PBDB Taxon

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Sulidae (booby)

Reptilia - Suliformes - Sulidae

Taxonomy

Sulidae was named by Reichenbach (1849). It is extant. Its type is Sula.

It was assigned to Steganopodes by Zittel (1890); to Pelecanoidea by Hay (1902); to Steganopodes by Case (1904); to Suloidae by Hay (1930); to Sulides by Lambrecht (1933); to Procellariiformes by Cheneval (1984); to Pelecani by Cheneval (1993); to Sulae by Howard (1968), Olson (1985), Carroll (1988) and Olson and Hasegawa (1996); to Pelecaniformes by Brodkorb (1955), Miller and Bowman (1958), Howard (1972), Olson (1975), Howard (1978), Olson and James (1982), Howard (1982), Olson (1985), Howard and Barnes (1987), Steadman et al. (1988), Chandler (1990), Olson and Rasmussen (2001), Stucchi (2003), Gohlich (2003) and Stucchi and Urbina (2004); to Pelecaniformes by Mayr (2005); to Suloidea by Wetmore (1940), Wetmore (1956), Mayr (2002), Mayr (2005), Bourdon et al. (2005) and Livezey and Zusi (2007); to Ardeiformes by Mlikovsky (2007); to Aequornithes by Mayr (2011); and to Suliformes by del Hoyo et al. (2014), Clements et al. (2017), Mayr (2022) and Figueiredo et al. (2023).

Subtaxa

Empheresula (syn. Parasula), Enkurosula (syn. Pseudosula), Eostega, Masillastega, Microsula, Miosula, Morus, Paleosula, Papasula, Prophalacrocorax, Ramphastosula, Sarmatosula, Sula (syn. Dysporus) (type genus)

Synonymy list

Year	Name and author
1849	Sulidae Reichenbach
1890	Sulidae Zittel p. 842
1902	Sulidae Hay p. 532
1904	Sulidae Case p. 58
1930	Sulidae Hay p. 291
1933	Sulidae Lambrecht p. 284
1940	Sulidae Wetmore p. 11
1955	Sulidae Brodkorb p. 7
1956	Sulidae Wetmore p. 14
1958	Sulidae Miller and Bowman p. 8
1968	Sulidae Howard p. 10
1972	Sulidae Howard p. 5
1975	Sulidae Olson p. 23
1978	Sulidae Howard p. 15
1982	Sulidae Howard p. 10
1982	Sulidae Olson and James p. 33
1984	Sulidae Cheneval p. 65
1985	Sulidae Olson p. 158
1987	Sulidae Howard and Barnes p. 3
1988	Sulidae Carroll
1988	Sulidae Steadman et al. p. 487
1990	Sulidae Chandler p. 112
1993	Sulidae Cheneval p. 89
1996	Sulidae Olson and Hasegawa p. 742
2001	Sulidae Olson and Rasmussen p. 260
2002	Sulidae Mayr p. 508
2003	Sulidae Gohlich p. 238
2003	Sulidae Stucchi p. 77
2004	Sulidae Stucchi and Urbina p. 974
2005	Sulidae Bourdon et al. p. 166
2005	Sulidae Mayr p. 67
2007	Sulidae Livezey and Zusi p. 91
2007	Sulidae Mlikovsky p. 20
2011	Sulidae Mayr p. 73 figs. Figure 5
2014	Sulidae del Hoyo et al.
2017	Sulidae Clements et al.
2022	Sulidae Mayr p. 119 figs. Fig. 7.1
2023	Sulidae Figueiredo et al. p. 26

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
—	Sauropsida
class	Reptilia
subclass	Eureptilia	()
—	Romeriida
—	Diapsida	()
—	Archosauromorpha	(Huene 1946)
—	Crocopoda

Rank	Name	Author
—	Archosauriformes	Gauthier 1986
—	Eucrocopoda
—	Archosauria	()
informal	Avemetatarsalia
—	Ornithodira
—	Dinosauromorpha
—	Dinosauriformes
—	Dinosauria	()
—	Theropoda	()
—	Neotheropoda
—	Averostra	Paul 2002
—	Tetanurae
—	Coelurosauria	()
—	Maniraptora
—	Paraves
class	Aves
subclass	Neornithes	Gadow 1893
—	Aequornithes
—	Feraequornithes
superorder	Pelecanimorphae	Huxley 1867
order	Suliformes	Sharpe 1891
family	Sulidae	Reichenbach 1849
family	Sulidae	Reichenbach 1849

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Sulidae Reichenbach 1849 [booby]

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G. †Empheresula Harrison 1975

†Empheresula arvernensis Milne-Edwards 1867

Invalid names: Parasula Harrison 1975 [synonym]

G. †Enkurosula Kašin 1977

Invalid names: Pseudosula Harrison 1975 [synonym]

G. †Eostega Lambrecht 1929

†Eostega lebedinskyi Lambrecht 1929

G. †Masillastega Mayr 2002

†Masillastega rectirostris Mayr 2002

G. †Microsula Wetmore 1938

†Microsula pygmaea Milne-Edwards 1874

G. †Miosula Miller 1925

†Miosula media Miller 1925

G. Morus Vieillot 1816 [gannet]

†Moris lompocana Miller 1925

†Moris loxostyla Cope 1870

†Moris reyana Howard 1936

†Morus atlanticus Shufeldt 1915

†Morus avitus Wetmore 1938

Morus bassanus Linnaeus 1758 [northern gannett]

Morus capensis Lichtenstein 1823 [Cape gannet]

†Morus humeralis Miller and Bowman 1958

†Morus magnus Howard 1978

†Morus olsoni Grigorescu and Kessler 1988

†Morus peninsularis Brodkorb 1955

†Morus peruvianus Stucchi 2003

†Morus poolensis Chandler 1963

†Morus recentior Howard 1949

†Morus reyanus Howard 1936

Morus serrator Gray 1843 [Australasian gannet]

†Morus vagabundus Wetmore 1930

G. †Paleosula Howard 1958

†Paleosula stocktoni Miller 1935

G. Papasula Olson and Warheit 1988

Papasula abbotti Ridgway 1893 [Abbott's booby]

†Papasula abbotti costelloi Steadman et al. 1988

G. †Prophalacrocorax Harrison 1975

†Prophalacrocorax ronzoni Gervais 1849

G. †Ramphastosula Stucchi and Urbina 2004

†Ramphastosula aguierrei Stucchi et al. 2016

†Ramphastosula ramirezi Stucchi and Urbina 2004

G. †Sarmatosula Grigorescu and Kessler 1977

†Sarmatosula dobrogensis Grigorescu and Kessler 1977

G. Sula Brisson 1760

Subg. Sula (Sula) Brisson 1760

†Sula brandi Stucchi et al. 2016

†Sula clarki Chandler 1990

Sula dactylatra Lesson 1831 [masked booby]

Sula dactylatra tasmani van Tets et al. 1988

†Sula figueroae Stucchi et al. 2016

Sula granti Rothschild 1902 [Nazca booby]

Sula leucogaster Boddaert 1783 [brown booby]

†Sula magna Stucchi 2003

Sula nebouxii Milne-Edwards 1882 [Blue-footed booby]

Sula piscator Linnaeus 1758

†Sula pohli Howard 1958

Sula sula Linnaeus 1766 [red-footed booby]

†Sula sulita Stucchi 2003

†Sula universitatis Brodkorb 1963

Sula variegata Tschudi 1843 [Peruvian booby]

†Sula willetti Miller 1925

Invalid names: Dysporus Illiger 1811 [synonym], Sula guano Brodkorb 1955 [nomen nudum], Sula phosphata Brodkorb 1955 [nomen nudum]

Diagnosis

No diagnoses are available