PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Mauremys (pond turtle)

Reptilia - Testudines - Geoemydidae

Taxonomy

Mauremys was named by Gray (1870). It is extant.

It was assigned to Emydidae by Carroll (1988) and Frank and Ramus (1996); to Batagurinae by Bickham (1975), Ernst and Barbour (1989) and Staesche et al. (2007); to Geoemydinae by van Dijk et al. (2014); and to Geoemydidae by Claude et al. (2012), Vlachos et al. (2015), Vlachos and Delfino (2016), Vlachos et al. (2019), Collareta et al. (2020) and Vlachos (2021).

Species lacking formal opinion data

M. batalleri

Synonyms

Paralichelys was named by Bergounioux (1935).

It was synonymized subjectively with Mauremys by Hervet (2004).

It was assigned to Pelomedusidae by Carroll (1988).
Emys prosigriz was named by Pallary (1901). Its type specimen is MNHN.F 1874-47, a partial shell (a partial steinkern of a carapace and plastron), and it is a cast. Its type locality is Mansourah, which is in a Pleistocene fluvial-lacustrine travertine/sandstone in Algeria.

It was considered a nomen dubium by Georgalis et al. (2021).
Clemmys precaspica was named by de Stefano (1902). Its type specimen is MNHN number unknown, a shell, and it is a 3D body fossil. Its type locality is Mansourah, which is in a Pleistocene fluvial-lacustrine travertine/sandstone in Algeria.

It was considered a nomen dubium by Georgalis et al. (2021).

Synonymy list

Year	Name and author
1870	Mauremys Gray
1901	Emys prosigriz Pallary p. 69
1902	Clemmys precaspica de Stefano p. 284
1935	Paralichelys Bergounioux
1975	Mauremys Bickham p. 203
1988	Mauremys Carroll
1988	Paralichelys Carroll
1989	Mauremys Ernst and Barbour
1996	Mauremys Frank and Ramus
2007	Mauremys Staesche et al.
2012	Mauremys Claude et al.
2014	Mauremys van Dijk et al.
2015	Mauremys Vlachos et al.
2016	Mauremys Vlachos and Delfino
2019	Mauremys Vlachos et al.
2020	Mauremys Collareta et al.
2021	Mauremys Vlachos

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha

Rank	Name	Author
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
—	Testudinata	(Oppel 1811)
order	Testudines	Batsch 1788
suborder	Cryptodira
—	Pantestudinoidea
superfamily	Testudinoidea
family	Geoemydidae	Theobald 1868
genus	Mauremys	Gray 1870

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

G. Mauremys Gray 1870 [pond turtle]

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Mauremys annamensis Siebenrock 1903 [Vietnamese pond turtle]

†Mauremys aristotelica Vlachos et al. 2019

Mauremys caspica Gmelin 1774 [Caspian turtle]

†Mauremys fenhoense Gray 1931

†Mauremys gaudryi Depéret 1885

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Invalid names: Clemmys romani Bergounioux 1935 [synonym], Emys etrusca Portis 1890 [synonym], Paralichelys carinata Bergounioux 1935 [synonym]

†Mauremys guntiana Roger 1902

†Mauremys isoclina Dubois 1908

Mauremys japonica Temminck and Schlegel 1835 [Japanese pond turtle]

Mauremys leprosa Schweigger 1812 [Mediterranean pond turtle]

†Mauremys massiliensis Bergounioux 1936

Mauremys mutica Cantor 1842 [yellow pond turtle]

Mauremys nigricans Gray 1834 [Chinese red-necked turtle]

†Mauremys portisi Sacco 1889

†Mauremys pygolopha Peters 1868

Mauremys reevesii Gray 1831 [Reeves turtle]

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Invalid names: Chinemys pani Tao 1985 [synonym]

Mauremys rivulata Valenciennes 1833 [Western Caspian turtle]

†Mauremys sarmatica Purschke 1885

Mauremys sinensis Gray 1834 [Chinese stripe-necked turtle]

†Mauremys sophiae Ammon 1911

†Mauremys thanhinensis Claude et al. 2007

†Mauremys yabei Shikama 1949

Invalid names: Clemmys precaspica de Stefano 1902 [nomen dubium], Emys prosigriz Pallary 1901 [nomen dubium], Paralichelys Bergounioux 1935 [synonym]

Diagnosis

Reference	Diagnosis
E. Vlachos et al. 2019	Member of Geoemydidae based on the following combination of characters: presence of anterior and posterior musk duct foramina; neural 1 quadrangular followed by hexagonal neurals with short anterolateral sides; rib heads wide and flat, placed near the neural/costal suture on the costals; 12th marginals divided, with their sulcus expanding on the suprapygal 2 that is crossed by the posterior border of vertebral 5.
E. Vlachos 2021	Mauremys can be distinguished by a vertebral I that contacts the marginal II, an entoplastron that is crossed by the humero-pectoral sulcus on its posterior part, short anal scutes, and a deep rounded anal notch.