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Taxonomy
Neurankylus was named by Lambe (1902). Its type is Neurankylus eximius.
It was assigned to Chelydridae by Lambe (1902); to Pleurosternidae by Gilmore (1916), Gilmore (1919), Hay (1930) and Gilmore (1935); to Neurankylidae by Carroll (1988); to Selmacryptodira by Sullivan et al. (1988); to Baenini by Hutchison and Holroyd (2003); to Paracryptodira by Anquetin et al. (2009); to Neurankylinae by Holroyd and Hutchison (2002) and Larson et al. (2013); to Testudinata by Perea et al. (2014); to Baenidae by Joyce and Lyson (2015); and to Baenidae by Hay (1908), Gaffney (1972), Bryant (1989), Joyce (2007), Lyson and Joyce (2010), Lyson and Joyce (2011), Anquetin (2012), Sullivan et al. (2012), Hutchison et al. (2013), Lively (2016), Lichtig and Lucas (2016), Lyson et al. (2016) and Panascí et al. (2019).
It was assigned to Chelydridae by Lambe (1902); to Pleurosternidae by Gilmore (1916), Gilmore (1919), Hay (1930) and Gilmore (1935); to Neurankylidae by Carroll (1988); to Selmacryptodira by Sullivan et al. (1988); to Baenini by Hutchison and Holroyd (2003); to Paracryptodira by Anquetin et al. (2009); to Neurankylinae by Holroyd and Hutchison (2002) and Larson et al. (2013); to Testudinata by Perea et al. (2014); to Baenidae by Joyce and Lyson (2015); and to Baenidae by Hay (1908), Gaffney (1972), Bryant (1989), Joyce (2007), Lyson and Joyce (2010), Lyson and Joyce (2011), Anquetin (2012), Sullivan et al. (2012), Hutchison et al. (2013), Lively (2016), Lichtig and Lucas (2016), Lyson et al. (2016) and Panascí et al. (2019).
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1902 | Neurankylus Lambe p. 42 |
| 1908 | Neurankylus Hay p. 93 |
| 1908 | Charitemys Hay p. 98 |
| 1916 | Neurankylus Gilmore p. 289 |
| 1919 | Neurankylus Gilmore p. 113 |
| 1930 | Neurankylus Hay p. 70 |
| 1930 | Charitemys Hay p. 73 |
| 1935 | Neurankylus Gilmore p. 165 |
| 1972 | Neurankylus Gaffney pp. 249, 291 |
| 1988 | Neurankylus Carroll |
| 1988 | Neurankylus Sullivan et al. p. 2 |
| 1989 | Neurankylus Bryant pp. 36-37 |
| 2002 | Neurankylus Holroyd and Hutchison p. 181 |
| 2003 | Neurankylus Hutchison and Holroyd |
| 2007 | Neurankylus Joyce |
| 2009 | Neurankylus Anquetin et al. |
| 2010 | Neurankylus Lyson and Joyce |
| 2011 | Neurankylus Lyson and Joyce p. 792 |
| 2012 | Neurankylus Anquetin p. 28 fig. 10 |
| 2012 | Neurankylus Sullivan et al. |
| 2013 | Neurankylus Hutchison et al. |
| 2013 | Neurankylus Larson et al. |
| 2014 | Neurankylus Perea et al. |
| 2015 | Neurankylus Joyce and Lyson p. 159 |
| 2016 | Neurankylus Lichtig and Lucas |
| 2016 | Neurankylus Lively |
| 2016 | Neurankylus Lyson et al. |
| 2019 | Neurankylus Panascí et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Neurankylus Lambe 1902
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†Neurankylus baueri Gilmore 1916
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Invalid names: Neurankylus hutchisoni Lively 2016 [synonym], Neurankylus utahensis Lively 2016 [synonym]
†Neurankylus eximius Lambe 1902
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Invalid names: Baena fluviatilis Parks 1933 [synonym], Charitemys captans Hay 1908 [synonym]
†Neurankylus lithographicus Larson et al. 2013
†Neurankylus notos Lichtig and Lucas 2016
†Neurankylus torrejonensis Lyson et al. 2016
Invalid names: Charitemys Hay 1908 [synonym]
Diagnosis
| Reference | Diagnosis | |
|---|---|---|
| O. P. Hay 1908 | A genus of uncertain position and known only from a portion of the carapace of the type species. Eighth neural large, followed by an expanded suprapygal. In the type a ninth pair of costal bones. The vertebral scutes nearly twice as wide as long. | |
| O. P. Hay 1908 (Charitemys) | A genus of Baenidae. So far as known like Thescelus, but with the axillary buttresses ascending to near the neural borders of the first costals. Inguinal buttresses probably ascending on inner surfaces of the fifth and sixth costals, but not to so great a height as did the axillary buttresses. | |
| E. S. Gaffney 1972 | 1) shell shape oval, sides parallel; 2) ornamentation mostly smooth, delicate striations marking growth lines in some specimens, large, raised ridges in others; 3) nuchal scute small, rectangular, bordered laterally by large first marginals; 4) no anterior emargination of carapace; peripherals not meeting in front of nuchal; 5) preneural bone absent; 6) first vertebral scute rectangular without supracostal scutes; 7) supramarginal scutes absent; 8) last vertebral scute not open posteriorly, last pair of marginals meeting medially behind vertebral, last marginals rectangular; 9) posterior edge of carapace not emarginated; 10) posterior edge of carapace sometimes with shallow scallops; 11) xiphiplastral notch absent; 12) paired intergulars meeting medially, gulars separated by intergulars. | |
| D. W. Larson et al. 2013 | Large relative to other baenid turtles
(carapace averaging 500–600 mm long and skull more than 74 mmlong in adult individuals); dorsolateral gutters along the peripherals of the carapace; shell ornamentation essentially smooth, but sometimes consisting of subtle parallel ridges and fine, low-density pitting; foramen stapediotemporale bordered by the opisthotic, quadrate, and prootic, but not the supraoccipital; posteriorly rounded short supraoccipital ridge consisting of only the paired parietals in dorsal view. | |
| W. G. Joyce and T. R. Lyson 2015 | Neurankylus is diagnosed as a baenid by the complete list of characters: presence of the following derived characters: pterygoid-basioccipital contact, axillary and inguinal buttresses that reach the costals and lack of epiplastral processes. Baenids symplesiomorphically possess a processus trochlearis oticum, a foramen posterius canalis carotici interni that is located halfway along the pterygoid-basisphenoid contact and mesoplastra. |
Measurements
No measurements are available
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| Source: c = class, subp = subphylum, uc = unranked clade | |||||
| References: Hendy et al. 2009, Carroll 1988 | |||||
Collections (60 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Coniacian | USA (Montana) | N. sp. (206447) | |
| Coniacian - Santonian | USA (New Mexico) | N. notos (182494) | |
| Late/Upper Santonian | Canada (Alberta) | N. lithographicus (154706 154707) | |
| Campanian | Canada (Alberta) | Baena fluviatilis (128457) | |
| Early/Lower Campanian - Middle Campanian | USA (Utah) | N. sp. (167916) | |
| Late/Upper Campanian | Canada (Alberta) | N. eximius (22697 45261 46953 128454 187581) | |
| Late/Upper Campanian | Mexico (Coahuila) | Chelonia indet. (35360) | |
| Late/Upper Campanian | USA (Utah) | N. sp. A (187657 187659 187693 187702 187706 187713) N. utahensis (219712) N. utahensis, N. sp. B (187674) | |
| Late/Upper Campanian | USA (New Mexico) | N. baueri (62317 62318 62481 154367 154372 154379 154382 154386 154419 154599 154600 154601) N. eximius (52052 128456) N. sp. (100582) | |
| Middle Campanian | USA (Montana) | N. sp. (14437) | |
| Middle Campanian - Late/Upper Campanian | USA (Montana) | Charitemys captans (128455) | |
| Lancian | USA (North Dakota) | N. sp. (137705 137706 137712 137715 137724 137730 137731) | |
| Lancian | USA (Montana) | N. eximius (14639) N. sp. (14512) | |
| Lancian | USA (Wyoming) | Aspideretoides sp. (75292) Baenidae indet. (75295) | |
| Late/Upper Maastrichtian | USA (Montana) | N. eximius (128458) N. sp. (189003) | |
| Late/Upper Maastrichtian | USA (South Dakota) | N. sp. (224484) | |
| Late/Upper Maastrichtian | Canada (Saskatchewan) | N. sp. (14555) | |
| Late/Upper Maastrichtian | USA (Colorado) | N. sp. (49551) | |
| Late/Upper Maastrichtian | USA (North Dakota) | N. sp. (137718) | |
| Puercan | USA (New Mexico) | N. eximius (74532) N. torrejonensis (197332) | |
| Puercan | USA (Colorado) | N. sp. (205861 205904 205906) | |
| Torrejonian | USA (New Mexico) | N. torrejonensis (179481) |