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Hyaenodontidae
Taxonomy
Hyaenodontidae was named by Leidy (1869). It is not extant.
It was assigned to Creodonta by Cope (1889); to Carnivora by Cope (1875) and Andrews (1906); to Creodonta by Flower and Lydekker (1891), Hay (1902) and Lambe (1908); to Creodonta by Carroll (1988), Sach and Heizmann (2001), Lange-Badré and Böhme (2005), Wang et al. (2005), Gheerbrant et al. (2006), Lavrov (2007), Morales et al. (2007), Morlo et al. (2007), Morales et al. (2008), Bajpai et al. (2009) and Rose et al. (2012); to Hyaenodontida by Solé et al. (2014) and Solé et al. (2014); to Hyaenodonta by Solé et al. (2015); to Hyaenodonta by Rana et al. (2015), Borths et al. (2016) and Zack (2019); and to Hyaenodontoidea by Van Valen (1967), Dubied et al. (2019) and Solé and Mennecart (2019).
It was assigned to Creodonta by Cope (1889); to Carnivora by Cope (1875) and Andrews (1906); to Creodonta by Flower and Lydekker (1891), Hay (1902) and Lambe (1908); to Creodonta by Carroll (1988), Sach and Heizmann (2001), Lange-Badré and Böhme (2005), Wang et al. (2005), Gheerbrant et al. (2006), Lavrov (2007), Morales et al. (2007), Morlo et al. (2007), Morales et al. (2008), Bajpai et al. (2009) and Rose et al. (2012); to Hyaenodontida by Solé et al. (2014) and Solé et al. (2014); to Hyaenodonta by Solé et al. (2015); to Hyaenodonta by Rana et al. (2015), Borths et al. (2016) and Zack (2019); and to Hyaenodontoidea by Van Valen (1967), Dubied et al. (2019) and Solé and Mennecart (2019).
Subtaxa
Acarictis, Acrohyaenodon, Ahknatenavus, Arfia (syn. Anthracoxyaena), Boritia, Cartierodon, Cynohyaenodon (syn. Pseudosinopa), Eurotherium, Galecyon, Gazinocyon, Hyaenaelurinae, Hyaenodontinae, Hyaenodontipus, Indohyaenodontinae, Ischnognathus, Koholiinae, Leonhardtina, Matthodon, Neohyaenodon, Oxyaenoides (syn. Francotherium), Paracynohyaenodon, Preregidens, Prionogalidae, Prodissopsalis (syn. Imperatoria), Protopsalis, Prototomus, Proviverroides, Pyrocyon, Quercytherium, Schizophagus, Sinopa (syn. Stypolophus, Triacodon), Thereutherium, Tinerhodon, Tritemnodon
Synonymy list
Year | Name and author |
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1869 | Hyaenodontidae Leidy |
1875 | Hyaenodontidae Cope p. 5–7 |
1889 | Hyaenodontidae Cope p. 876 |
1891 | Hyaenodontidae Flower and Lydekker p. 91 |
1902 | Hyaenodontidae Hay p. 756 |
1906 | Hyaenodontidae Andrews p. 218 |
1908 | Hyaenodontidae Lambe |
1967 | Hyaenodontidae Van Valen |
1988 | Hyaenodontidae Carroll |
2001 | Hyaenodontidae Sach and Heizmann p. 27 |
2005 | Hyaenodontidae Lange-Badré and Böhme p. 312 |
2005 | Hyaenodontidae Wang et al. p. 2 |
2006 | Hyaenodontidae Gheerbrant et al. p. 476 |
2007 | Hyaenodontidae Lavrov |
2007 | Hyaenodontidae Morales et al. p. 72 |
2007 | Hyaenodontidae Morlo et al. p. 146 |
2008 | Hyaenodontidae Morales et al. |
2009 | Hyaenodontidae Bajpai et al. p. 103 |
2012 | Hyaenodontidae Rose et al. |
2014 | Hyaenodontidae Solé et al. |
2014 | Hyaenodontidae Solé et al. |
2015 | Hyaenodontidae Rana et al. |
2015 | Hyaenodontidae Solé et al. |
2016 | Hyaenodontidae Borths et al. |
2019 | Hyaenodontidae Dubied et al. |
2019 | Hyaenodontidae Solé and Mennecart |
2019 | Hyaenodontidae Zack p. 17 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Fm. †Hyaenodontidae Leidy 1869
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G. †Acrohyaenodon Ameghino 1904
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†Acrohyaenodon acutidens Rovereto 1914
†Acrohyaenodon pungens Ameghino 1904
G. †Ahknatenavus Holroyd 1999
Subfm. †Arfiinae Solé et al. 2014
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G. †Arfia Van Valen 1965
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†Arfia gingerichi Smith and Smith 2001
†Arfia junnei Gingerich 1989
†Arfia langebadreae Lavrov and Lopatin 2004
†Arfia opisthotoma Matthew 1901
†Arfia palustris Tong and Wang 2006
†Arfia shoshoniensis Matthew and Granger 1915
†Arfia zele Gingerich and Deutsch 1989
Invalid names: Anthracoxyaena Tong and Wang 2006 [synonym]
G. †Cynohyaenodon Filhol 1873
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†Cynohyaenodon cayluxi Filhol 1873
†Cynohyaenodon lautricensis Lange-Badré 1978
†Cynohyaenodon magnus Crochet 1988
†Cynohyaenodon minor Filhol 1877
†Cynohyaenodon ruetimeyeri Lange-Badré 1979
†Cynohyaenodon trux Solé et al. 2014
Invalid names: Pseudosinopa Deperet 1917 [synonym]
G. †Eurotherium Polly and Lange-Badré 1993
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†Eurotherium matthesi Lange-Badré and Haubold 1990
†Eurotherium theriodis Van Valen 1965
Subfm. †Hyaenodontinae Trouessart 1885
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G. †Hyaenodon Laizer and Parieu 1838
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Subg. †Hyaenodon (Isohyaenodon) Savage 1965
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†Hyaenodon (Isohyaenodon) andrewsi Savage 1965
†Hyaenodon (Isohyaenodon) matthewi Savage 1965
†Hyaenodon (Isohyaenodon) pilgrimi Savage 1965
†Hyaenodon americanus Bravard 1858
†Hyaenodon bavaricus Dehm 1935
†Hyaenodon brachycephalus Osborn 1909
†Hyaenodon brevirostrus Macdonald 1970
†Hyaenodon chunkhtensis Dashzeveg 1985
†Hyaenodon crucians Leidy 1853
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Invalid names: Hyaenodon minutus Douglass 1902 [synonym], Hyaenodon paucidens Osborn and Wortman 1894 [synonym]
†Hyaenodon cruentus Leidy 1853
†Hyaenodon dubius Filhol 1873
†Hyaenodon eminus Matthew and Granger 1925
†Hyaenodon gigas Dashzeveg 1985
†Hyaenodon horridus Leidy 1853
†Hyaenodon leptocephalus Scott 1888
†Hyaenodon leptorhynchus Laizer and Parieu 1838
†Hyaenodon megaloides Mellett 1977
†Hyaenodon microdon Mellett 1977
†Hyaenodon minor Lange-Badré 1979
†Hyaenodon mongoliensis Dashzeveg 1964
†Hyaenodon montanus Douglass 1902
†Hyaenodon mustelinus Scott 1894
†Hyaenodon neimongoliensis Huang and Zhu 2002
†Hyaenodon pervagus Matthew and Granger 1924
†Hyaenodon raineyi Gustafson 1986
†Hyaenodon weilini Wang et al. 2005
†Hyaenodon yuanchuensis Young 1937
Invalid names: Protohyaenodon Stock 1933 [synonym], Pseudopterodon Schlosser 1887 [synonym], Taxotherium Blainville 1841 [synonym]
G. †Isphanatherium Lavrov and Averianov 1998
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†Isphanatherium ferganensis Lavrov and Averianov 1998
G. †Megalopterodon Dashzeveg 1964
G. †Propterodon Martin 1906
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†Propterodon morrisi Matthew and Granger 1924
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Invalid names: Propterodon irdinensis Matthew and Granger 1925 [synonym]
†Propterodon panganensis de Bonis et al. 2018
†Propterodon tongi Liu and Huang 2002
†Propterodon witteri Zack 2019
G. †Hyaenodontipus
Subfm. †Indohyaenodontinae Solé et al. 2014
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Subfm. †Koholiinae Crochet 1988
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G. †Leonhardtina Matthes 1952
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†Leonhardtina godinoti Solé et al. 2014
†Leonhardtina gracilis Matthes 1952
†Leonhardtina meridianum Solé et al. 2020
G. †Matthodon Lange-Badré and Haubold 1990
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†Matthodon menui Rich 1971
†Matthodon peignei Solé et al. 2020
†Matthodon tritens Lange-Badré and Haubold 1990
G. †Neohyaenodon Thorpe 1922
G. †Oxyaenoides Matthes 1967
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†Oxyaenoides aumelasiensis Solé et al. 2020
†Oxyaenoides bicuspidens Matthes 1967
†Oxyaenoides lindgreni Rich 1971
†Oxyaenoides schlosseri Rütimeyer 1891
Invalid names: Francotherium Rich 1971 [synonym]
G. †Paracynohyaenodon Martin 1906
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†Paracynohyaenodon magnus Crochet 1991
†Paracynohyaenodon schlosseri Martin 1906
Unr. †Prionogalidae Morales et al. 2008
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G. †Prionogale Schmidt-Kittler and Heizmann 1991
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†Prionogale breviceps Schmidt-Kittler and Heizmann 1991
G. †Prodissopsalis Matthes 1950
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†Prodissopsalis eocaenicus Matthes 1952
Invalid names: Imperatoria Matthes 1952 [synonym]
G. †Quercytherium Filhol 1880
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†Quercytherium simplicidens Solé et al. 2014
†Quercytherium tenebrosum Solé et al. 2014
G. †Schizophagus Lange-Badre 1975
Subfm. †Sinopinae Solé et al. 2014
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G. †Galecyon Gingerich and Deutsch 1989
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†Galecyon chronius Zack 2011
†Galecyon gallus Solé et al. 2013
†Galecyon mordax Matthew and Granger 1915
†Galecyon morloi Smith and Smith 2001
†Galecyon peregrinus Zack 2011
G. †Gazinocyon Polly 1996
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†Gazinocyon vulpeculus Matthew and Granger 1915
†Gazinocyon whitiae Cope 1882
G. †Prototomus Cope 1874
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†Prototomus deimos Gingerich and Deutsch 1989
†Prototomus girardoti Smith and Smith 2001
†Prototomus martis Gingerich and Deutsch 1989
†Prototomus minimus Smith and Smith 2001
†Prototomus multicuspis Cope 1875
†Prototomus phobos Gingerich and Deutsch 1989
†Prototomus robustus Matthew and Granger 1915
†Prototomus secundarius Cope 1875
G. †Pyrocyon Gingerich and Deutsch 1989
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†Pyrocyon dioctetus Gingerich and Deutsch 1989
†Pyrocyon strenuus Gingerich and Deutsch 1989
G. †Sinopa Leidy 1871
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†Sinopa aculeatus Cope 1872
†Sinopa ethiopica Andrews 1906
†Sinopa eximia Leidy 1873
†Sinopa grangeri Matthew 1906
†Sinopa insectivorus Cope 1871
†Sinopa jilinia Morlo et al. 2014
†Sinopa lania Matthew 1909
†Sinopa major Wortman 1902
†Sinopa minor Wortman 1902
†Sinopa pungens Cope 1872
†Sinopa viverrinus Cope 1874
Invalid names: Stypolophus Cope 1871 [synonym], Triacodon Marsh 1871 [synonym]
G. †Tritemnodon Matthew 1906
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†Tritemnodon agilis Marsh 1872
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Invalid names: Sinopa gracilis Wortman 1902 [synonym], Stypolophus brevicolcarabus Cope 1872 [synonym]
†Tritemnodon hians Cope 1877
†Tritemnodon strenuus Cope 1875
Invalid names: Hyaenaelurinae Pilgrim 1932 [empty]
Diagnosis
Reference | Diagnosis | |
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E. Gheerbrant et al. 2006 | Creodonta with upper molars either three or reduced to two; two front upper molars specialised as carnassial teeth either tuberculo-sectorial or completely sectorial; last upper molar, when present, transversely extended; all the lower molars specialised as carnassial teeth ; P\ two-rooted, except in some specialised genera; primitive forms with long and slender skulls; tail long and heavy; later forms with more robust skull, claws blunt; cursorial adaptations to a varying extent, [after Pilgrim 1932]. |
Measurements
No measurements are available
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Source: f = family, subc = subclass, c = class, subp = subphylum | |||||
References: Gunnell 1998, Lillegraven 1979, Carroll 1988, Ji et al. 2002, Hendy et al. 2009 |
Age range: base of the Selandian to the top of the Middle Miocene or 61.60000 to 11.60800 Ma
Collections (535 total)
Oldest occurrences | |||
Time interval | Ma | Country or state | Original ID and collection number |
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Selandian | Morocco | Boualitomus marocanensis (68176) Lahimia selloumi (133217) | |
Thanetian | Morocco (Meknès-Tafilalet) | Tinerhodon disputatum (88301 133241) | |
Thanetian - Early/Lower Eocene | France (Marne) | Hyaenodontidae indet., Arfia gingerichi (133367) | |
Late/Upper Paleocene | Mongolia | Sinopa sp. (133399) | |
Ypresian | France (Aude) | Preregidens langebadrae (78236) Proviverrinae n. gen. (182641) | |
Ypresian | India (Gujarat) | Indohyaenodon raoi (55494 180566) | |
Ypresian | France (Marne) | Francotherium lindgreni, Matthodon menui (181653) Matthodon menui (181639) Matthodon menui, Leonhardtina godinoti (35539) Oxyaenoides lindgreni, Oxyaena menui (35540) Preregidens langebadrae (35535) Prototomus girardoti (35537) | |
Ypresian | France (Oise) | Arfia gingerichi (182643) Prototomus minimus, Arfia gingerichi, Hyaenodontidae indet. (68174) | |
Ypresian | United Kingdom (England) | Prototomus minimus, Prototomus girardoti (38868) | |
Ypresian | France | Prototomus girardoti (125597) | |
Youngest occurrences |
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Time interval | Ma | Country or state | PBDB collection number |
Chattian | Switzerland (Canton Solothurn) | Hyaenodon compressus, Hyaenodon filholi (182805) | |
Early/Lower Miocene | Namibia (Karas) | H. unnamed species (168036) H. unnamed species, Namasector soriae (146601) | |
Early/Lower Miocene | Mongolia | Hyaenodontidae indet. (182090) | |
Burdigalian | Kenya | Hyaenodon andrewsi, Hyaenodon matthewi, Hyaenodon pilgrimi (21725) Hyaenodon matthewsi, Hyaenodon andrewsi, Hyaenodon pilgrimi (21726) Hyaenodon sp. (27858 38060 87341) | |
Burdigalian | Kenya (Northern Frontier District) | Hyaenodontidae indet., Hyaenodon andrewsi (122454) | |
Burdigalian | Uganda | Prionogale breviceps (28441) | |
Burdigalian | China (Gansu) | Hyaenodon sp. (42065) | |
Middle Miocene | Kenya | Hyaenodon andrewsi (21724) | |
Middle Miocene | Nepal | Hyaenodontidae indet. (48712) | |
Serravallian - Tortonian | Tunisia (Gafsa) | Hyaenodontidae indet. (35277 35281) |