PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Scaritinae

Insecta - Coleoptera - Carabidae

Taxonomy

Scaritides was named by Bonelli (1810). It is extant. Its type is Scarites.

It was corrected as Scaritinae by Lawrence and Newton (1995), Tamutis et al. (2011), Bouchard et al. (2011) and Cai et al. (2022).

It was assigned to Carabidae by Lawrence and Newton (1995), Tamutis et al. (2011), Bouchard et al. (2011) and Cai et al. (2022).

Subtaxa

Clivinini, Dyschiriini, Dyschiriomimus, Palaeoaxinidiini, Scaritini

Synonyms

Promecognathinae was named by Le Conte (1853). It is extant.

It was reranked as the tribe Promecognathini by LeConte (1861); it was synonymized subjectively with Scaritinae by Bouchard et al. (2011).

It was assigned to Carabidae by LeConte (1861); and to Carabidae by McKay (1991).

Synonymy list

Year	Name and author
1853	Promecognathinae Le Conte
1861	Promecognathini LeConte p. 11
1991	Promecognathinae McKay p. 3
1995	Scaritinae Lawrence and Newton, Jr.
2011	Scaritinae Bouchard et al.
2011	Scaritinae Tamutis et al.
2022	Scaritinae Cai et al.

Is something missing? Join the Paleobiology Database and enter the data

Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924

Rank	Name	Author
infraclass	Neoptera	Martynov 1923
—	Holometabola
—	Aparaglossata
—	Neuropteriformia
—	Coleopterida
order	Coleoptera
—	Mesocoleoptera
—	Metacoleoptera
suborder	Adephaga
family	Carabidae	()
subfamily	Scaritinae	()
subfamily	Scaritinae	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Subfm. Scaritinae Bonelli 1810

show all | hide all

Tr. Clivinini Rafinesque 1815

hide

G. Camptodontus Dejean 1826

Subtr. Clivinina Rafinesque 1815

G. Clivina Latreille 1802

Tr. Dyschiriini Kolbe 1880

hide

G. Dyschiriodes Jeannel 1941

Dyschiriodes aeneus Dejean 1825

Dyschiriodes globosus Herbst 1784

Dyschiriodes luticola Chaudoir 1850

Dyschiriodes nigricornis Motschulsky 1844

Invalid names: Dyschirius septentrionum Munster 1923 [synonym]

G. Dyschirius Bonelli 1810

Dyschirius laevifasciatus Horn 1878

Dyschirius melancholicus Putzeys 1867

Dyschirius montanus Le Conte 1879

Dyschirius thoracicus Rossi 1790

Invalid names: Dyschirius arenosus Stephens 1827 [synonym]

Dyschirius varidens Fall 1910

Invalid names: Helophorus rigescens Scudder 1890 [nomen dubium]

G. †Dyschiriomimus Yablokov-Khnzorian 1960

hide

†Dyschiriomimus stackelbergi Yablokov-Khnzorian 1960

Tr. †Palaeoaxinidiini McKay 1991

hide

G. †Palaeoaxinidium McKay 1991

†Palaeoaxinidium orapensis McKay 1991

Tr. Scaritini Bonelli 1810

hide

Subtr. Scaritina Bonelli 1810

G. Scarites Fabricius 1775

†Scarites mancus Zhang 1989

†Scarites robustiventris Theobald 1937

Invalid names: Scarites haidingeri Heer 1861 [nomen nudum]

Invalid names: Promecognathinae Le Conte 1853 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Turonian to the top of the Late/Upper Pleistocene or 93.90000 to 0.01170 Ma
Minimum age of oldest fossil (stem group age): 89.8 Ma

Collections (42 total)

Time interval	Ma	Country or state	Original ID and collection number
Turonian	93.9 - 89.8	Botswana	Palaeoaxinidium orapensis (123430)
MP 11	48.6 - 40.4	Germany (Hessen)	Scaritinae indet. (120798)
Priabonian	38.0 - 33.9	Lithuania	Dyschiriomimus stackelbergi (113470)
Priabonian	38.0 - 33.9	France	Scarites robustiventris (113709)
Burdigalian	20.44 - 15.97	China (Shandong)	Scarites mancus (128550)
Late/Upper Miocene	11.608 - 5.333	Russian Federation	Dyschiriodes sp. (113713 153434)
Messinian	7.246 - 5.333	USA (Alaska)	Dyschirius sp. (123352)
Gelasian	2.588 - 1.806	Greenland	Dyschirius spp. (160108 160144 160146 160151 160153) Dyschirius spp., Dyschirius varidens (160109)
Early/Lower Pleistocene	2.588 - 0.781	Russian Federation	Dyschiriodes nigricornis (163069)
Middle Pleistocene	0.774 - 0.129	United Kingdom	Dyschirius globosus (168265 168266 168267 168268 168271 168272) Dyschirius globosus, Dyschirius aeneus (168264 168269 168270)
Middle Pleistocene	0.774 - 0.129	United Kingdom (Warwickshire)	Dyschirius globosus (169389 169390)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Wisconsin)	Dyschirius sp. (169406)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Pennsylvania)	Dyschirius nigricornis (137209) Dyschirius sp. (137208)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Washington)	Dyschiriodes sp., Dyschiriodes laevifasciatus (157092) Dyschiriodes sp., Dyschiriodes montanus, Dyschiriodes laevifasciatus (167662)
Late/Upper Pleistocene	0.129 - 0.0117	Canada (British Columbia)	Dyschirius sp. (167673)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom (England)	Dyschirius globosus (164904 164906)
Late/Upper Pleistocene	0.129 - 0.0117	Russian Federation (Astrakhan)	Dyschirius sp., Dyschirius luticola (125794)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Colorado)	Dyschirius sp. (169336)
Late/Upper Pleistocene	0.129 - 0.0117	United Kingdom (Worcestershire)	Dyschirius sp., Dyschirius arenosus, Dyschirius septentrionum, Dyschirius globosus (163070)
Late/Upper Pleistocene	0.129 - 0.0117	USA (Ohio)	Helophorus rigescens (152574)
Tarantian	0.129 - 0.0117	USA (Pennsylvania)	Dyschirius integer group (137205) Dyschirius integer group, Dyschirius sp. (137206) Dyschirius sp. (137204) Dyschirius sp., Dyschirius nigricornis (137201)