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Echmatemys
Taxonomy
Echmatemys was named by Hay (1906). It is not extant. Its type is Emys septarius.
It was assigned to Emydidae by Hay (1906), Gilmore (1915), Hay (1930), Carroll (1988) and Hutchison (1992); to Bataguridae by Hutchison (1996) and Holroyd et al. (2001); to Ptychogasteridae by Hervet (2004); to Geoemydidae by Lichtig and Lucas (2015); and to Geoemydidae by Claude et al. (2012), Lichtig and Lucas (2015), Vlachos (2018) and Vlachos (2020).
It was assigned to Emydidae by Hay (1906), Gilmore (1915), Hay (1930), Carroll (1988) and Hutchison (1992); to Bataguridae by Hutchison (1996) and Holroyd et al. (2001); to Ptychogasteridae by Hervet (2004); to Geoemydidae by Lichtig and Lucas (2015); and to Geoemydidae by Claude et al. (2012), Lichtig and Lucas (2015), Vlachos (2018) and Vlachos (2020).
Species
E. arethusa, E. callopyge, E. cibollensis, E. depressa, E. haydeni, E. jeanesi, E. lativertebralis (syn. E. douglassi, E. hollandi, Chrysemys inornata, Emys shaughnessiana, Graptemys cordifera, E. rivalis), E. naomi (syn. E. obscura), E. stevensoniana (syn. E. cyane, E. aegle, Emys septarius, E. ocyrrhoe), Palaeotheca polycypha, Palaeotheca terrestris
Synonyms
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Synonymy list
Year | Name and author |
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1872 | Palaeotheca polycypha Cope p. 463 |
1872 | Palaeotheca terrestris Cope p. 464 |
1873 | Emys polycypha Cope |
1873 | Emys terrestris Cope |
1884 | Emys polycypha Cope |
1884 | Emys terrestris Cope |
1902 | Emys polycypha Hay p. 448 |
1902 | Emys terrestris Hay p. 448 |
1906 | Echmatemys Hay p. 27 |
1915 | Echmatemys Gilmore p. 123 |
1915 | Echmatemys depressa Gilmore p. 139 figs. 2; 16 |
1930 | Echmatemys Hay p. 95 |
1930 | Echmatemys depressa Hay p. 95 |
1930 | Palaeotheca polycypha Hay p. 96 |
1930 | Palaeotheca terrestris Hay p. 96 |
1988 | Echmatemys Carroll |
1992 | Echmatemys Hutchison |
1996 | Echmatemys Hutchison p. 340 |
2001 | Echmatemys Holroyd et al. |
2004 | Echmatemys Hervet |
2012 | Echmatemys Claude et al. |
2015 | Echmatemys Lichtig and Lucas |
2018 | Echmatemys Vlachos |
2020 | Echmatemys Vlachos p. 13 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
G. †Echmatemys Hay 1906
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†Echmatemys arethusa Hay 1908
†Echmatemys callopyge Hay 1908
†Echmatemys cibollensis Cope 1877
†Echmatemys haydeni Leidy 1870
†Echmatemys jeanesi Leidy 1870
†Echmatemys lativertebralis Cope 1877
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Invalid names: Chrysemys inornata Loomis 1904 [synonym], Echmatemys douglassi Gilmore 1915 [synonym], Echmatemys hollandi Gilmore 1915 [synonym], Echmatemys rivalis Hay 1909 [synonym], Emys shaughnessiana Cope 1882 [synonym], Graptemys cordifera Clark 1937 [synonym]
†Echmatemys stevensoniana Leidy 1870
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Invalid names: Echmatemys aegle Hay 1908 [synonym], Echmatemys cyane Hay 1908 [synonym], Echmatemys ocyrrhoe Hay 1908 [synonym], Emys septarius Cope 1873 [synonym]
Invalid names: Echmatemys depressa Gilmore 1915 [nomen dubium], Palaeotheca polycypha Cope 1872 [nomen dubium], Palaeotheca terrestris Cope 1872 [nomen dubium]
Diagnosis
Reference | Diagnosis | |
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E. Vlachos 2018 | Echmatemys can be diagnosed as a member of Geoemydidae based on by the presence of musk ducts and associated musk duct foramina, wide rib heads, a narrow costo-vertebral tunnel, and paired marginals XII that expand on the suprapygal II. Echmatemys differs from other Geoemydidae based on the presence of narrow triturating surfaces, a short cervical scute, mostly hexagonal neurals II–VIII with short antero-lateral sides, a posteriorly flared carapace margin, anals that are shorter medially than the femorals, and well-developed axillary and inguinal buttresses. |
Measurements
No measurements are available
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Source: g = genus, subo = suborder, c = class, subp = subphylum | |||||
References: Holroyd et al. 2001, Uetz 2005, Hendy et al. 2009, Carroll 1988 |
Age range: base of the Wasatchian to the top of the Duchesnean or 55.80000 to 37.00000 Ma
Collections (85 total)
Time interval | Ma | Country or state | Original ID and collection number |
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Lysitean | USA (Colorado) | E. stevensoniana (15654) | |
Wasatchian | USA (Colorado) | E. lativertebralis (176837) | |
Wasatchian | USA (Wyoming) | E. lativertebralis (46718) E. rivalis (177320) E. sp. (15458 15563 15566 15587 15759 46650 46772 59904 59905) | |
Wasatchian | USA (New Mexico) | E. lativertebralis (15443 176872 176873 176877 176879 176880 176882 176885 176886 176887 176888 176890 176891 176894 176898 176899 176900 176902 176909 176911 176912 176914) Emys cibollensis, Emys lativertebralis (131867) | |
Wasatchian - Bridgerian | USA (New Mexico) | E. sp. (25762) | |
Wasatchian - Bridgerian | USA (Wyoming) | E. sp. (16134) | |
Bridgerian | USA (Wyoming) | E. arethusa, E. aegle, E. cyane (16228) E. callopyge, E. wyomingensis (74504) E. cibollensis (70264) E. naomi (16255 206092) E. ocyrrhoe (16201) E. septaria (15837 16261 16262 16265 16267 16270 16273 16405 55207) E. sp. (16209 16296) Emys haydeni, Palaeotheca polycypha, Palaeotheca terrestris, Emys shaugnessiana (130809) Emys jeanesi, Emys haydeni, Emys stevensonianus (128447) Emys septarius (191004) | |
Bridgerian | USA (Colorado) | E. stevensoniana (16242) | |
Bridgerian | USA (Utah) | E. sp. (16396) | |
Middle Eocene | USA (Texas) | E. sp. (37559) | |
Early/Lower Uintan | USA (Colorado) | E. sp., E. septaria (16623) | |
Early/Lower Uintan | USA (Utah) | E. callopyge (186888) | |
Late/Upper Uintan | USA (Utah) | E. callopyge (109166) | |
Uintan | USA (Utah) | E. callopyge (185685 185688 186896 205622 213059 213066) E. depressa (185693) E. douglassi (185696) E. hollandi (185697) E. obscura (185689) E. septaria (74505) E. sp. (222799 222802 222803 222807) E. sp., E. callopyge (222801 222805) | |
Duchesnean | USA (Utah) | E. sp. (16877) | |
Chadronian | USA (South Dakota) | Chrysemys inornata (16900) Graptemys cordifera (16938 17013) |