PBDB Taxon

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Neurankylus

Reptilia - Baenidae

Taxonomy

Neurankylus was named by Lambe (1902). Its type is Neurankylus eximius.

It was assigned to Chelydridae by Lambe (1902); to Pleurosternidae by Gilmore (1916), Gilmore (1919), Hay (1930) and Gilmore (1935); to Neurankylidae by Carroll (1988); to Selmacryptodira by Sullivan et al. (1988); to Baenini by Hutchison and Holroyd (2003); to Paracryptodira by Anquetin et al. (2009); to Neurankylinae by Holroyd and Hutchison (2002) and Larson et al. (2013); to Testudinata by Perea et al. (2014); to Baenidae by Joyce and Lyson (2015); and to Baenidae by Hay (1908), Gaffney (1972), Bryant (1989), Joyce (2007), Lyson and Joyce (2010), Lyson and Joyce (2011), Anquetin (2012), Sullivan et al. (2012), Hutchison et al. (2013), Lively (2016), Lichtig and Lucas (2016), Lyson et al. (2016) and Panascí et al. (2019).

Species

N. baueri (syn. N. hutchisoni, N. utahensis), N. eximius (syn. Charitemys captans, Baena fluviatilis) (type species), N. lithographicus, N. notos, N. torrejonensis

Synonyms

Charitemys was named by Hay (1908). Its type is Charitemys captans.

It was synonymized subjectively with Neurankylus by Gaffney (1972).

It was assigned to Baenidae by Hay (1908) and Hay (1930).

Synonymy list

Year	Name and author
1902	Neurankylus Lambe p. 42
1908	Neurankylus Hay p. 93
1908	Charitemys Hay p. 98
1916	Neurankylus Gilmore p. 289
1919	Neurankylus Gilmore p. 113
1930	Neurankylus Hay p. 70
1930	Charitemys Hay p. 73
1935	Neurankylus Gilmore p. 165
1972	Neurankylus Gaffney pp. 249, 291
1988	Neurankylus Carroll
1988	Neurankylus Sullivan et al. p. 2
1989	Neurankylus Bryant pp. 36-37
2002	Neurankylus Holroyd and Hutchison p. 181
2003	Neurankylus Hutchison and Holroyd
2007	Neurankylus Joyce
2009	Neurankylus Anquetin et al.
2010	Neurankylus Lyson and Joyce
2011	Neurankylus Lyson and Joyce p. 792
2012	Neurankylus Anquetin p. 28 fig. 10
2012	Neurankylus Sullivan et al.
2013	Neurankylus Hutchison et al.
2013	Neurankylus Larson et al.
2014	Neurankylus Perea et al.
2015	Neurankylus Joyce and Lyson p. 159
2016	Neurankylus Lichtig and Lucas
2016	Neurankylus Lively
2016	Neurankylus Lyson et al.
2019	Neurankylus Panascí et al.

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda

Rank	Name	Author
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
—	Testudinata	(Oppel 1811)
—	Paracryptodira
family	Baenidae	Cope 1882
genus	Neurankylus

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. †Neurankylus Lambe 1902

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†Neurankylus baueri Gilmore 1916

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Invalid names: Neurankylus hutchisoni Lively 2016 [synonym], Neurankylus utahensis Lively 2016 [synonym]

†Neurankylus eximius Lambe 1902

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Invalid names: Baena fluviatilis Parks 1933 [synonym], Charitemys captans Hay 1908 [synonym]

†Neurankylus lithographicus Larson et al. 2013

†Neurankylus notos Lichtig and Lucas 2016

†Neurankylus torrejonensis Lyson et al. 2016

Invalid names: Charitemys Hay 1908 [synonym]

Diagnosis

Reference	Diagnosis
O. P. Hay 1908	A genus of uncertain position and known only from a portion of the carapace of the type species. Eighth neural large, followed by an expanded suprapygal. In the type a ninth pair of costal bones. The vertebral scutes nearly twice as wide as long.
O. P. Hay 1908 (Charitemys)	A genus of Baenidae. So far as known like Thescelus, but with the axillary buttresses ascending to near the neural borders of the first costals. Inguinal buttresses probably ascending on inner surfaces of the fifth and sixth costals, but not to so great a height as did the axillary buttresses.
E. S. Gaffney 1972	1) shell shape oval, sides parallel; 2) ornamentation mostly smooth, delicate striations marking growth lines in some specimens, large, raised ridges in others; 3) nuchal scute small, rectangular, bordered laterally by large first marginals; 4) no anterior emargination of carapace; peripherals not meeting in front of nuchal; 5) preneural bone absent; 6) first vertebral scute rectangular without supracostal scutes; 7) supramarginal scutes absent; 8) last vertebral scute not open posteriorly, last pair of marginals meeting medially behind vertebral, last marginals rectangular; 9) posterior edge of carapace not emarginated; 10) posterior edge of carapace sometimes with shallow scallops; 11) xiphiplastral notch absent; 12) paired intergulars meeting medially, gulars separated by intergulars.
D. W. Larson et al. 2013	Large relative to other baenid turtles (carapace averaging 500–600 mm long and skull more than 74 mmlong in adult individuals); dorsolateral gutters along the peripherals of the carapace; shell ornamentation essentially smooth, but sometimes consisting of subtle parallel ridges and fine, low-density pitting; foramen stapediotemporale bordered by the opisthotic, quadrate, and prootic, but not the supraoccipital; posteriorly rounded short supraoccipital ridge consisting of only the paired parietals in dorsal view.
W. G. Joyce and T. R. Lyson 2015	Neurankylus is diagnosed as a baenid by the complete list of characters: presence of the following derived characters: pterygoid-basioccipital contact, axillary and inguinal buttresses that reach the costals and lack of epiplastral processes. Baenids symplesiomorphically possess a processus trochlearis oticum, a foramen posterius canalis carotici interni that is located halfway along the pterygoid-basisphenoid contact and mesoplastra.