidium australe Tillyard 1935
G. †Parapsocidium Zalessky 1937
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Parapsocidium uralicum Zalessky 1937
G. †Stenopsocidium Tillyard 1935
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Stenopsocidium elongatum Tillyard 1935
Invalid names: Dichentomidae Carpenter 1932 [synonym]
Fm. †Westphalopsocidae Azar et al. 2013
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G. †Westphalopsocus Azar et al. 2013
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Westphalopsocus pumilio Azar et al. 2013
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Marsdenian to the top of the Holocene or 321.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 320.0 Ma

Collections (1385 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Marsdenian321.3 - 320.0Belgium Protoprosbole straeleni (164479)
Westphalian C - Westphalian D314.6 - 307.5France Aviorrhyncha magnifica, Westphalothripides oudardi, Westphalopsocus pumilio (122504)
Gzhelian303.7 - 298.9Russian Federation Archeglyphis crassinervis (124589) Blattoprosbole tomiensis (124585)
Asselian298.9 - 295.5USA (Colorado) Cercopyllis delicatula, Cercopyllis justiciae, Cercopyllis adolescens (106345)
Asselian298.9 - 295.5Germany (Rhineland-Palatinate) Permoridium fresenaci (156248)
Asselian298.9 - 295.5USA (New Mexico) Dichentomum arroyo, Arroyoscyta novaemexicana (124386) Dichentomum sp. (124387)
Permian298.9 - 251.902China (Nei Mongol) Palaeocicadopsis chinensis (156333)
Artinskian290.1 - 279.3USA (Kansas) Dichentomum complexum (151412) Dichentomum tinctum (21979) Lithopsocidium permianum, Permopsocus latipennis, Permopsocus ovatus, Dichentomum tinctum, Permopsylla minuta, Permopsylla americana, Permopsylla permiana, Permopsylla grandis, Archescytina permiana, Permoscytina kansasensis (114151) Orthopsocus singularis, Progonopsocus permianus, Lithopsocidium permianum, Permopsocus latipennis, Dichentomum tinctum, Dichentomum minimum, Dichentomum latum, Dichentomum parvulum, Permopsylla minuta, Permopsylla americana, Permopsylla permiana, Archescytina permiana, Permoscytina kansasensis, Permoscytina muiri, Paleoscytina brevistigma, Cyphoneura permiana, Cyphoneura reducta (111217) Permopsocus latipennis, Permopsylla minuta, Permopsylla americana (114152) Progonopsocus permianus, Lithopsocidium permianum, Permopsocus latipennis, Dichentomum tinctum, Dichentomum grande, Permopsylla americana, Permopsylla anomala, Permopsylla grandis, Permopsylla sp., Lithoscytina cubitalis (111319) Progonopsocus permianus, Permopsocus latipennis, Dichentomum tinctum, Permopsylla americana (111320) Progonopsocus pusillus, Progonopsocus permianus, Permopsocus congener, Permopsocus enderleini, Ancylopsocus insolitus, Permopsocus latipennis, Psocidium permianum, Psocidium abnormale, Psocidium kansasense, Chaetopsocidium sellardsi, Permentomum tenuiforme, Pentapsocidium indistinctum, Metapsocidium loxoneurum, Psocidium robustum, Dichentomum tinctum, Psocidium sp., Psocidium minimum, Permopsylla americana, Archescytina permiana, Permoscytina kansasensis, Permoscytina muiri (111219)
Artinskian290.1 - 279.3France (Languedoc) Lutevanaphis permiana (105044) Permopsyllidium lesclansis (122883)
Artinskian290.1 - 279.3Uruguay Paracicadopsis mendezalzolai (124521)
Artinskian290.1 - 279.3USA (Oklahoma) Cyphoneurodes patriciae (104818) Dichentomum tinctum, Permopsylla americana, Parapermopsylla tricubita (104826)
Kungurian279.3 - 272.3France (Languedoc) Pervestigia veteris, Thripida indet. (105045) Prosbole dio (122920)
Kungurian279.3 - 272.3Russian Federation (Perm) Scytoneurella sp. (173727)
Kungurian279.3 - 272.3Russian Federation (Suksun) Archescytina tshekardaensis, Archescytinidae indet. (116641) Dichentomum sp., Dinoscyta microcephala, Tschekardus hispidus (108206) Tshekardaella tshekardaensis (124241)
Kungurian279.3 - 272.3Russian Federation Cicadopsis sp. (115214) Maueria pusillus (196229) Maueria rhynchota (155821) Parapsocidium uralicum, Permopsyllopsis maueriaeformis (196228) Parapsocidium uralicum, Scytoneurella major, Scytoneurella minor, Maueria sylvensis, Permoscytinopsis maueriaeformis, Maueria intermedia, Maueria rhynchota, Uraloscytina prosbolioides, Maueria pusillus, Permopsyllopsis rossica, Maripsocus ambiguus (102504) Parapsocidium uralicum, Uraloscytina similis (155819) Uraloscytina multinervosa (152060)
Kungurian279.3 - 272.3South Africa (Gauteng) Protopsyllidiidae indet. (122863)
Kungurian279.3 - 272.3Russian Federation (Kemerovo) Sarbaloptera sarbalensis (104951) Sarbaloscytina angustipennis, Kaltanoscytina latistigmata, Sarbaloptera sarbalensis, Scytinoptera sibirica, Prosbole sp., Lophioneurodes sarbalensis (112708)
Kungurian279.3 - 272.3France Archescytina sp. A, Euroscytina lutevanorum (103637) Prosbole garrici (174415)
Kungurian279.3 - 272.3Brazil (Rio Grande do Sul) Gondwanaptera capsii, Fulgoringruo kukalovae, Prosbole iratiensis, Prosbolecicada gondwanica (116121)
Guadalupian273.01 - 259.51Russian Federation Pereboria bella (173159)