Basic info Taxonomic history Classification Included Taxa
Morphology Ecology and taphonomy External Literature Search Age range and collections

Carnivora (carnivoran)

Mammalia - Carnivora

Taxonomy
Carnivora was named by Bowdich (1821). It is extant.

It was reranked as the suborder Carnivora by Ameghino (1897); it was synonymized subjectively with Ferae by Hay (1902).

It was assigned to Unguiculata by Cope (1889); to Eutheria by Flower and Lydekker (1891); to Sarcobora by Ameghino (1897); to Holotheria by Jaekel (1911); to Monodelphia by Hay (1930); to Eutheria by Hooker (1986); to Zooamata by Waddell et al. (1999); to Ferae by Lopatin (2006); to Laurasiatheria by Madsen et al. (2001) and Silcox et al. (2010); to Ferae by McKenna (1975), Waddell et al. (2001) and Asher and Helgen (2010); to Carnivoramorpha by Wesley-Hunt and Werdelin (2005), Spaulding et al. (2009), Spaulding et al. (2010) and Tomiya (2011); to Eutheria by Flower (1883), Gadow (1898), Carroll (1988), Nowak (1991) and Hunt (2011); to Eutheria by Vianey-Liaud et al. (2014); to Carnivoraformes by Solé et al. (2016); to Mammalia by Leidy (1873), Lydekker (1885), Andrews (1906), Lambe (1908), Tedford (1976), Barnes (1979), Thurmond and Jones (1981), Barnes (1987), Barnes (1988), Shoshani et al. (1989), Barnes (1989), Kohno (1992), Barnes (1992), Tedford et al. (1994), Barnes and Hirota (1995), Kohno (1996), Ginsburg and Janvier (1999), Sepkoski (2002), Wang et al. (2004), Davis et al. (2004), Wilson and Reeder (2005), Sotnikova (2006), Barnes et al. (2006), Hulbert and Whitmore (2006), Kohno (2006), García (2008), Wang and Tedford (2008), Tedford and Wang (2008), Kohno and Ray (2008), Mazak et al. (2011), Wang et al. (2015), Tomiya and Tseng (2016) and Bōgner and Samuels (2022); and to Mammalia by Hontecillas et al. (2023).

Synonyms
Synonymy list
YearName and author
1758Viverra memphitis Linnaeus p. 44
1821Carnivora Bowdich
1873Carnivora Leidy p. 315
1883Carnivora Flower p. 186
1885Carnivora Lydekker p. 196
1889Carnivora Cope p. 875
1891Carnivora Flower and Lydekker p. 91
1897Carnivora Ameghino p. 501
1898Carnivora Gadow p. 50
1906Carnivora Andrews p. 218
1908Carnivora Lambe
1911Carnivora Jaekel p. 227
1930Carnivora Hay p. 456
1945Eosictis Scott
1975Carnivora McKenna p. 42
1976Carnivora Tedford p. 372
1979Carnivora Barnes p. 8
1981Carnivora Thurmond and Jones p. 174
1986Carnivora Hooker p. 326
1987Carnivora Barnes p. 3
1988Carnivora Barnes p. 2
1988Carnivora Carroll
1989Carnivora Barnes p. 3
1989Carnivora Shoshani et al. p. 436
1991Carnivora Nowak
1992Carnivora Barnes p. 3
1992Carnivora Kohno p. 16
1994Carnivora Tedford et al. p. 12
1995Carnivora Barnes and Hirota p. 330
1996Carnivora Kohno p. 389
1999Carnivora Ginsburg and Janvier p. 170
1999Carnivora Waddell et al. p. 3
2001Carnivora Madsen et al. p. 610
2001Carnivora Waddell et al. p. 148
2002Carnivora Sepkoski
2004Carnivora Davis et al. p. 365
2004Carnivora Wang et al.
2005Carnivora Wesley-Hunt and Werdelin p. 842
2005Carnivora Wilson and Reeder
2006Carnivora Barnes et al. p. 59
2006Carnivora Hulbert and Whitmore p. 5
2006Carnivora Kohno p. 411
2006Carnivora Lopatin
2006Carnivora Sotnikova
2008Carnivora García
2008Carnivora Kohno and Ray p. 44
2008Carnivora Tedford and Wang p. 273
2008Carnivora Wang and Tedford
2009Carnivora Spaulding et al.
2010Carnivora Asher and Helgen p. 3 figs. Figure 1
2010Carnivora Silcox et al.
2010Carnivora Spaulding et al. p. 815
2011Carnivora Hunt, Jr. p. 21
2011Carnivora Mazak et al.
2011Carnivora Tomiya p. 3
2014Carnivora Vianey-Liaud et al. p. 610
2015Carnivora Wang et al.
2016Carnivora Solé et al.
2016Carnivora Tomiya and Tseng p. 5
2022Carnivora Bōgner and Samuels
2023Carnivora Hontecillas et al. p. 6

Is something missing? Join the Paleobiology Database and enter the data

RankNameAuthor
kingdomAnimalia()
—Bilateria
—EubilateriaAx 1987
—Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
—Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
—Tetrapoda
—Reptiliomorpha
—Anthracosauria
subclassAmphibiosauriaKuhn 1967
—Cotylosauria()
—Amniota
subclassSynapsida
—Therapsida()
RankNameAuthor
infraorderCynodontia()
—Mammaliamorpha
—Mammaliaformes
classMammalia
—Cladotheria
—Zatheria
subclassTribosphenida()
subclassTheria
—Eutheria()
—Placentalia
—Boreoeutheria
—Laurasiatheria
—Scrotifera
—Ferae()
—CarnivoramorphaWyss and Flynn 1993
—CarnivoraformesFlynn et al.
orderCarnivora
orderCarnivora

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Or. Carnivora Bowdich 1821 [carnivoran]
show all | hide all
G. †Adracon Filhol 1884
Fm. Canidae Fischer de Waldheim 1817 [canid]
+
Subfm. †Borophaginae Simpson 1945 [bone-crushing dog]
+
G. †Archaeocyon Wang et al. 1999
+
†Archaeocyon falkenbachi Wang et al. 1999
†Archaeocyon leptodus Schlaikjer 1935
†Archaeocyon pavidus Stock 1933
Tr. †Borophagini Simpson 1945
+
Subtr. †Aelurodontina Wang et al. 1999
+
G. †Aelurodon Leidy 1858
+
Invalid names: Prohyaena Schlosser 1890 [synonym], Strobodon Webb 1969 [synonym]
G. †Tomarctus Cope 1873
Subtr. †Borophagina Wang et al. 1999
+
G. †Carpocyon Webb 1969
G. †Epicyon Leidy 1858
G. †Paratomarctus Wang et al. 1999
G. †Protepicyon Wang et al. 1999
G. †Cormocyon Wang and Tedford 1992
+
†Cormocyon copei Wang and Tedford 1992
†Cormocyon haydeni Wang et al. 1999
Subtr. †Cynarctina Wang et al. 1999
+
G. †Cynarctus Matthew 1902
G. †Euoplocyon Matthew 1924
G. †Metatomarctus Wang et al. 1999
G. †Microtomarctus Wang et al. 1999
G. †Paracynarctus Wang et al. 1999
G. †Protomarctus Wang et al. 1999
G. †Psalidocyon Wang et al. 1999
G. †Tephrocyon Merriam 1906
G. †Desmocyon Wang et al. 1999
+
†Desmocyon matthewi Wang et al. 1999
†Desmocyon thomsoni Matthew 1907
+
Invalid names: Nothocyon regulus Cook and Macdonald 1962 [synonym]
Invalid names: Cynodesmus petersoni Radinsky 1973 [nomen nudum]
G. †Borophagus Cope 1892
+
+
Invalid names: Felis hillanus Cope 1893 [synonym], Hyaenognathus matthewi Freudenberg 1910 [synonym], Hyaenognathus pachyodon Merriam 1903 [synonym], Hyaenognathus solus Stock 1932 [synonym], Porthocyon dubius Merriam 1903 [synonym]
†Borophagus dudleyi White 1941
†Borophagus hilli Johnston 1939
+
Invalid names: Osteoborus crassapineatus Olsen 1956 [synonym], Osteoborus progressus Hibbard 1944 [synonym]
†Borophagus littoralis VanderHoof 1931
+
Invalid names: Osteoborus diabloensis Richey 1938 [synonym]
†Borophagus orc Webb 1969
†Borophagus parvus Wang et al. 1999
†Borophagus pugnator Cook 1922
+
Invalid names: Osteoborus galushai Webb 1969 [synonym]
†Borophagus secundus VanderHoof 1931
+
Invalid names: Hyaenognathus cyonoides Martin 1928 [synonym], Hyaenognathus direptor Matthew 1924 [synonym]
Invalid names: Cynogulo [synonym], Hyaenognathus Merriam 1903 [synonym], Osteoborus Stirton and VanderHoof 1933 [synonym], Pliogulo White 1941 [synonym], Porthocyon Merriam 1903 [synonym]
G. †Otarocyon Wang et al. 1999
+
†Otarocyon cooki Macdonald 1963
†Otarocyon cuspidatus Green 1954
†Otarocyon macdonaldi Wang et al. 1999
G. †Oxetocyon Green 1954
+
†Oxetocyon cuspidatus Green 1954
Tr. †Phlaocyonini Wang et al. 1999
+
G. †Cynarctoides McGrew 1938
+
†Cynarctoides acridens Barbour and Cook 1914
+
Invalid names: Cynarctus mustelinus Matthew 1932 [synonym]
†Cynarctoides emryi Wang et al. 1999
†Cynarctoides gawnae Wang et al. 1999
†Cynarctoides harlowi Loomis 1932
†Cynarctoides lemur Cope 1879
†Cynarctoides luskensis Wang et al. 1999
†Cynarctoides roii Macdonald 1963
†Cynarctoides whistleri Wang and Tedford 2008
G. †Phlaocyon Matthew 1899
+
†Phlaocyon achoros Frailey 1979
†Phlaocyon annectens Peterson 1907
+
Invalid names: Phlaocyon willistoni Peterson 1923 [synonym]
†Phlaocyon latidens Cope 1881
†Phlaocyon leucosteus Matthew 1899
†Phlaocyon mariae Wang et al. 1999
†Phlaocyon marslandensis McGrew 1941
†Phlaocyon minor Matthew 1907
†Phlaocyon multicuspus Romer and Sutton 1927
†Phlaocyon taylori Hayes 2000
†Phlaocyon yatkolai Wang et al. 1999
Invalid names: Aletocyon Romer and Sutton 1927 [synonym], Bassariscops Peterson 1928 [synonym]
G. †Rhizocyon Wang et al. 1999
+
†Rhizocyon oregonensis Merriam 1906
Subfm. Caninae Gill 1872 [canine]
+
G. †Aenocyon Merriam 1918
+
†Aenocyon dirus Leidy 1858 [dire wolf]
+
Invalid names: Canis ayersi Sellards 1916 [synonym], Canis indianensis Leidy 1869 [synonym], Canis mississippiensis Allen 1876 [synonym], Canis nehringi Ameghino 1902 [synonym], Canis primaevus Hodgson 1833 [synonym]
Invalid names: Stereocyon Mercerat 1917 [synonym]
Tr. †Canini Fischer de Waldheim 1817
+
Subtr. Canina Gray 1825
+
G. †Nurocyon Sotnikova 2006
Subtr. †Cerdocyonina Tedford et al. 2009
+
G. Atelocynus Cabrera 1940 [short-eared dog]
G. Cerdocyon Hamilton-Smith 1839 [crab-eating fox]
+
Invalid names: Carcinocyon Allen 1905 [synonym]
G. †Dusicyon Hamilton Smith 1839 [Falkland Island wolf]
G. Speothos Lund 1839 [bush dog]
+
Invalid names: Abathmodon Lund 1843 [synonym]
G. †Eucyon Tedford and Qiu 1996
+
†Eucyon adoxus Martin 1971
†Eucyon davisi Merriam 1911
+
Invalid names: Canis condoni Shotwell 1956 [synonym], Leptocyon shermanensis Hibbard 1937 [synonym]
†Eucyon debonisi Montoya et al. 2009
†Eucyon ferox Miller and Carranza-Castaneda 1998
†Eucyon intrepidus Morales et al. 2005
†Eucyon khoikhoi Valenciano et al. 2022
†Eucyon kuta Werdelin et al. 2014
†Eucyon marinae Spassov and Rook 2006
†Eucyon minor Teilhard de Chardin and Piveteau 1930
†Eucyon monticinensis Rook 1992
†Eucyon odessanus Odintzov 1967
†Eucyon skinneri Tedford et al. 2009
†Eucyon wokari García 2008
†Eucyon zhoui Tedford and Qiu 1996
G. †Sinicuon Kretzoi 1941
+
†Sinicuon dubius Teilhard de Chardin 1940
+
Invalid names: Sinicuon peii Kretzoi 1941 [objective synonym]
G. †Xenocyon Kretzoi 1938
+
†Xenocyon africanus Pohle 1928
+
Invalid names: Canis atrox Broom 1946 [synonym]
†Xenocyon antonii Zdansky 1924
†Xenocyon brevicephalus Qiu et al. 2004
†Xenocyon dubius Teilhard de Chardin 1940
†Xenocyon falconeri Forsyth-Major 1877
†Xenocyon lycaonoides Kretzoi 1938
+
Invalid names: Cuon rosi Pons-Moyá and Moyá-Solá 1978 [synonym], Cuon stehlini Thenius 1954 [synonym]
†Xenocyon texanus Troxell 1915
G. Canis Linnaeus 1758 [canine]
+
Subg. †Canis (Epicyon) Leidy 1858
†Canis accitanus Garrido and Arribas 2008
†Canis adoxus Martin 1971
†Canis apolloniensis Koufos and Kostopoulos 1997
†Canis armbrusteri Gidley 1913
†Canis arnensis Del Campana 1913
Canis aureus Linnaeus 1758 [golden jackal]
†Canis borjgali Bartolini-Lucenti et al. 2020
Canis cana Blanford 1877 [Blanfords fox]
†Canis cedazoensis Mooser and Dalquest 1975
Canis ceylonicus Shaw 1800
†Canis chihliensis Zdansky 1924
†Canis chihliensis palmidens Teilhard de Chardin and Piveteau 1930
†Canis cipio Crusafont-Pairó 1950
Canis culpaeus Molina 1782 [culpeo]
Canis dingo Blumenbach 1780 [dingo]
†Canis edwardii Gazin 1942
†Canis etruscus Forsyth Major 1877
+
Invalid names: Canis majori Del Campana 1913 [synonym], Canis olivolanus Del Campana 1913 [synonym]
Canis familiaris Linnaeus 1758 [dog]
+
Canis familiaris sagax Linnaeus 1758
Invalid names: Pachycyon robustus Allen 1885 [synonym]
†Canis feneus Tedford et al. 2009
†Canis geismarianus Cope 1878
†Canis gezi Kraglievich 1928
+
Invalid names: Macrocyon chapalmalensis Frenguelli 1929 [synonym]
†Canis hewitti Fourvel and Frerebeau 2023
†Canis incertus D'Orbigny and Laurillard 1842
†Canis latidentatus Cope 1899
Canis latrans Say 1823 [coyote]
+
Invalid names: Canis andersoni Merriam 1910 [synonym], Canis caneloensis Skinner 1942 [synonym], Canis clepticus Eliot 1903 [synonym], Canis estor Merriam 1897 [synonym], Canis frustror Woodhouse 1851 [synonym], Canis goldmani Merriam 1904 [synonym], Canis hondurensis Goldman 1936 [synonym], Canis impavidus Allen 1903 [synonym], Canis irvingtonensis Savage 1951 [synonym], Canis jamesi Townsend 1912 [synonym], Canis lestes Merriam 1897 [synonym], Canis mearnsi Merriam 1897 [synonym], Canis microdon Merriam 1897 [synonym], Canis nebrascensis Merriam 1898 [synonym], Canis ochropus Eschscholtz 1829 [synonym], Canis orcutti Merriam 1910 [synonym], Canis peninsulae Merriam 1897 [synonym], Canis riviveronis Hay 1917 [synonym], Canis vigilis Merriam 1897 [synonym], Lyciscus cagottis Hamilton-Smith 1839 [synonym]
†Canis lepophagus Johnston 1938
†Canis longdanensis Qiu et al. 2004
Canis lupaster Hemprich and Eherenberg 1832 [African wolf]
+
Invalid names: Canis anthus Cuvier 1820 [synonym]
Canis lupus Linnaeus 1758 [gray wolf]
+
Invalid names: Canis chanco Gray 1863 [synonym], Canis furlongi Hay 1927 [synonym], Canis hodophilax Temminck 1839 [synonym], Canis lycaon Schreber 1775 [synonym], Canis milleri Merriam 1912 [synonym], Canis niger Sclater 1874 [synonym], Canis nubilus Say 1823 [synonym], Canis occidentalis Richardson 1829 [synonym], Canis pallipes Sykes 1831 [synonym], Canis pambasileus Elliot 1905 [synonym], Canis petrolei Stock 1938 [synonym], Canis tundrarum Miller 1912 [synonym], Canis variabilis Wied-Neuwied 1841 [synonym], Lupus altaicus Noak 1911 [synonym], Lupus filchneri Matschie 1908 [synonym], Lupus gigas Townsend 1850 [synonym], Lupus karanorensis Matschie 1908 [synonym], Lupus laniger Hodgson 1847 [synonym], Lupus orientalis Wagner 1841 [synonym], Lupus tschiliensis Matschie 1908 [synonym]
†Canis lydekkeri Ameghino 1889
†Canis megamastoides Pomel 1842
†Canis michauxi Martin 1971
†Canis monticinensis Rook 1992
†Canis mosbachensis Soergel 1925
+
+
Invalid names: Canis cyonoides Pei 1934 [synonym]
†Canis osorum Cope 1873
†Canis othmanii Amri et al. 2017
†Canis palaeoplatensis Ameghino 1891
†Canis palmidens Teilhard de Chardin and Piveteau 1930
†Canis proplatensis Ameghino 1891
Canis rufus Audubon and Bachman 1851 [red wolf]
+
Invalid names: Canis floridanus Miller 1912 [synonym], Canis priscolatrans Cope 1899 [synonym], Lupus niger Bartram 1791 [synonym]
Canis sechurae Thomas 1900 [Sechuran fox]
†Canis senezensis Martin 1971
Canis simensis Rüppell 1835 [Ethiopian wolf]
†Canis tarijensis Ameghino 1902
†Canis teilhardi Qiu et al. 2004
†Canis thooides Tedford et al. 2009
†Canis troglodytes Lund 1938
+
Invalid names: Canis spelaeus Lund 1837 [replaced]
†Canis variabilis Pei 1934
Canis vetulus Lund 1842 [Hoary fox]
Invalid names: Lyciscus Hamilton-Smith 1839 [synonym], Nyctereutes terblanchei Broom 1948 [nomen dubium], Pachycyon Allen 1885 [synonym]
G. Chrysocyon Hamilton-Smith 1839 [maned wolf]
+
Chrysocyon brachyurus Illiger 1815 [maned wolf]
+
Invalid names: Canis isodactylus Ameghino 1906 [synonym], Canis jubatus Desmarest 1820 [synonym], Vulpes cancrosa Oken 1816 [synonym]
†Chrysocyon nearcticus Tedford et al. 2009
G. †Leptocyon Matthew 1918
+
†Leptocyon delicatus Loomis 1932
†Leptocyon douglassi Tedford et al. 2009
†Leptocyon gregorii Matthew 1907
†Leptocyon leidyi Tedford et al. 2009
†Leptocyon matthewi Tedford et al. 2009
†Leptocyon mollis Merriam 1906
†Leptocyon tejonensis Tedford et al. 2009
†Leptocyon vafer Leidy 1858
†Leptocyon vulpinus Matthew 1907
Invalid names: Neocynodesmus Macdonald 1963 [synonym]
G. Lycalopex Burmeister 1854 [South American fox]
+
Lycalopex culpaeus Molina 1782 [culpeo]
†Lycalopex ensenadensis Ameghino 1888
Lycalopex fulvipes Martin 1837 [Darwin's fox]
Lycalopex griseus Gray 1837 [South American gray fox]
Lycalopex gymnocercus Fischer 1814 [pampas fox]
Lycalopex sechurae Thomas 1900 [Sechuran fox]
Lycalopex vetulus Lund 1842 [Hoary fox]
+
Invalid names: Canis fulvicaudus Lund 1843 [synonym], Canis parvidens Mivart 1890 [synonym], Canis urostictus Mivart 1890 [synonym], Eunothocyon sladeni Thomas 1904 [synonym], Pseudalopex vetulus Lund 1842 [synonym]
Invalid names: Eunothocyon Allen 1905 [synonym], Pseudalopex Burmeister 1856 [synonym]
G. Nyctereutes Temminck 1839 [raccoon dog]
+
†Nyctereutes abdeslami Geraads 1997
†Nyctereutes barryi Werdelin and Dehghani 2010
†Nyctereutes donnezani Depérer 1890
†Nyctereutes lockwoodi Geraads et al. 2010
+
Invalid names: Canis borbonicus Gervais 1859 [objective synonym], Canis mitzopouli Paraskevaidis 1953 [synonym]
Nyctereutes procyonoides Gray 1834 [raccoon dog]
†Nyctereutes sinensis Schlosser 1903
†Nyctereutes tingi Tedford and Qiu 1991
†Nyctereutes vulpinus Soria and Aguirre 1976
G. †Protocyon Giebel 1855
+
†Protocyon orocualensis Ruiz-Ramoni et al. 2022
†Protocyon scagliarum Kraglievich 1952
†Protocyon tarijensis Ameghino 1902
†Protocyon troglodytes Lund 1938
+
Invalid names: Canis lycodes Lund 1843 [synonym], Canis musculosus Ameghino 1882 [synonym], Palaeocyon validus Lund 1843 [synonym], Paleospeothos colomae Spillmann 1942 [synonym], Protocyon orcesi Hoffstetter 1952 [synonym]
Invalid names: Paleospeothos Spillmann 1941 [nomen nudum], Pleurocyon Mercerat 1917 [synonym]
G. †Theriodictis Mercerat 1891
+
†Theriodictis floridanus Tedford et al. 2009
†Theriodictis platensis Mercerat 1891
+
Invalid names: Macrocyon morenoi Lydekker 1894 [synonym], Palaeocyon troglodytes Revilliod 1926 [synonym]
†Theriodictis tarijensis Ameghino 1902
Invalid names: Dinocynops Ameghino 1898 [synonym]
Tr. Vulpini Hemprich and Ehrenberg 1832
+
G. †Ferrucyon Ruiz-Ramoni et al. 2020
+
†Ferrucyon avius Torres and Ferrusquia 1981
G. †Metalopex Tedford and Wang 2008
+
†Metalopex bakeri Tedford et al. 2009
†Metalopex macconnelli Tedford et al. 2009
†Metalopex merriami Tedford and Wang 2008
G. Otocyon Müller 1835 [bat-eared fox]
+
Otocyon megalotis Desmarest 1822 [bat-eared fox]
Invalid names: Agriodus Smith 1840 [objective synonym]
G. †Prototocyon Pohle 1928
+
†Prototocyon recki Pohle 1928
G. Urocyon Baird 1857 [gray fox]
+
Urocyon cinereoargenteus Schreber 1775 [gray fox]
+
Invalid names: Canis virginianus Schreber 1775 [synonym], Urocyon californicus Merriam 1899 [synonym], Urocyon guatemalae Miller 1899 [synonym], Urocyon parvidens Miller 1899 [synonym], Urocyon seminolensis Simpson 1929 [synonym]
†Urocyon citrinus Tedford et al. 2009
†Urocyon galushai Tedford et al. 2009
+
Invalid names: Urocyon atwaterensis Getz 1960 [synonym]
Urocyon littoralis Baird 1857 [island gray fox]
†Urocyon minicephalus Martin 1974
†Urocyon progressus Stevens 1965
†Urocyon webbi Tedford et al. 2009
G. †Chailicyon Chow 1975
+
†Chailicyon crassidens Chow 1975
G. †Cubacyon Arredondo and Varona 1974
+
†Cubacyon transversidens Arredondo and Varona 1974
G. Cuon Hodgson 1838 [dhole]
+
Cuon alpinus Pallas 1811 [dhole]
†Cuon alpinus europaeus Bourguignat 1875
†Cuon bourreti Harlé 1891
†Cuon dukhunensis Sykes 1831
†Cuon majori Del Campana 1913
†Cuon priscus Thenius 1954
†Cuon sumatrensis Hardwicke 1822
Invalid names: Cyon Agassiz 1846 [objective synonym]
G. †Cynotherium Studiati 1857
+
†Cynotherium malatestai Madurell-Malapeira et al. 2015
†Cynotherium sardous Studiati 1857
G. Dysodus Cope 1879
+
Dysodus pravus Cope 1879 [Japanese sleeve dog]
Subfm. †Hesperocyoninae Tedford 1978
+
G. †Caedocyon Wang 1994
+
†Caedocyon tedfordi Wang 1994
G. †Cynodesmus Scott 1893
+
†Cynodesmus martini Wang 1994
†Cynodesmus thooides Scott 1893
+
Invalid names: Brachyrhynchocyon douglasi Macdonald 1970 [synonym], Mesocyon drummondanus Douglass 1903 [synonym], Sunkahetanka sheffleri Macdonald 1967 [synonym]
G. †Ectopocynus Wang 1994
+
†Ectopocynus antiquus Wang 1994
G. †Enhydrocyon Cope 1879
+
†Enhydrocyon basilatus Cope 1879
†Enhydrocyon crassidens Matthew 1907
†Enhydrocyon pahinsintewakpa Macdonald 1963
+
Invalid names: Enhydrocyon oregonensis Thorpe 1922 [synonym]
Invalid names: Hyaenocyon Cope 1879 [synonym], Hyaenocyon sectorius Cope 1883 [nomen vanum]
G. †Hesperocyon Scott 1890
+
†Hesperocyon gregarius Cope 1873
+
Invalid names: Amphicyon gracilis Leidy 1856 [synonym], Cynodictis hylactor Hay 1899 [synonym], Cynodictis paterculus Matthew 1903 [synonym], Nanodelphys mcgrewi Russell 1972 [synonym]
Invalid names: Alloeodectes Russell 1984 [synonym], Pseudocynodictis Schlosser 1902 [synonym]
G. †Mesocyon Scott 1890
+
†Mesocyon brachyops Merriam 1906
+
Invalid names: Mesocyon baileyi Stock 1933 [synonym]
†Mesocyon coryphaeus Cope 1879
†Mesocyon temnodon Wortman and Matthew 1899
Invalid names: Hypotemnodon Eyerman 1894 [synonym], Mesocyon josephi secundus Thorpe 1922 [invalid subgroup]
G. †Osbornodon Wang 1994
+
†Osbornodon brachypus Cope 1881
†Osbornodon fricki Wang 1994
†Osbornodon iamonensis Sellards 1916
+
Invalid names: Cynodesmus nobilis Simpson 1932 [synonym], Paradaphoenus tropicalis White 1942 [synonym]
†Osbornodon renjiei Wang 1994
†Osbornodon scitulus Hay 1924
†Osbornodon sesnoni Macdonald 1967
†Osbornodon wangi Hayes 2000
G. †Paraenhydrocyon Wang 1994
+
+
Invalid names: Mesocyon hortulirosae Schlaikjer 1935 [synonym]
†Paraenhydrocyon robustus Matthew 1907
+
Invalid names: Temnocyon venator Cook 1909 [synonym]
G. †Philotrox Merriam 1906
+
†Philotrox condoni Merriam 1906
G. †Prohesperocyon Wang 1994
+
†Prohesperocyon wilsoni Gustafson 1986
G. †Sunkahetanka Macdonald 1963
+
†Sunkahetanka geringensis Barbour and Schultz 1935
G. †Hydrocyon Lartet 1851
G. †Icticyon Lund 1839
G. Lupulella Hilzheimer 1906
+
Lupulella mesomelas Schreber 1775 [black-backed jackal]
†Lupulella mohibi Geraads 2011
†Lupulella paralius Geraads 2011
G. †Lupus
G. Lycaon Brookes 1827
+
†Lycaon magnus Ewer and Singer 1956
Lycaon pictus Temminck 1820 [African wild dog]
†Lycaon sekowei Hartstone-Rose et al. 2010
G. †Lycorus Bourguignat 1875
+
†Lycorus nemesianus Bourguignat 1875
G. Megalotis Smith 1839
G. †Palaeocyon Lund 1843
G. †Plesiocyon Schlosser 1887
G. Schaeffia Hilzheimer 1906
+
Schaeffia adusta Sundevall 1847 [side-striped jackal]
G. Simenia Gray
+
Simenia simensis Rüppell 1835 [Ethiopian wolf]
G. †Synagodus Cope 1879
+
†Synagodus mansuetus Cope 1879
G. Thous Hamilton Smith 1839
+
Thous fulvipes Martin 1837 [Darwin's fox]
G. Vulpes Frisch 1775 [fox]
+
†Vulpes alopecoides Forsyth Major 1877
†Vulpes angustidens Thenius 1954
†Vulpes beihaiensis Qiu and Tedford 1990
Vulpes bengalensis Shaw 1800 [Bengal fox]
Vulpes cana Blanford 1877 [Blanfords fox]
Vulpes chama Smith 1833 [Cape fox]
†Vulpes chikushanensis Young 1930
Vulpes corsac Linnaeus 1768 [Corsac fox]
Vulpes ferrilata Hodgson 1842 [Tibetan fox]
†Vulpes galaticus Ginsburg 1998
Vulpes griffithi Blyth 1854
†Vulpes hassani Geraads 2011
Vulpes japonica Ognev 1823
Vulpes karagan Erxleben 1777
†Vulpes kernensis Tedford et al. 2009
Vulpes lagopus Linnaeus 1758 [Arctic fox]
Vulpes macrotis Merriam 1888 [kit fox]
†Vulpes mathisoni Geraads et al. 2015
Vulpes montana Pearson 1836
Vulpes niloticus Geoffroy Saint-Hilaire 1803
†Vulpes odessana Odintzov 1967
Vulpes pallida Cretzschmar 1826 [pale fox]
Vulpes pensylvanicus Boddaert 1784
†Vulpes praecorsac Kormos 1932
†Vulpes praeglacialis Kormos 1932
+
Invalid names: Vulpes marinosi Sickenberg 1971 [synonym]
Vulpes pusilla Blyth 1854
†Vulpes riffautae de Bonis et al. 2007
†Vulpes rooki Bartolini Lucenti 2021
Vulpes rueppellii Schinz 1825 [Ruuppells fox]
†Vulpes sinensis Schlosser 1903
†Vulpes skinneri Hartstone-Rose et al. 2013
†Vulpes stenognathus Savage 1941
Vulpes velox Say 1823 [swift fox]
Vulpes vulgaris Oken 1816
Vulpes vulpes Linnaeus 1758 [red fox]
+
Invalid names: Canis alopex Linnaeus 1758 [synonym], Canis vulpes montana Pearson 1835 [synonym], Vulpes fulva Desmarest 1829 [synonym], Vulpes palmaria Hay 1917 [synonym], Vulpes vulpes cascadensis Merriam 1900 [replaced]
Vulpes zerda Zimmermann 1780 [fennec fox]
Invalid names: Alopex Kaup 1829 [synonym], Fennecus Desmarest 1804 [synonym]
Invalid names: Amphicyon angustidens Marsh 1871 [nomen vanum], Otocyoninae Trouessart 1885 [empty]
Subor. Caniformia Kretzoi 1943
+
Fm. †Amphicyonidae Haeckel 1886 [bear-dog]
+
G. †Amphicyanis Springhorn 1977
Subfm. †Amphicyoninae Trouessart 1885
+
G. †Afrocyon Arambourg 1961
+
†Afrocyon burolleti Arambourg 1961
†Afrocyon macrodon Savage 1965
G. †Amphicyon Lartet 1836
+
Subg. †Amphicyon (Euroamphicyon) Ginsburg 1999
†Amphicyon ambiguus Filhol 1877
†Amphicyon argentinus Ameghino 1904
†Amphicyon frendens Matthew 1924
†Amphicyon galushai Hunt, Jr. 2003
†Amphicyon giganteus Schinz 1825
†Amphicyon gracilis Pomel 1847
†Amphicyon ingens Matthew 1924
†Amphicyon intermedius White 1940
†Amphicyon lathanicus Ginsburg 2000
†Amphicyon laugnacensis Ginsburg 1999
†Amphicyon longiramus White 1942
†Amphicyon major De Blainville 1841
+
Invalid names: Amphicyon cultridens Pomel 1853 [objective synonym]
†Amphicyon olisiponensis Antunes and Ginsburg 1977
†Amphicyon palaeindicus Falconer and Lydekker 1876
†Amphicyon pontoni Simpson 1930
†Amphicyon reinheimeri Cook 1926
†Amphicyon riggsi McGrew 1939
†Amphicyon zhanxiangi Jiangzuo et al. 2018
Tr. †Amphicyonini Trouessart 1885
+
G. †Cynelos Jourdan 1862
+
Invalid names: Absonodaphoenus Olsen 1958 [synonym], Hecubides Savage 1965 [synonym]
G. †Paludocyon Morales et al. 2021
G. †Goupilictis Ginsburg 1969
+
†Goupilictis minor Dehm 1935
G. †Ischyrocyon Matthew and Gidley 1904
+
†Ischyrocyon gidleyi Matthew 1902
+
Invalid names: Hadrocyon mohavensis Stock and Furlong 1926 [synonym], Ischyrocyon hyaenodus Matthew and Gidley 1904 [synonym], Pliocyon walkerae Johnston and Christian 1941 [synonym]
Invalid names: Hadrocyon Stock and Furlong 1926 [synonym]
G. †Magericyon Peigné et al. 2008
+
†Magericyon anceps Peigné et al. 2008
†Magericyon castellanus Fraile 1997
G. †Megamphicyon Kuss 1965
+
G. †Mogharacyon Morales and Pickford 2022
+
†Mogharacyon anubisi Morlo and Miller 2019
G. †Myacyon Sudre and Hartenberger 1992
+
†Myacyon kiptalami Morales and Pickford 2005
†Myacyon peignei Werdelin 2019
G. †Namibiocyon Morales and Pickford 2022
+
†Namibiocyon ginsburgi Morales et al. 1998
G. †Pliocyon Matthew 1918
+
†Pliocyon medius Matthew 1918
†Pliocyon ossifragus Douglass 1903
†Pliocyon robustus Berta and Galiano 1984
Tr. †Pseudarctini Morales et al. 2021
+
G. †Dehmicyon Morales et al. 2021
G. †Ictiocyon Crusafont Pairo et al. 1955
G. †Pseudarctos Schlosser 1899
G. †Pseudocyon Lartet 1851
+
†Pseudocyon caucasicus Gabunia 1973
†Pseudocyon sansaniensis Lartet 1851
Invalid names: Amphicyonopsis Viret 1929 [synonym]
G. †Tartarocyon Solé et al. 2022
+
†Tartarocyon cazanavei Solé et al. 2022
G. †Vishnucyon Pilgrim 1932
Invalid names: Thaumastocyonini Ginsburg 1977 [empty]
G. †Angelarctocyon Tomiya and Tseng 2016
+
†Angelarctocyon australis Gustafson 1986
G. †Arctamphicyon Pilgrim 1932
G. †Aymardia Trouessart 1904
G. †Bonisicyon Werdelin 2009
+
†Bonisicyon illacabo Werdelin 2009
G. †Brachycyon Filhol 1872
+
†Brachycyon gaudryi Ginsburg 1966
G. †Cynodictis Bravard and Pomel 1850
+
Subg. †Cynodictis (Elocyon) Aymard 1849
†Cynodictis cayluxensis Filhol 1876
†Cynodictis crassus Teilard de Chardin 1915
†Cynodictis exilis Teilard de Chardin 1915
†Cynodictis ferox Filhol 1876
†Cynodictis lacustris Gervais 1852
+
Invalid names: Cynodictis intermedius Filhol 1876 [synonym]
†Cynodictis longirostris Filhol 1872
†Cynodictis parisiensis Pomel 1853
†Cynodictis peignei Le Verger et al. 2020
G. †Cyotherium Aymard
Subfm. †Daphoeninae Hunt, Jr. 1998
+
G. †Adilophontes Hunt, Jr. 2002
+
†Adilophontes brachykolos Hunt, Jr. 2002
G. †Brachyrhynchocyon Scott and Jepsen 1936
+
+
Invalid names: Daphoenocyon minor Clark 1967 [synonym], Daphoenus canadensis Russell 1934 [synonym], Miacis matthewi Schlaikjer 1935 [synonym]
†Brachyrhynchocyon montanus Hunt, Jr. 1996
Invalid names: Brachicyon Loomis 1931 [replaced], Daphoenocyon Hough 1948 [synonym]
G. †Daphoenictis Hunt, Jr. 1974
+
†Daphoenictis tedfordi Hunt, Jr. 1974
G. †Daphoenodon Peterson 1909
+
Subg. †Daphoenodon (Borocyon) Peterson 1910
†Daphoenodon falkenbachi Hunt, Jr. 2002
†Daphoenodon notionastes Frailey 1979
†Daphoenodon skinneri Hunt, Jr. 2002
†Daphoenodon superbus Peterson 1907
+
Invalid names: Daphoenodon periculosus Cook 1909 [synonym]
G. †Daphoenus Leidy 1853
+
†Daphoenus demilo Dawson 1980
†Daphoenus felinus Scott 1898
†Daphoenus lambei Russell 1934
†Daphoenus robustus Scott 1897
†Daphoenus ruber Stock 1932
†Daphoenus socialis Thorpe 1922
†Daphoenus transversus Wortman and Matthew 1899
†Daphoenus vetus Leidy 1853
+
Invalid names: Proamphicyon nebrascensis Hatcher 1902 [synonym]
Invalid names: Canis lippincottianus Cope 1873 [nomen dubium], Pericyon Thorpe 1922 [synonym], Proamphicyon Hatcher 1902 [synonym]
G. †Paradaphoenus Wortman and Matthew 1899
+
+
Invalid names: Amphicyon entoptychi Cope 1879 [synonym]
†Paradaphoenus minimus Hough 1948
†Paradaphoenus tooheyi Hunt, Jr. 2001
G. †Guangxicyon Zhai et al. 2003
+
†Guangxicyon sinoamericanus Zhai et al. 2003
G. †Gustafsonia Tomiya and Tseng 2016
+
†Gustafsonia cognita Gustafson 1986
Subfm. †Haplocyoninae
+
Tr. †Aktaucyonini Kordikova et al. 2000
+
G. †Aktaucyon Kordikova et al. 2000
G. †Gobicyon Colbert 1839
G. †Haplocyon Schlosser 1901
+
†Haplocyon crucians Filhol 1879
†Haplocyon dombrowski Helbing 1928
†Haplocyon elegans de Bonis 1966
G. †Haplocyonoides Hürzeler 1940
+
†Haplocyonoides mordax Hürzeler 1940
G. †Haplocyonopsis de Bonis 1973
G. †Sarcocyon Ginsburg 1966
G. †Harpagophagus De Bonis 1971
+
†Harpagophagus sanguinensis De Bonis 1971
G. †Notoamphicyon Ameghino 1904
+
†Notoamphicyon paranensis Ameghino 1904
G. †Protemnocyon Hatcher 1901
+
†Protemnocyon inflatus Hatcher 1902
G. †Pseudamphicyon Schlosser 1887
G. †Pseudocyonopsis Kuss 1965
+
†Pseudocyonopsis ambiguus Ginsburg 1965
†Pseudocyonopsis antiquus Ginsburg 1966
†Pseudocyonopsis quercensis Ginsburg 1965
G. †Storchictis de Bonis 2020
+
†Storchictis miacinus de Chardin 1915
G. †Symplectocyon Springhorn 1979
Subfm. †Temnocyoninae Hunt, Jr. 1998
+
G. †Delotrochanter Hunt, Jr. 2011
+
†Delotrochanter major Hunt, Jr. 2011
†Delotrochanter oryktes Hunt, Jr. 2011
†Delotrochanter petersoni Hunt, Jr. 2011
G. †Mammacyon Loomis 1936
+
†Mammacyon ferocior Hunt, Jr. 2011
†Mammacyon obtusidens Loomis 1936
G. †Rudiocyon Hunt, Jr. 2011
+
†Rudiocyon amplidens Hunt, Jr. 2011
G. †Temnocyon Cope 1878
+
†Temnocyon altigenis Cope 1878
†Temnocyon ferox Eyermann 1896
†Temnocyon fingeruti Hunt, Jr. 2011
†Temnocyon macrogenys Hunt, Jr. 2011
†Temnocyon percussor Cook 1909
†Temnocyon subferox Hunt, Jr. 2011
†Temnocyon typicus Loomis 1936
Subfm. †Thaumastocyoninae Hurzeller 1940
+
G. †Agnotherium Kaup 1833
+
†Agnotherium antiquum Kaup 1832
+
Invalid names: Amphicyon agnotus Pomel 1853 [synonym]
†Agnotherium grivense Viret 1929
G. †Ammitocyon Morales et al. 2021
+
†Ammitocyon kainos Morales et al. 2021
G. †Crassidia Heizmann and Kordikova 2000
+
†Crassidia intermedia von Meyer 1849
G. †Peignecyon Morales et al. 2019
+
†Peignecyon felinoides Morales et al. 2019
G. †Thaumastocyon Stehlin and Helbing 1925
+
†Thaumastocyon bourgeoisi Stehlin and Helbing 1925
†Thaumastocyon dirus Ginsburg et al. 1981
G. †Tomocyon Viret 1929
+
†Tomocyon grivensis Viret 1929
G. †Ysengrinia Ginsburg 1965
+
†Ysengrinia americana Wortman 1901
†Ysengrinia depereti Mayet 1908
†Ysengrinia gerandiana Viret 1929
†Ysengrinia tolosana Noulet 1876
†Ysengrinia valentiana Belinchon and Morales 1989
Invalid names: Aphicyoninae Trouessart 1885 [empty]
Infraor. Canoidea Simpson 1931
+
Superfm. Arctoidea Flower 1869
+
G. †Allocyon Merriam 1930
+
†Allocyon loganensis Merriam 1930
G. †Amphicticeps Matthew and Granger 1924
+
†Amphicticeps dorog Wang et al. 2005
†Amphicticeps makhchinus Wang et al. 2005
†Amphicticeps shackelfordi Matthew and Granger 1924
Unr. Mustelida Tedford 1976
+
Fm. Ailuridae Bonaparte 1850 [red panda]
G. †Bavarictis Mödden 1991
G. †Chaprongictis Peigné et al. 2006
Fm. Mephitidae Bonaparte 1845 [skunk]
G. †Mustelictis Lange 1969
Fm. Mustelidae Fischer 1817 [mustelid carnivore]
G. †Peignictis de Bonis et al. 2019
Fm. Procyonidae Gray 1825 [procyonid carnivore]
+
Invalid names: Cercoleptidae Bonaparte 1850 [synonym]
G. †Pseudobassaris Pohle 1917
Invalid names: Oligobunidae [empty], Procyonoidea Bonaparte 1850 [empty]
Superfm. Musteloidea Fischer de Waldheim 1817
+
G. †Proputorius Filhol 1890
Invalid names: Semantoridae Orlov 1932 [empty]
G. †Nothocyon Wortman and Matthew 1899
+
Subg. †Nothocyon (Galecynus) Owen 1847
Unr. †Panpinnipedia Wolsan et al. 2020
+
Unr. Pinnipedimorpha Berta et al. 1989 [pinniped]
G. †Potamotherium Geoffroy 1833
G. †Puijila Rybczynski et al. 2009
Unr. Ursida Tedford 1976
+
Superfm. Ursoidea Fischer de Waldheim 1817
Invalid names: Arctomorpha Wolson 1993 [empty]
G. †Lycophocyon Tomiya 2011
+
†Lycophocyon hutchisoni Tomiya 2011
†Lycophocyon tabrumi Lofgren et al. 2020
Invalid names: Cynoidea Flower 1869 [empty]
Subor. Feliformia Kretzoi 1945
+
Infraor. Aeluroida Flynn and Galiano 1982
+
G. †Ictidopappus Simpson 1935
+
†Ictidopappus mustelinus Simpson 1935
Invalid names: Viverravoidea Wortman and Matthew 1899 [empty]
Unr. Aeluroidea Flower 1869
+
G. †Anictis Kretzoi 1945
+
†Anictis simplicidens Schlosser 1890
G. †Fejfarictis de Bonis et al. 2024
+
†Fejfarictis valecensis de Bonis et al. 2024
G. †Haplogale
+
†Haplogale media Filhol 1882
G. †Herpestides de Beaumont 1967
+
†Herpestides aegypticus Morlo et al. 2007
†Herpestides aequatorialis Schmidt-Kittler 1987
†Herpestides antiquus de Blainville 1842
†Herpestides compactus Morlo 1996
G. †Palaeoprionodon Filhol 1880
+
G. †Shandgolictis Hunt, Jr. 1998
+
†Shandgolictis constans Matthew and Granger 1924
G. †Stenoplesictis Filhol 1880
+
†Stenoplesictis cayluxi Filhol 1880
†Stenoplesictis crocheti Peigné and de Bonis 1999
†Stenoplesictis elegans Matthew and Granger 1924
†Stenoplesictis minor Filhol 1882
†Stenoplesictis muhoronii Schmidt-Kittler 1987
Infraor. Viverroidea Gray 1821
+
Fm. Eupleridae Chenu 1850
+
G. Cryptoprocta Bennett 1833 [fossa]
G. Eupleres Doyère 1835
G. Fossa Gray 1864 [Malagasy civet]
Subfm. Galidiinae Gray 1864
Fm. †Herpestidae Bonaparte 1845 [mongoose]
+
G. Bdeogale Peters 1850
G. Crossarchus Cuvier 1825 [dwarf mongoose]
G. Cynictis Ogilby 1833 [yellow mongoose]
G. Dologale Thomas 1920
G. Galerella Gray 1864 [slender mongoose]
G. Helogale Gray 1861 [dwarf mongoose]
G. Herpestes Illliger 1811
+
Invalid names: Calictis Gray 1864 [synonym], Calogale Gray 1864 [synonym], Onychogale Gray 1864 [synonym], Taeniogale Gray 1864 [synonym], Urva Hodgson 1837 [synonym]
Subfm. Herpestinae Bonaparte 1845
G. Ichneumia Geoffroy 1837
+
Invalid names: Herpestes Cuvier 1829 [replaced]
G. †Legetetia Schmidt-Kittler 1987
G. Liberiictis Hayman 1958
G. Mungos Geoffroy Saint Hilaire and Cuvier 1795
+
Invalid names: Ariela Gray 1864 [synonym]
G. †Paracynictis Pocock 1916
G. Rhynchogale Thomas 1894
+
Invalid names: Rhinogale Gray 1865 [replaced]
G. Suricata Desmarest 1804 [meerkat]
+
Invalid names: Ryzaena Illiger 1811 [synonym]
G. †Ugandictis Morales et al. 2007
G. Xenogale Allen 1919
Fm. Hyaenidae Gray 1821 [hyaena]
+
G. †Adcrocuta Kretzoi 1938
+
Invalid names: Hyena [replaced]
G. †Allohyaena Kretzoi 1938
+
Invalid names: Xenohyaena Kretzoi 1938 [synonym]
G. †Belbus Werdelin and Solunias 1991
G. †Chasmaporthetes Hay 1921
+
Invalid names: Ailuraena Stirton and Christian 1940 [synonym]
G. Crocuta Erxleben 1777 [spotted hyaena]
G. †Euryboas Schaub 1941
G. Hyaena Brisson 1762
+
Invalid names: Euhyaena Falconer 1868 [objective synonym]
G. †Hyaenictis Gaudry 1861
G. †Hyaenictitherium Kretzoi 1938
G. †Hyaenotherium Semenov 1989
Subfm. †Ictitheriinae Dietrich 1927
G. †Ictitherium Wagner 1848
G. †Ikelohyaena Werdelin and Solunias 1991
G. †Leecyaena Young and Liu 1948
G. †Lepthyaena Lydekker 1884
G. †Lycaena Hensel 1863
G. †Lycyaena Hensel 1863
G. †Lycyaenops Kretzoi 1938
G. †Mesoviverrops de Beaumont and Mein 1972
G. †Miohyaenotherium Semenov 1989
G. Pachycrocuta Kretzoi 1938 [brown hyaena]
G. †Palinhyaena Qiu et al. 1979
G. Parahyaena Hendey 1974
G. †Pliocrocuta Kretzoi 1938
G. †Plioviverrops Kretzoi 1938
G. Proteles Geoffroy Saint-Hilaire 1824
+
Invalid names: Geocyon Wagler 1830 [synonym]
G. †Protoviverrops de Beaumont and Mein 1972
G. †Thalassictis Nordmann 1850
+
Invalid names: Miohyaena Kretzoi 1938 [synonym], Palhyaena Gervais 1859 [synonym]
G. †Tongxinictis Werdelin and Solunias 1991
G. †Tungurictis Colbert 1939
Invalid names: Hyaeninae Mivart 1882 [empty], Protelidae Flower 1869 [synonym]
Fm. Viverridae Gray 1821 [civet]
+
G. †Africanictis Morales et al. 1998
G. †Euboictis Fejfar and Schmidt-Kittler 1984
Subfm. Galidictinae Mivart 1882
Subfm. Genettinae Gray 1864
Subfm. Hemigalinae Gray 1864
G. †Kanuites Dehghani and Werdelin 2008
G. †Ketketictis Morlo et al. 2007
G. †Leptoplesictis Forsyth-Major 1903
Subfm. †Lophocyoninae Fejfar and Schmidt-Kittler 1987
G. †Megaviverra Qiu 1980
G. †Mioprionodon Schmidt-Kittler 1987
Subfm. Paradoxurinae Gray 1864
G. †Progenetta Deperet 1892
G. †Sahelictis Peigné 2008
G. †Vishnuictis Pilgrim 1932
Subfm. Viverrinae Gray 1821
Invalid names: Arctogalidiinae Pocock 1933 [empty], Cryptoproctinae Gray 1865 [empty], Cynogalinae Gray 1864 [empty], Herpestina Bonaparte 1850 [empty], Hyaenina Gray 1825 [empty], Prionodontinae Gray 1864 [invalid subgroup]
Invalid names: Stenoplesictinae Schlosser 1923 [invalid subgroup]
G. †Alagtsavbaatar Egi et al. 2016
+
†Alagtsavbaatar indigenus Dashzeveg 1996
G. †Asiavorator Spassov and Lange-Badré 1995
+
†Asiavorator gracilis Matthew and Granger 1924
+
Invalid names: Asiavorator altidens Spassov and Lange-Badré 1995 [synonym], Stenoplesictis simplex Dashzeveg 1996 [synonym]
Superfm. Feloidea Hay 1930
+
Fm. Felidae Gray 1821 [cat]
+
G. †Aeluropsis Lydekker 1884
G. Ailurin Gervais 1855
G. Catolynx Severtzov 1858
G. Cynaelurus Gloger 1841
G. Cynailurus Wagler
G. †Diamantofelis Morales et al. 1998
+
†Diamantofelis ferox Morales et al. 1998
†Diamantofelis minor Morales et al. 1998
G. †Drepanodon Nesti 1826
G. †Epimachairodus Kretzoi 1929
Subfm. Felinae Trouessart 1885
+
G. Acinonyx Brookes 1828 [cheetah]
G. Caracal Gray 1843
G. Catopuma Severtzov 1858
G. Felis Linnaeus 1758 [cat]
G. Herpailurus Severtzov 1858
G. Leopardus Gray 1842
G. Leptailurus Severtzov 1858
G. †Leptofelis Salesa et al. 2017
G. Lynx Kerr 1792 [lynx]
+
Invalid names: Lyncus Gray 1825 [synonym]
G. †Magerifelis Salesa et al. 2024
G. †Miopanthera Kretzoi 1938
G. †Miracinonyx Adams 1979 [American cheetah]
G. Neofelis Gray 1867 [clouded leopard]
G. Otocolobus Brandt 1842
G. Panthera Oken 1816
+
Invalid names: Leo Brehm 1829 [synonym], Tigris Gray 1843 [synonym]
G. Pardofelis Severtzov 1858
G. †Pratifelis Hibbard 1934
G. Prionailurus Severtzov 1858
G. †Pristifelis Salesa et al. 2012
G. Puma Jardine 1834 [cougar]
G. †Sivapanthera Kretsoi
G. †Styriofelis Kretzoi 1929
Invalid names: Felini Gray 1825 [empty]
G. †Hyperailurictis Kretzoi 1929
G. †Jansofelis Bonifay 1971
+
†Jansofelis vaufreyi Bonifay 1971
G. †Katifelis Adrian et al. 2018
+
†Katifelis nightingalei Adrian et al. 2018
G. †Machaerodus Kaup 1833
+
†Machaerodus ensenadensis Ameghino 1888
†Machaerodus niobrarensis Thorpe 1922
Subfm. †Machairodontinae Gill 1872
+
G. †Dinofelis Zdansky 1924
G. †Hemimachairodus Koeningswald 1974
Tr. †Machairodontini Gill 1872
G. †Paramachaerodus Pilgrim 1913
+
Invalid names: Promegantereon Kretzoi 1938 [synonym]
Tr. †Smilodontini Kretzoi 1929
G. †Mellivorodon Lydekker 1884
G. †Namafelis Morales et al. 2003
G. Plethaelurus Cope 1882
G. †Proailurus Filhol 1879
+
†Proailurus lemanensis Filhol 1879
G. †Profelis Severtzov 1858
G. †Propontosmilus Kretzoi 1929
G. †Pseudaelurus Leidy 1858
+
†Pseudaelurus aeluroides Macdonald 1954
†Pseudaelurus africanus Andrews 1914
†Pseudaelurus cuspidatus Wang et al. 1998
†Pseudaelurus edwardsi Filhol 1872
†Pseudaelurus guangheensis Cao et al. 1990
†Pseudaelurus marshi Thorpe 1922
†Pseudaelurus quadridentatus de Blainville 1843
†Pseudaelurus romieviensis Roman and Viret 1934
†Pseudaelurus skinneri Rothwell 2003
†Pseudaelurus stouti Schultz and Martin 1972
†Pseudaelurus validus Rothwell 2001
G. †Sivaelurus Pilgrim 1913
G. †Sivafelis Pilgrim 1932
G. †Sivasmilus Kretzoi 1929
G. Uncia Schreber 1775
G. †Vinayakia Pilgrim 1932
G. †Viretailurus Hemmer 1965
G. †Vishnufelis Pilgrim 1932
Invalid names: Pantherinae Pocock 1917 [empty]
Fm. Prionodontidae Horsfield 1824 [asiatic linsang]
+
G. Prionodon Horsfield 1822 [banded linsang]
+
Prionodon gracilis Horsfield 1824
Prionodon linsang Hardwicke 1821 [banded linsang]
+
Invalid names: Prionodon maculosus Blanford 1878 [synonym]
Prionodon pardicolor Hodgson 1842 [spotted linsang]
Invalid names: Linsang Müller 1839 [synonym], Linsanga Lydekker 1896 [synonym], Pardictis Thomas 1925 [synonym]
G. †Stenogale Schlosser 1888
+
†Stenogale bransatensis de Bonis et al. 1999
†Stenogale gracilis Filhol 1877
†Stenogale intermedia Filhol 1877
G. †Viretictis de Bonis et al. 1999
+
†Viretictis gaillardi Viret 1929
†Viretictis julieni Filhol 1879
Fm. Nandiniidae Pocock 1929
+
G. Nandinia Gray 1843 [African palm civet]
+
Nandinia binotata Gray 1832 [African palm civet]
+
Invalid names: Paradoxurus hamiltonii Gray 1852 [synonym]
Fm. †Nimravidae Cope 1880 [false sabre-tooth]
+
G. †Aelurogale Filhol 1872
Subfm. †Barbourofelinae Schultz et al. 1970
+
Tr. †Afrosmilini Morales et al. 2001
+
G. †Afrosmilus Kretzoi 1929
+
†Albanosmilus jourdani Filhol 1883
G. †Barbourofelis Schultz et al. 1970
+
†Barbourofelis fricki Schultz et al. 1970
†Barbourofelis loveorum Baskin 1981
†Barbourofelis morrisi Schultz et al. 1970
†Barbourofelis piveteaui Ozansoy 1965
†Barbourofelis vallesiensis de Beaumont and Crusafont-Pairo 1982
†Barbourofelis whitfordi Barbour and Cook 1915
+
Invalid names: Ischyrosmilus osborni Merriam 1919 [synonym]
G. †Ginsburgsmilus Morales et al. 2001
+
†Ginsburgsmilus napakensis Morales et al. 2001
G. †Oriensmilus Wang et al. 2020
+
†Oriensmilus liupanensis Wang et al. 2020
G. †Prosansanosmilus Heizmann et al. 1980
+
†Prosansanosmilus eggeri Morlo et al. 2004
†Prosansanosmilus peregrinus Heizmann et al. 1980
G. †Sansanosmilus Kretzoi 1929
+
†Sansanosmilus jourdani Filhol 1883
†Sansanosmilus palmidens Blainville 1841
G. †Syrtosmilus Ginsburg 1978
+
†Syrtosmilus syrtensis Ginsburg 1978
G. †Vampyrictis Kurten 1976
G. †Dinictis Leidy 1854
+
†Dinictis felina Leidy 1854
+
Invalid names: Daptophilus squalidens Cope 1873 [synonym], Dinictis bombifrons Adams 1895 [synonym], Dinictis cyclops Cope 1879 [synonym], Dinictis fortis Adams 1895 [synonym], Dinictis paucidens Riggs 1896 [synonym]
Invalid names: Daptophilus Cope 1873 [synonym]
Tr. †Hoplophoneini Kretzoi 1929
+
G. †Eusmilus Gervais 1876
G. †Hoplophoneus Cope 1874
+
†Hoplophoneus adelos Barrett 2021
†Hoplophoneus bidentatus Filhol 1872
+
Invalid names: Ekgmoiteptecela olsontau Macdonald 1963 [synonym], Hoplophoneus belli Stock 1933 [synonym]
†Hoplophoneus crassidens Cragin 1892
†Hoplophoneus dakotensis Hatcher 1895
+
Invalid names: Dinotomius atrox Williston 1895 [synonym]
†Hoplophoneus oharrai Jepsen 1926
†Hoplophoneus primaevus Leidy and Owen 1851
+
Invalid names: Hoplophoneus insolens Adams 1896 [synonym], Hoplophoneus latidens Thorpe 1920 [synonym], Hoplophoneus marshi Thorpe 1920 [synonym], Hoplophoneus mentalis Sinclair 1921 [synonym], Hoplophoneus molossus Thorpe 1920 [synonym], Hoplophoneus oreodontis Cope 1873 [synonym], Hoplophoneus robustus Adams 1896 [synonym]
†Hoplophoneus sicarius Sinclair and Jepsen 1927
†Hoplophoneus villebramarensis Peigné and Brunet 2001
Invalid names: Dinotomius Williston 1895 [synonym], Ekgmoiteptecela Macdonald 1963 [synonym]
G. †Nanosmilus Martin 1992
+
†Nanosmilus kurteni Martin 1992
G. †Maofelis Averianov et al. 2016
+
†Maofelis cantonensis Averianov et al. 2016
Subfm. †Nimravinae Trouessart 1885
+
G. †Dinailurictis Helbing 1922
+
†Dinailurictis bonali Helbing 1922
+
Invalid names: Ailurictis helbingi Kretzoi 1929 [synonym]
G. †Eofelis Kretzoi 1938
+
†Eofelis edwardsii Filhol 1872
†Eofelis giganteus Peigné 2000
G. †Quercylurus Ginsburg 1979
+
†Quercylurus major Ginsburg 1979
Invalid names: Hoplophoneinae [synonym]
Tr. †Nimravini Cope 1880
+
G. †Dinaelurus Eaton 1922
+
†Dinaelurus crassus Eaton 1922
G. †Nimravus Cope 1879
+
†Nimravus brachyops Cope 1878
+
Invalid names: Archaelurus debilis Cope 1879 [synonym], Dinictis major Lucas 1898 [synonym], Nimravus altidens Macdonald 1950 [synonym], Nimravus bumpensis Scott and Jepsen 1936 [synonym], Nimravus confertus Cope 1880 [synonym], Nimravus gomphodus Cope 1880 [synonym], Nimravus meridianus Stock 1933 [synonym], Nimravus sectator Matthew 1907 [synonym]
†Nimravus intermedius Filhol 1872
†Nimravus mongoliensis Gromova 1959
Invalid names: Archaelurus Cope 1879 [synonym], Nimravinus Kretzoi 1945 [synonym]
G. †Pangurban Poust et al. 2022
+
†Pangurban egiae Poust et al. 2022
G. †Pogonodon Cope 1880
+
†Pogonodon davisi Merriam 1906
+
Invalid names: Pogonodon eileenae Macdonald 1970 [synonym], Pogonodon serrulidens Eaton 1922 [synonym]
†Pogonodon platycopis Cope 1879
+
Invalid names: Pogonodon cismontanus Thorpe 1920 [synonym]
Invalid names: Eusmilus insignis Filhol 1876 [nomen nudum]
Fm. †Palaeogalidae Martin and Lim 2001
+
G. †Cryptailurus Martin and Lim 2001
+
†Cryptailurus tedfordi Martin and Lim 2001
†Cryptailurus tinaynakti Barrett et al. 2020
G. †Palaeogale von Meyer 1846
+
†Palaeogale dorothiae Macdonald 1963
†Palaeogale evanoffi Welsh 2021
†Palaeogale felina Filhol 1877
†Palaeogale hyaenoides Dehm 1950
†Palaeogale minuta Gervais 1848
†Palaeogale sanguinarius Loomis 1932
†Palaeogale sectoria Gervais 1848
+
Invalid names: Bunaelurus infelix Matthew 1903 [synonym], Bunaelurus parvulus Matthew and Granger 1924 [synonym], Bunaelurus ulysses Matthew and Granger 1924 [synonym]
Invalid names: Bunaelurus Cope 1873 [synonym]
Invalid names: Didymictida Flynn and Galiano 1982 [empty]
Subor. Fissipedia Blumenbach 1791
+
Invalid names: Arctictidae Cope 1889 [empty], Bassarididae Cope 1889 [empty], Cynictidae Cope 1882 [empty]
Superfm. †Miacoidea Cope
+
Fm. †Miacidae Cope 1880
+
G. †Eogale Beard and Dawson 2009
+
†Eogale parydros Beard and Dawson 2009
Subfm. †Miacinae Cope 1880
+
G. †Paroodectes Springhorn 1980
G. †Miocyon Matthew 1909
+
†Miocyon magnus Bryant 1992
†Miocyon major Matthew 1909
†Miocyon scotti Wortman and Matthew 1899
†Miocyon vallisrubrae Friscia and Rasmussen 2010
G. †Pappidictops Chiu and Li 1977
G. †Simamphicyon Viret 1942
+
†Simamphicyon helveticus Pictet and Humbert 1869
Subfm. †Viverravinae Matthew 1909
+
G. †Pappictidops Qiu and Li 1977
+
†Pappictidops acies Wang 1978
†Pappictidops obtusus Wang 1978
†Pappictidops orientalis Qiu and Li 1977
G. †Monachiopsis Kretzoi 1941
Fm. †Percrocutidae Werdelin and Solounias 1991
+
G. †Dinocrocuta Schmidt-Kittler 1976
+
†Dinocrocuta algeriensis Arambourg 1959
†Dinocrocuta gigantea Schlosser 1903
G. †Percrocuta Kretzoi 1938
+
†Percrocuta australis Hendey 1974
†Percrocuta carnifex Pilgrim 1913
†Percrocuta hebeiensis Chen and Wu 1976
†Percrocuta miocenica Pavlovic and Thenius 1965
†Percrocuta primordialis Qiu and Cao 1988
†Percrocuta tobieni Crusafont-Pairo and Aguirre 1971
†Percrocuta xixiaensis Xiong 2023
G. †Pseudailurus Gervais 1848
G. †Pristinictis Fox and Youzwyshyn 1994
+
†Pristinictis connata Fox and Youzwyshyn 1994
G. †Ravenictis Fox and Youzwyshyn 1994
+
†Ravenictis krausei Fox and Youzwyshyn 1994
G. †Thos Bogachev
+
†Thos terblanchei Broom 1948
Invalid names: Eosictis Scott 1945 [nomen dubium], Felidi Jaekel 1911 [empty], Phocidi Jaekel 1911 [empty], Protelidi Jaekel 1911 [empty], Protoferi Jaekel 1911 [empty], Ursina Gray 1825 [empty], Ursocanidi Jaekel 1911 [empty], Viverra memphitis Linnaeus 1758 [nomen vanum]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Maximum body mass: 300 kgs
Minimum body mass: 100 kgs
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: ground dwellingo
Diet: carnivoreo
Diet 2: omnivoreo
Reproduction: viviparoussubc
Comments: life habit and diet based on Nowak 1999 o
Created: 2005-08-26 13:31:06
Modified: 2005-08-26 15:31:06
Source: s = species, o = order, subc = subclass, c = class, subp = subphylum
References: Carroll 1988, Nowak 1999, Martin 1990, Hendy et al. 2009, Ji et al. 2002, Lillegraven 1979

Age range

Maximum range based only on fossils: base of the Torrejonian to the top of the Holocene or 63.80000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 63.3 Ma

Collections (5093 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Puercan66.0 - 63.3Canada (Saskatchewan) Carnivora indet., Ravenictis krausei (14768)
Early/Lower Paleocene66.0 - 61.7China (Anhui) Pappictidops orientalis (132943 146556)
Early/Lower Paleocene66.0 - 61.7China (Guangdong) Pappictidops acies, Pappictidops obtusus (140273 176607)
Danian66.0 - 61.6USA (North Dakota) Miacidae indet., Viverravinae indet. (14962)
Cenozoic66.0 - 0.0Mongolia (South Gobi Aimag) Carnivora indet. (137861)
Torrejonian63.8 - 60.9USA (Montana) Ictidopappus mustelinus (14906)
Torrejonian63.8 - 60.9USA (Texas) Protictis sp. (14887)
Ti161.7 - 56.8Canada (Alberta) Pristinictis connata (14939)
Tiffanian61.7 - 56.8USA (North Dakota) Miacidae indet. (14931)
Tiffanian61.7 - 56.8USA (Wyoming) Didymictis sp. (14999) Viverravinae indet. (15032 15049)
Tiffanian61.7 - 56.8USA (Montana) Protictis sp. (15048)
Ypresian56.0 - 47.8United Kingdom Miacidae indet. (87950)
Ypresian56.0 - 47.8France (Île-de-France) Miacidae indet. (68437)
Ypresian56.0 - 47.8France (Marne) Miacidae indet. (35533 35535)
Ypresian56.0 - 47.8United Kingdom (England) Miacidae indet., Viverridae indet. (133368)
Sharamurunian56.0 - 37.71Russian Federation Carnivora indet. (192038)
Sharamurunian56.0 - 37.71China (Shanxi) Chailicyon crassidens (41116)
Graybullian55.8 - 50.3USA (Wyoming) Carnivora indet. (15878)
Lostcabinian55.8 - 50.3USA (Wyoming) Carnivora indet. (15917) Uintacyon sp. (15915 15930 16078) Viverravinae indet. (16073)
Lysitean - Lostcabinian55.8 - 50.3USA (Wyoming) Miacidae indet. (15451)