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Hardella

Reptilia - Testudines - Geoemydidae

Synonymy list
YearName and author
1870Hardella Gray
1889Hardella Lydekker pp. 119-120
1988Hardella Carroll
1989Hardella Ernst and Barbour
1996Hardella Frank and Ramus
2014Hardella van Dijk et al.
2020Hardella Garbin et al. p. 25

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
RankNameAuthor
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
Sauropsida
classReptilia
Testudinata(Oppel 1811)
orderTestudinesBatsch 1788
suborderCryptodira
Pantestudinoidea
superfamilyTestudinoidea
familyGeoemydidaeTheobald 1868
Palatochelydia
genusHardellaGray 1870

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Hardella Gray 1870
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Hardella siamensis Claude et al. 2007
Hardella thurjii Gray 1831
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Invalid names: Batagur falconeri Lydekker 1885 [synonym], Clemmys watsoni Lydekker 1886 [synonym]
Diagnosis
ReferenceDiagnosis
R. Garbin et al. 2020Hardella thurjii can be differentiated from other geoemydids by large carapace size (up to 60 cm in females), presence of hexagonal, nearly square second to fourth vertebrals, a first vertebral scute that is wider posteriorly, presence of an inflection at the margin of the gulohumeral sulcus, and an entoplastron that is not intersected by the humeropectoral sulcus.