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Taxonomy
Sciaraedes was named by Billberg (1820). It is extant. Its type is Sciara.
It was corrected as Sciaridae by Zhang (1989), Carpenter (1992), Cook and Ross (1996) and Yeates et al. (2007).
It was assigned to Diptera by Zhang (1989); to Nematocera by Carpenter (1992); to Mycetophiloidea by Cook and Ross (1996); and to Sciaroidea by Yeates et al. (2007).
It was corrected as Sciaridae by Zhang (1989), Carpenter (1992), Cook and Ross (1996) and Yeates et al. (2007).
It was assigned to Diptera by Zhang (1989); to Nematocera by Carpenter (1992); to Mycetophiloidea by Cook and Ross (1996); and to Sciaroidea by Yeates et al. (2007).
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1904 | Sciara morosa Meunier p. 78 |
| 1915 | Sciara lacoei Cockerell p. 491 figs. Pl 62, fig 6 |
| 1989 | Sciaridae Zhang p. 357 |
| 1992 | Sciaridae Carpenter |
| 1994 | Lycoriella (Lycoriella) morosa Evenhuis |
| 1996 | Sciaridae Cook and Ross |
| 2000 | Lycoriella morosa Menzel and Mohrig p. 376 |
| 2007 | Sciaridae Yeates et al. |
| 2019 | Sciara lacoei Blagoderov p. 541 figs. Pl 7j |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. Sciaridae Billberg 1820 [dark-winged fungus gnat]
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G. Archicratyna Mohrig 2005
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†Archicratyna arcana Mohrig and Röschmann 2005
†Archicratyna subarcana Mohrig and Röschmann 2005
†Archicratyna trichoarcana Mohrig and Röschmann 2005
Subfm. Chaetosciarinae Shin 2019
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Subfm. Cratyninae Enderlein 1911
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G. Corynoptera Winnertz 1867
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†Corynoptera curiosa Meunier 1904
†Corynoptera dubia Meunier 1904
†Corynoptera evenhuisi Röschmann and Mohrig 1995
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Invalid names: Sciara difficilis Meunier 1904 [replaced]
†Corynoptera iocosa Mohrig and Röschmann 2005
†Corynoptera klebsii Meunier 1904
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Invalid names: Bradysia electra Meunier 1904 [synonym], Bradysia infernalis Meunier 1904 [synonym], Bradysia morosoides Meunier 1904 [synonym], Sciara bella Meunier 1904 [synonym], Sciara hirticornis Loew 1850 [replaced], Sciara minuscula Meunier 1904 [synonym], Sciara palmnickii Meunier 1904 [synonym], Sciara variabilis Meunier 1904 [synonym]
†Corynoptera koenigsbergensis Mohrig and Röschmann 1994
†Corynoptera neutralis Röschmann and Mohrig 1995
†Corynoptera prisca Mohrig and Röschmann 2005
†Corynoptera pristina Röschmann and Mohrig 2005
†Corynoptera ritzkowskii Röschmann and Mohrig 1995
†Corynoptera schumanni Mohrig and Röschmann 1994
†Corynoptera subagilis Mohrig and Röschmann 1994
†Corynoptera tritospinosula Röschmann and Mohrig 1995
†Corynoptera umbrosa Meunier 1904
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Invalid names: Bradysia agilis Meunier 1904 [synonym], Bradysia conwentzii Meunier 1904 [synonym]
Invalid names: Sciara spinulosa Meunier 1899 [nomen nudum]
G. Cratyna Winnertz 1867
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Subg. Cratyna (Peyerimhoffia) Kieffer 1903
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†Cratyna (Peyerimhoffia) keilbachi Mohrig and Röschmann 1994
†Cratyna interposita Mohrig and Röschmann 2005
†Cratyna villosoantennata Mohrig and Röschmann 2005
Invalid names: Plastosciara Berg 1899 [synonym]
G. Epidapus Haliday 1851
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Subg. Epidapus (Epidapus) Haliday 1851
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†Epidapus (Epidapus) brevihalteratus Röschmann and Mohrig 1995
†Epidapus (Epidapus) palaeogracilis Roschmann and Mohrig 1993
†Epidapus (Epidapus) unistylatus Röschmann and Mohrig 1995
Subg. Epidapus (Pseudoaptanogyna) Vimmer 1926
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†Epidapus (Pseudoaptanogyna) primarius Röschmann and Mohrig 1995
†Epidapus (Pseudoaptanogyna) succinellus Mohrig and Röschmann 1994
†Epidapus adstrictosetus Mohrig and Röschmann 2005
†Epidapus longisetus Mohrig and Röschmann 2005
†Epidapus macrospinatus Mohrig and Röschmann 2005
†Epidapus microspinus Mohrig and Röschmann 2005
Subfm. Megalosphyinae Enderlein 1911
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G. Bradysia Winnertz 1867
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†Bradysia aliciae Ramírez and Alonso 2014
†Bradysia antiqua Mohrig and Röschmann 2005
†Bradysia dimidiata Mohrig and Röschmann 2005
†Bradysia exoleta Mohrig and Röschmann 2005
†Bradysia oligocenica Quiévreux 1938
†Bradysia tertiaria Quiévreux 1938
G. Zygoneura Meigen 1830
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†Zygoneura curta Hong 1981
Invalid names: Apatemosciaritis Hong 2002 [synonym]
G. †Protolycoriella Röschmann and Mohrig 1995
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†Protolycoriella aliena Röschmann and Mohrig 1995
†Protolycoriella gigaspina Röschmann and Mohrig 1995
†Protolycoriella neogenica Röschmann and Mohrig 1995
G. Pseudosciara Schiner 1866
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†Pseudosciara bifurcata Meunier 1904
Invalid names: Heeriella Meunier 1904 [synonym]
Subfm. Sciarinae Billberg 1820
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G. Leptosciarella Tuomikoski 1960
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Subg. Leptosciarella (Leptosciarella) Tuomikoski 1960
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†Leptosciarella (Leptosciarella) botuli Meunier 1904
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Invalid names: Sciara sendelina Meunier 1904 [synonym], Sciara tanypeza Meunier 1899 [synonym]
†Leptosciarella (Leptosciarella) infausta Mohrig and Röschmann 1994
†Leptosciarella (Leptosciarella) macrociliata Mohrig and Röschmann 1994
†Leptosciarella (Leptosciarella) manifesta Mohrig and Röschmann 2005
†Leptosciarella (Leptosciarella) microacantha Röschmann and Mohrig 1995
†Leptosciarella (Leptosciarella) microciliata Mohrig and Röschmann 1994
†Leptosciarella (Leptosciarella) orientalis Meunier 1904
†Leptosciarella (Leptosciarella) protorotunda Mohrig and Röschmann 1994
†Leptosciarella (Leptosciarella) pseudorecens Röschmann and Mohrig 1995
†Leptosciarella (Leptosciarella) resinae Röschmann and Mohrig 1995
†Leptosciarella (Leptosciarella) tertiaria Meunier 1904
†Trichosia (Leptosciarella) macrociliata Mohrig and Röschmann 1994
G. Sciara Meigen 1803
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†Sciara acuminata Theobald 1937
†Sciara acuminata Heer 1849
†Sciara amicula Statz 1944
†Sciara antennaria Statz 1944
†Sciara atavina von Heyden 1870
†Sciara atropha Zhang 1989
†Sciara aulica Statz 1944
†Sciara austera Statz 1944
†Sciara bella Statz 1944
†Sciara berckhemeri Armbruster 1938
†Sciara bicolora Statz 1944
†Sciara braeuhaeuseri Armbruster 1938
†Sciara branconis Armbruster 1938
†Sciara brevipetiola Zhang 1989
†Sciara burmitina Cockerell 1917
†Sciara compta Statz 1944
†Sciara cryptocephala Statz 1944
†Sciara defectuosa Meunier 1904
†Sciara deffneri Armbruster 1938
†Sciara defossa von Heyden 1870
†Sciara deleta Heer 1864
†Sciara deperdita Scudder 1878
†Sciara dietleni Armbruster 1938
†Sciara dormitans Melander 1949
†Sciara ehrati Armbruster 1938
†Sciara electriphila Cockerell 1910
†Sciara endrissi Armbruster 1938
†Sciara exilis Statz 1944
†Sciara femuralis Statz 1944
†Sciara flichei Theobald 1937
†Sciara fraasi Armbruster 1938
†Sciara gracilenta Statz 1944
†Sciara gracilior Statz 1944
†Sciara hennigi Armbruster 1938
†Sciara heydeni Meunier 1917
†Sciara hirtella Heer 1849
†Sciara hoelzlei Armbruster 1938
†Sciara ignorata Meunier 1904
†Sciara inflata Theobald 1937
†Sciara janassa von Heyden 1870
†Sciara joossi Armbruster 1938
†Sciara jucheti Camier and Nel 2019
†Sciara jucunda Statz 1944
†Sciara kluepfeli Armbruster 1938
†Sciara laticornis Statz 1944
†Sciara lengersdorfi Statz 1944
†Sciara lengersdorfi Armbruster 1938
†Sciara longa Statz 1944
†Sciara longipennata Statz 1944
†Sciara lugens Statz 1944
†Sciara martii Novák 1878
†Sciara melanopa Statz 1944
†Sciara milleri Armbruster 1938
†Sciara minuscula Theobald 1937
†Sciara minuscula Statz 1944
†Sciara minutula Heer 1849
†Sciara oblonga Theobald 1937
†Sciara ovalipennata Statz 1944
†Sciara pelidua Statz 1944
†Sciara pennata Statz 1944
†Sciara pompeckii Armbruster 1938
†Sciara protogaea Heer 1856
†Sciara quenstedti Armbruster 1938
†Sciara radians Meunier 1910
†Sciara rara Meunier 1904
†Sciara requieta Melander 1949
†Sciara rottensis von Heyden 1870
†Sciara saportana Theobald 1937
†Sciara schremmeri Bachmayer 1961
†Sciara scopuli Scudder 1890
†Sciara seemanni Armbruster 1938
†Sciara sepulta Lewis 1969
†Sciara sopora Melander 1949
†Sciara splendida Meunier 1904
†Sciara strausi Kohring and Schlüter 1993
†Sciara thoracica Statz 1944
†Sciara villosoides Meunier 1904
†Sciara wageri Armbruster 1938
†Sciara weylandi Statz 1944
†Sciara willershausensis Kohring and Schlüter 1993
†Sciara winnertzii von Heyden 1870
Invalid names: Sciara basalis Meunier 1899 [nomen nudum], Sciara brachycera Meunier 1899 [nomen nudum], Sciara dasycera Meunier 1899 [nomen nudum], Sciara macrocera Meunier 1899 [nomen nudum], Sciara troglodytes Heer 1861 [nomen nudum]
G. Trichosia Winnertz 1867
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Subg. †Trichosia (Archaeosciara) Mohrig and Röschmann 1994
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†Trichosia (Archaeosciara) ruebsaamenia Meunier 1904
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Invalid names: Sciara robusta Meunier 1904 [synonym]
†Trichosia (Archaeosciara) venohirsuta Röschmann and Mohrig 1995
Subg. †Trichosia (Palaeotrichosia) Mohrig and Röschmann 1994
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†Trichosia (Palaeotrichosia) diabolica Meunier 1904
†Trichosia (Palaeotrichosia) errans Meunier 1904
†Trichosia (Palaeotrichosia) kedingi Röschmann and Mohrig 1995
†Trichosia (Palaeotrichosia) preciosa Meunier 1904
†Trichosia (Palaeotrichosia) voelsgeni Röschmann and Mohrig 1995
Subg. Trichosia (Trichosia) Winnertz 1867
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†Trichosia (Trichosia) meunieri Cockerell 1910
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Invalid names: Sciara prolifica Meunier 1904 [replaced]
Invalid names: Lycoriidae [synonym], Sciara lacoei Cockerell 1915 [nomen dubium], Sciara morosa Meunier 1904 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Kiessling 2004, Bush and Bambach 2015 | |||||
Age range
Maximum range based only on fossils: base of the Late/Upper Hauterivian to the top of the Holocene or 136.40000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 130.0 Ma
Minimum age of oldest fossil (stem group age): 130.0 Ma
Collections (62 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Late/Upper Hauterivian | United Kingdom (England) | Sciaridae indet. (119697) | |
| Albian | Jordan | Sciaridae indet. (124847) | |
| Early/Lower Cenomanian | Myanmar | Sciara burmitina (123020) | |
| Ypresian | France (Oise) | Sciara jucheti (123850) | |
| Ypresian | China (Liaoning) | Lycoria brachyceroidis, Lycoria curta (124530) Mixosciaritis brachyceroidis (124820) | |
| Bridgerian | USA (Wyoming) | Sciara scopuli (122090) | |
| Lutetian | USA (Montana) | Eosciarites hermes (135666) | |
| Priabonian | United Kingdom (England) | Sciara lacoei (123959) | |
| Priabonian | Poland | Trichosia sp., Trichosia botuli, Trichosia macrociliata, Corynoptera sp., Trichosia errans, Corynoptera umbrosa, Trichosia ruebsaamenia, Sciara hirticornis, Corynoptera schumanni, Epidapus succinellus (109625) | |
| Priabonian | Russian Federation | Trichosia sp., Corynoptera sp., Trichosia meunieri, Trichosia microciliata, Corynoptera hirticornis, Corynoptera schumanni, Epidapus menzeli, Sciaridae indet. (123419) | |
| Priabonian | France | Sciara sp. (113709) | |
| Priabonian | Canada (British Columbia) | Sciara deperdita (121480) | |
| Priabonian | Germany (Halle) | Trichosia sp., Epidapus primarius, Trichosia voelsgeni, Trichosia botuli, Trichosia diabolica, Trichosia macrociliata, Trichosia orientalis, Corynoptera sp., Corynoptera klebsi, Corynoptera subagilis, Sciaridae indet. (139214) Trichosia sp., Trichosia kedingi, Trichosia botuli, Corynoptera klebsi, Sciaridae indet. (139215) Trichosia sp., Trichosia voelsgeni, Trichosia botuli, Trichosia diabolica, Trichosia macrociliata, Trichosia orientalis, Corynoptera sp., Corynoptera subagilis, Trichosia errans, Trichosia meunieri, Trichosia tertiaria, Trichosia microciliata, Trichosia ruebsaamenia, Corynoptera koenigsbergensis, Corynoptera schumanni, Trichosia microacantha, Trichosia pseudorecens, Trichosia resinae, Trichosia venohirsuta, Cratyna archaica, Lycoriella aliena, Lycoriella gigaspina, Lycoriella neogenica, Corynoptera neutralis, Corynoptera ritzkowskii, Corynoptera tritospinosula, Epidapus brevihalteratus, Epidapus unistylatus, Succinosciara acuminata, Corynoptera klebsii, Epidapus menzeli, Epidapus palaeogracilis, Epidapus menzeli, Sciaridae indet. (129227) | |
| Priabonian | Russian Federation (Kaliningrad) | Sciara electriphila (140082) Sciara sendelina, Sciara verticillata, Sciara variabilis, Sciara preciosa, Sciara palmnickii (124073) Sciaridae indet. (124275) Trichosia sp., Sciara botuli, Sciara diabolica, Trichosia macrociliata, Sciara orientalis, Corynoptera sp., Sciara klebsii, Corynoptera subagilis, Sciara prolifica, Sciara splendida, Sciara errans, Sciara villosa, Sciara sendelina, Sciara difficilis, Sciara verticillata, Sciara variabilis, Sciara eocenica, Sciara rara, Sciara bella, Sciara ignorata, Sciara preciosa, Sciara rubsaamenia, Sciara tertiaria, Sciara robusta, Sciara morosa, Sciara palmnickii, Sciara minuscula, Sciara villosoides, Sciara tanypeza, Bradysia curiosa, Bradysia electra, Bradysia morosoides, Bradysia infernalis, Bradysia agilis, Bradysia umbrosa, Bradysia conwentzii, Heeriella bifurcata, Sciara defectuosa, Trichosia infausta, Trichosia microciliata, Trichosia protorotunda, Corynoptera hirticornis, Corynoptera koenigsbergensis, Corynoptera schumanni, Plastosciara keilbachi, Corynoptera dubia (123927) | |
| Chadronian | USA (Colorado) | Sciara dormitans (106379) Sciara requieta (106381) Sciara sopora (124681) | |
| MP 23 | France | Sciara sp., Sciara flichei (122535) | |
| Rupelian | France | Lycoria oligocoenica, Lycoria tertiaria, Protosciara alsatica (146907) | |
| Late/Upper Oligocene | France (Bouches-du-Rhone) | Sciara protogaea (122464) | |
| MP 30 | Germany | Sciara spp., Sciara janassa, Sciara defossa, Sciara atavina, Sciara rottensis, Sciara winnertzii (123945) | |
| MP 30 | Germany (Nordrhein-Westfalen) | Sciara janassa, Sciara heydeni (138696) | |
| Chattian | Germany (North Rhine-Westphalia) | Lycoria aulica, Lycoria weylandi, Lycoria austera, Lycoria pennata, Lycoria pelidua, Lycoria cryptocephala, Lycoria longa, Lycoria antennaria, Lycoria laticornis, Lycoria longipennata, Lycoria jucunda, Lycoria ovalipennata, Lycoria melanopa, Lycoria lengersdorfi, Lycoria gracilenta, Lycoria lugens, Lycoria bicolora, Lycoria compta, Lycoria femuralis, Lycoria thoracica, Lycoria gracilior, Lycoria amicula, Lycoria bella, Lycoria minuscula, Lycoria exilis (122010) | |
| Chattian | France (Provence-Alpes-Côte d'Azur) | Sciara inflata, Sciara acuminata, Sciara oblonga, Sciara minuscula (122487) Sciara protogaea, Sciara saportana, Sciara inflata, Sciara acuminata (122488) | |
| Chattian | France (Bouches-Du-Rhone) | Cecidomyia protogaea (122877) Sciara oblonga (122501) Sciara sp. (138838) | |
| Early/Lower Miocene | Mexico | Schwenckfeldina archoica (21977) | |
| Early/Lower Miocene | Mexico (Chiapas) | Bradysia sp. (124467 140214 140215) Sciara sp., Bradysia sp. (140260 140262) | |
| Early/Lower Miocene | Japan | Sciara sp. (114003) | |
| Burdigalian | China (Shandong) | Sciara atropha (128551) Sciara brevipetiola (128550) | |
| Burdigalian - Langhian | Dominican Republic | Archicratyna arcana (125970 155411) Corynoptera pristina, Trichosia nova, Leptosciarella manifesta, Cratyna tempestiva, Cratyna villosoantennata, Archicratyna trichoarcana, Archicratyna subarcana, Chaetosciara obsoleta, Bradysiopsis repentina, Bradysia antiqua, Bradysia exoleta, Corynoptera prisca, Epidapus microspinus, Epidapus macrospinatus, Epidapus adstrictosetus, Epidapus longisetus (122472) Cratyna interposita, Bradysia exoleta, Bradysia dimidiata, Corynoptera prisca, Corynoptera iocosa (123422) Trichosia nova, Leptosciarella manifesta, Cratyna interposita, Bradysia exoleta, Corynoptera prisca, Corynoptera iocosa, Epidapus microspinus (122982) | |
| MN 4 | Czech Republic | Sciara martii (117612) | |
| Langhian | USA (Idaho) | Sciara sepulta (97616) | |
| MN 5 | Germany (Baden-Württemberg) | Lycoria deffneri, Lycoria branconis, Lycoria endrissi, Lycoria quenstedti, Lycoria pompeckii, Lycoria fraasi, Lycoria hoelzlei, Lycoria dietleni, Lycoria berckhemeri, Lycoria braeuhaeuseri, Lycoria seemanni, Lycoria hennigi, Lycoria kluepfeli, Lycoria ehrati, Lycoria joossi, Lycoria wageri, Lycoria milleri, Lycoria lengersdorfi (127613) | |
| Middle Miocene - Late/Upper Miocene | Thailand | Sciaridae indet. (140983) | |
| Sarmatian | Croatia | Sciara acuminata, Sciara minutula (117761) Sciara hirtella (117250) | |
| Sarmatian | Germany (Baden-Württemberg) | Sciara deleta (117252) | |
| Messinian | Austria | Lycoria schremmeri (127417) | |
| Piacenzian | Germany (Niedersachsen) | Sciara willershausensis, Sciara strausi (109196) | |
| Piacenzian | Japan | Sciara sp. A, Sciara sp. B, Sciaridae indet. (188976) | |
| Middle Pleistocene | Japan | Sciaridae indet. (186848) | |
| Late/Upper Pleistocene | Argentina | Bradysia aliciae (183973) | |
| Holocene | Tanzania | Sciara radians (140788) Sciara variabilis (152189) |