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Basilosauridae

Mammalia - Artiodactyla - Basilosauridae

Taxonomy
Basilosauridae was named by Cope (1867). It was considered paraphyletic by Uhen and Gingerich (2001), Uhen (2004) and Uhen (2008) and Martinez-Cáceres and Muizon (2011).

It was synonymized subjectively with Zeuglodontidae by Mchedlidze (1970).

It was assigned to Zeuglodontia by Gill (1871), Gill (1872) and Gill (1873); to Zeuglodonta by Trouessart (1904); to Zeuglodontes by Hay (1902) and Hay (1930); to Cetacea by Sherman (1952) and Thewissen et al. (2001); to Archaeoceti by Bouetel and Muizon (2006); to Archaeoceti by Bianucci and Landini (2007); to Basilosauroidea by Rice (1998), Gingerich (2005), Rice (2009) and Abu El-kheir et al. (2013); to Archaeoceti by Miller (1923), Kellogg (1928), Kellogg (1936), Simpson (1945), Thurmond and Jones (1981), Barnes et al. (1985), Hooker (1986), Carroll (1988), Benton (1993), Fordyce and Barnes (1994), Fordyce et al. (1995), Uhen (1996), Kohler and Fordyce (1997), McKenna and Bell (1997), Bergqvist et al. (1999), Uhen and Gingerich (2001), Fordyce and de Muizon (2001), Rice (2002), Fordyce (2003), Geisler and Sanders (2003), Uhen (2004), Gingerich (2007), McLeod and Barnes (2008), Uhen (2008), Fordyce and Roberts (2009), Fitzgerald (2010), Gingerich (2012), Gingerich and Zouhri (2015), Gingerich (2015) and Gao and Ni (2015); and to Pelagiceti by Uhen (2008), Uhen (2010), Martinez-Cáceres and Muizon (2011), Uhen et al. (2011), Uhen (2013), Goldin et al. (2014), Marx et al. (2016), Buono et al. (2016), Martínez-Cáceres et al. (2017) and Berta (2017).

Synonyms
Synonymy list
YearName and author
1849Zeuglodontidae Bonaparte p. 618
1849Zeuglodon brachyspondylus Müller pp. pp. 20-33
1850Hydrarchidae Bonaparte p. 1
1850Hydrarchina Bonaparte p. 1
1851Zeuglodon trachyspondylus Koch p. 57
1852Pontogeneus Leidy
1852Pontogeneus priscus Leidy p. 52
1853Zeuglodontae Bronn p. 763
1853Zeuglodon brachyspondylus Bronn p. 770
1854Pontogeneus priscus Leidy p. 8
1867Basilosauridae Cope
1871Basilosauridae Gill p. 122
1872Basilosauridae Gill p. 93
1873Stegorhinidae Brandt p. 334
1873Basilosauridae Gill p. 27
1876Zeuglodon wanklyni Seeley
1881Balaenoptera juddi Seeley p. 278
1887Balaenoptera juddi Lydekker p. 41
1887Zeuglodontidae Lydekker p. 49
1889Zeuglodontidae Cope p. 876
1890Zeuglodontidae Cope p. 601
1890Zeuglodon brachyspondylus Cope p. 602
1890Zeuglodontidae Thompson p. 1
1891Zeuglodontidae Flower and Lydekker p. 89
1894Zeuglodontidae Zittel p. 167
1894Zeuglodon wanklyni Zittel p. 168
1898Zeuglodontidae Trouessart p. 1009
1898Zeuglodon brachyspondylus Trouessart p. 1010
1898Zeuglodon wanklyni Trouessart p. 1010
1898Balaenoptera juddi Trouessart p. 1077
1899Berardiopsis miocaenus Koch p. 209 figs. Fig. 1-2
1902Basilosauridae Hay p. 586
1902Basilosaurus brachyspondylus Hay p. 586
1903Zeuglodon wanklyni Stromer p. 87
1904Zeuglodontidae Case p. 4
1904Pontogeneus Palmer p. 558
1904Basilosauridae Trouessart p. 753
1904Basilosaurus brachyspondylus Trouessart p. 753
1904Basilosaurus wanklyni Trouessart p. 753
1904Balaenoptera juddi Trouessart p. 782
1906Zeuglodontidae Andrews p. 234
1908Zeuglodon brachyspondylus Stromer p. 31
1908Zeuglodontidae Stromer p. 148
1914Zeuglodon wanklyni Abel p. 220
1914Zeuglodontidae Abel p. 220
1919Zeuglodontidae Abel p. 751
1923Dorudontidae Miller
1923Basilosauridae Miller p. 40
1923Dorudontidae Miller p. 40
1925Zeuglodontidae Zittel p. 84
1925Zeuglodon brachyspondylus Zittel p. 85
1925Zeuglodon wanklyni Zittel p. 85
1928Basilosauridae Kellogg p. 32 figs. Table 1
1928Dorudontidae Kellogg p. 32 figs. Table 1
1930Basilosauridae Hay p. 569
1930Basilosaurus brachyspondylus Hay p. 570
1930Dorudontidae Hay p. 657
1931Zeuglodontidae Van Deinse p. 33
1936Zygorhiza juddi Kellogg
1936Basilosauridae Kellogg p. 13
1936Dorudontidae Kellogg p. 100
1936Pontogeneus brachyspondylus Kellogg p. 101
1936Zygorhiza wanklyni Kellogg p. 176
1936Pontogeneus Kellogg p. 248
1936Dorudontinae Slijper p. 540
1936Zeuglodontidae Slijper p. 540
1937Zeuglodontidae Benham p. 1
1945Basilosauridae Simpson p. 100
1945Dorudontidae Simpson p. 100
1952Basilosauridae Sherman p. 99
1952Basilosaurus brachyspondylus Sherman p. 99
1954Prozeuglodontidae Moustafa p. 87
1970Zeuglodontidae Mchedlidze p. 50
1973Dorudontidae Keyes p. 389
1977Dorudontidae Whitmore and Sanders p. 305
1981Dorudontidae Thurmond and Jones p. 192
1981Basilosauridae Thurmond and Jones p. 193
1981Pontogeneus Thurmond and Jones p. 196
1981Pontogeneus brachyspondylus Thurmond and Jones p. 196
1982Dorudontinae Fordyce p. 422
1985Basilosauridae Barnes et al. p. 25
1985Dorudontinae Barnes et al. p. 25
1986Basilosauridae Hooker p. 214
1986Dorudontidae Hooker p. 214
1986Zygorhiza wanklyni Hooker p. 342
1988Basilosauridae Carroll
1988Pontogeneus Carroll
1993Basilosauridae Benton p. 761
1994Basilosauridae Fordyce and Barnes p. 428 figs. Table 1
1994Dorudontinae Fordyce and Barnes p. 428
1995Basilosauridae Fordyce et al. p. 379
1995Dorudontinae Fordyce et al. p. 379
1996Basilosauridae Uhen p. 39
1996Dorudontinae Uhen p. 41
1997Basilosauridae Kohler and Fordyce p. 575
1997Dorudontinae Kohler and Fordyce p. 575
1997Basilosauridae McKenna and Bell p. 369
1997Dorudontidae McKenna and Bell p. 369
1997Pontogeneus McKenna and Bell p. 369
1998Basilosauridae Rice
1998Dorudontidae Rice
1998Dorudontinae Uhen p. 34
1998Pontogeneus Uhen p. 37
1998Pontogeneus brachyspondylus Uhen p. 37
1999Basilosauridae Bergqvist et al. p. 31
2001Dorudontidae Fordyce and de Muizon p. 104
2001Basilosauridae Fordyce and de Muizon p. 176
2001Basilosauridae Thewissen et al. p. 351 figs. Table 1
2001Basilosauridae Uhen and Gingerich p. 2
2001Dorudontinae Uhen and Gingerich p. 2
2002Basilosauridae Rice p. 231 figs. Table 1
2002Dorudontidae Rice p. 231 figs. Table 1
2002Pontogeneus Sepkoski, Jr.
2002Dorudontinae Uhen p. 79
2002Pontogeneus Uhen p. 79
2002Pontogeneus brachyspondylus Uhen p. 79
2003Basilosauridae Fordyce p. 156 figs. Figure 9.1
2003Basilosauridae Geisler and Sanders p. 27
2004Basilosauridae Uhen pp. 11-12
2004Dorudontinae Uhen p. 12
2004Pontogeneus Uhen p. 12
2004Pontogeneus brachyspondylus Uhen p. 12
2005Basilosauridae Gingerich p. 237 figs. Table 15.1
2006Basilosauridae Bouetel and Muizon p. 383
2006Dorudontinae Bouetel and Muizon p. 383
2007Basilosauridae Bianucci and Landini p. 45 figs. Table 2.1
2007Dorudontinae Bianucci and Landini p. 45 figs. Table 2.1
2007Basilosauridae Gingerich p. 366
2008Basilosauridae McLeod and Barnes p. 93
2008Dorudontinae McLeod and Barnes p. 93
2008Zygorhiza juddi McLeod and Barnes p. 93
2008Zygorhiza wanklyni McLeod and Barnes p. 93
2008Pontogeneus McLeod and Barnes p. 94
2008Pontogeneus brachyspondylus McLeod and Barnes p. 94
2008Basilosauridae Uhen p. 561
2008Dorudontinae Uhen p. 561
2008Basilosauridae Uhen p. 592 figs. Figure 4
2009Basilosauridae Fordyce and Roberts p. 553
2009Dorudontinae Fordyce and Roberts p. 553
2009Basilosauridae Rice p. 235 figs. Table 1
2009Dorudontinae Uhen p. 93
2010Basilosauridae Fitzgerald p. 370 figs. Table 1
2010Basilosauridae Uhen p. 206
2011Basilosauridae Martinez-Cáceres and Muizon
2011Pontogeneus Schouten p. 18
2011Pontogeneus brachyspondylus Schouten p. 18
2011Dorudontinae Uhen et al. p. 960
2011Basilosauridae Uhen et al. p. 966 figs. Figure 10
2012Basilosauridae Gingerich p. 313 figs. Figure 4
2012Basilosaurus wanklyni Kalmykov p. 179
2013Basilosauridae Abu El-kheir et al. p. 462
2013Basilosauridae Uhen
2014Pontogeneus Cooper et al. p. 91
2014Basilosauridae Goldin et al.
2015Basilosauridae Gao and Ni p. 156 figs. Table 1
2015Dorudontinae Gao and Ni p. 156 figs. Table 1
2015Basilosauridae Gingerich p. 164
2015Pontogeneus Gingerich p. 184
2015Pontogeneus brachyspondylus Gingerich p. 184
2015Pontogeneus priscus Gingerich p. 184
2015Basilosauridae Gingerich and Zouhri p. 278
2016Basilosauridae Buono et al. p. 299
2016Basilosauridae Marx et al. p. 101
2017Basilosauridae Berta p. 159
2017Pontogeneus Berta p. 159
2017Basilosauridae Martínez-Cáceres et al. p. 11

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
RankNameAuthor
Amniota
Synapsida()
Therapsida()
infraorderCynodontia()
Epicynodontia
infraorderEucynodontia
Probainognathia
Mammaliamorpha
Mammaliaformes
classMammalia
orderArtiodactyla()
Cetacea()
Pelagiceti
familyBasilosauridae
familyBasilosauridae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
M. D. Uhen 2004Members of the Basilosauridae lack M3 and sacral vertebrae.. Basilosaurids have accessory denticles on their adult premolars and molars and have a very reduced hindlimb. Members of the other families of archaeocetes, Protocetidae, Remingtonocetidae, and Ambulocetidae retain M3, have sacral vertebrae, and have large hind limbs. They also lack accessory denticles on their adult cheek teeth.
M. D. Uhen 2008Basilosauridae are distinguished from all other archaeocetes by: the loss of M3, the presence of multiple accessory denticles on the cheek teeth, loss of the sacrum, reduction of the hind limb, and rotation of the pelvis (Uhen, 1998a).
M. D. Uhen 2013Members of the family Basilo- sauridae possess two upper molars rather than the three typical of other archaeocetes, as well as multiple acces- sory denticles on their cheek teeth. Most basilosaurids also have longer bodies than other archaeocetes due to an increased numbers of thoracic and lumbar vertebrae. The thoracic vertebral count is 16 in Basilosaurus (Bebej et al., 2012), 20 in Cynthiacetus (Martínez-Cáceres and de Muizon, 2011), 17 in Dorudon (Uhen, 2004) and Zygorhiza (pers. obs. contra (Kellogg, 1936)). The complete thorac- ic count is unknown for Ancalecetus (probably 17, see Gin- gerich and Uhen, 1996 for discussion), Chrysocetus (at least 13), Masracetus, Ocucajea, Saghacetus (at least 15, pers. obs.). The thoracic vertebral count for Stromerius and Supayace- tus are unknown. The exception to this generalization is Basilotritus wardii, which has a thoracic vertebral count of 12 (Uhen, 2001), more similar to the thoracic count of protocetids than other basilosaurids (Uhen et al., 2011).
The lumbar vertebral count is 16 in Cynthiacetus (Martínez-Cáceres and de Muizon, 2011), 20 in Basilosau- rus (Gingerich et al., 1990) and Dorudon, (Uhen, 2004). The lumbar vertebral count is unknown for Basilotritus (Gol’din and Zvonok, 2013). All basilosaurids also have forelimbs modified into flippers. These forelimbs include: a fan shaped scapular blade; robust humerus, radius and ulna; elbow joints with limited rotation (which exclude the ability to pronate and supinate the forearm); and car- pal joints which also limit a wide range of motion. In basi- losaurids for which they are known (Basilosaurus (Lucas, 1900; Gingerich et al., 1990), Chrysocetus (Uhen and Gin- gerich, 2001), and Dorudon (Uhen, 2004)), the hindlimbs are greatly reduced and do not retain a bony connection to the vertebral column. The innominate of Basilotritus is not as reduced as those of other basilosaurids (see Uhen, 1999). See Uhen (1998, 2009) for thorough discussions of basilosaurid characteristics.
M. Martínez-Cáceres et al. 2017Basilosaurids are medium-to-large sized archaeocetes characterized principally by: 1) a at palate narrowing at the level of P4; 2) absence of M3; 3) anterior edge of the orbit dorsal to the level of P4 or P4/M1 diastema; 4) anterior process of the periotic strongly elliptical in cross-section; 5) triangular expan- sion on the posterolateral corner of the pars cochlearis; and 6) well developed ventrolateral tuberosity of the periotic.
Measurements
No measurements are available
Composition: hydroxyapatiteo
Form: roller-shapedo
Ontogeny: modification of partso
Environment: marineo
Locomotion: actively mobileo
Life habit: aquatico
Depth habitat: surfaceo
Diet: carnivoreo
Reproduction: viviparouso
Created: 2005-03-06 14:19:05
Modified: 2017-04-17 10:41:18
Source: o = order
Reference: Uhen 2004

Age range: base of the Bartonian to the top of the Kaiatan or 41.30000 to 36.00000 Ma

Collections (216 total)


Time interval Ma Country or state Original ID and collection number
Tertiary66.0 - 2.588Poland Zeuglodon puschii (77355)
Eocene56.0 - 33.9Ukraine Basilosaurus sp. (132383 132385 132389 132391) Platyosphys einori (57767) Zeuglodon sp. (132386)
Eocene56.0 - 33.9USA (Florida) Basilosauridae indet. (132705)
Eocene56.0 - 33.9USA (North Carolina) Basilosauridae indet. (132701 133023)
Eocene56.0 - 33.9Russian Federation Archaeoceti indet. (114208)
Eocene56.0 - 33.9USA (Mississippi) Basilosauridae indet. (131912) Basilosaurus cetoides (131913)
Middle Eocene48.6 - 37.2USA (North Carolina) Basilosauridae indet. (41822)
Lutetian - Bartonian47.8 - 38.0Antarctica (Seymour Island) Basilosauridae indet. (53666)
Lutetian - Bartonian47.8 - 38.0Ukraine Eocetus sp. (137757)
Bortonian43.0 - 37.2New Zealand (South Canterbury) Zygorhiza sp. (32900)
Bortonian - Kaiatan43.0 - 36.0New Zealand (South Canterbury) Archaeoceti indet. (79346)
Bortonian - Kaiatan43.0 - 36.0New Zealand (North Canterbury) Dorudontinae indet. (124710)
Bartonian41.3 - 38.0Austria Basilosauridae indet. (45817)
Bartonian41.3 - 38.0Antarctica Zeuglodon sp. (77358)
Bartonian41.3 - 38.0Egypt Basilotritus sp. (45626)
Bartonian41.3 - 38.0USA (Mississippi) Zygorhiza kochii (45625 61249)
Bartonian41.3 - 38.0Ukraine (Kirovograd) Basilosauridae indet. (132381) Basilotritus sp. (141002)
Bartonian41.3 - 38.0Pakistan (Punjab) Basilosaurus drazindai (11596) Basiloterus hussaini (13064)
Bartonian41.3 - 38.0United Kingdom (England) Basilosaurus sp. (42641) Zeuglodon wanklyni, Basilosaurus sp. (32915)
Bartonian41.3 - 38.0Ukraine (Luhansk) Basilosauridae indet., Basilotritus uheni (132393)
Bartonian41.3 - 38.0Germany (Bavaria) Eocetus sp. (71821)
Bartonian41.3 - 38.0USA (Virginia) Eocetus wardii (107512)
Bartonian41.3 - 38.0USA (North Carolina) Basilosauridae indet. (5386) Cynthiacetus maxwelli (60452) Eocetus wardii (7297 7299 75518 99753 132664) Zygorhiza kochii (5387)
Bartonian41.3 - 38.0Peru (Ica) Ocucajea picklingi (90022) Protocetidae indet. (73780) Supayacetus muizoni (90021)
Bartonian41.3 - 38.0Western Sahara Chrysocetus fouadassii, Platyosphys aithai (175734)
Bartonian41.3 - 38.0USA (Florida) Basilosaurus sp. (49882)
Bartonian - Priabonian41.3 - 33.9Egypt Dorudon atrox (124995 124996 124997 124998 125003 125014)
Bartonian - Priabonian41.3 - 33.9USA (Florida) Basilosaurus sp. (81604)
Bartonian - Priabonian41.3 - 33.9Netherlands Basilosauridae indet. (168386)
Priabonian38.0 - 33.9Peru Archaeoceti indet. (63838) Basilosauridae indet. (73779)
Priabonian38.0 - 33.9Russian Federation Basilosaurus sp. (123876) Zeuglodon paulsonii (77227)
Priabonian38.0 - 33.9Italy (Parma) Dorudon sp. (32992)
Priabonian38.0 - 33.9USA (Louisiana) Basilosaurus cetoides (32907) Basilosaurus cetoides, Zygorhiza kochii (32925) Basilosaurus cetoides, Zygorhiza kochii, Pontogeneus priscus (75638) Zygorhiza kochii (55251)
Priabonian38.0 - 33.9USA (Georgia) Basilosauridae indet. (132552 133024) Basilosauridae indet., Basilosaurus cetoides (132660) Basilosaurus sp. (138667) Cynthiacetus maxwelli, Zygorhiza kochii (49885) Dorudon serratus, Archaeoceti indet. (60451) Zygorhiza kochii (27508) Zygorhiza kochii, Basilosaurus cetoides (45640)
Priabonian38.0 - 33.9Antarctica Basilosauridae indet. (100411 179418) Basilosauridae indet., Zygorhiza sp. (45723)
Priabonian38.0 - 33.9Jordan Basilosaurus isis (32899)
Priabonian38.0 - 33.9USA (Alabama) Basilosauridae indet. (131914) Basilosaurinae indet. (124815) Basilosaurus cetoides (28986 45638 76685 89613 131804 131832) Cynthiacetus maxwelli, Basilosaurus sp. (45637) Hydrarchos harlani (77362) Zeuglodon cetoides (86949) Zeuglodon sp. (86948) Zeuglodon sp., Basilosaurus kochii (84754) Zygorhiza kochii (124707 131909 131910) Zygorhiza kochii, Dorudon sp., Basilosaurus cetoides, Pontogeneus brachyspondylus (28983)
Priabonian38.0 - 33.9USA (Tennessee) Basilosaurus cetoides (32902)
Priabonian38.0 - 33.9USA (Arkansas) Basilosaurus cetoides (122718) Basilosaurus cetoides, Zygorhiza kochii (16549)
Priabonian38.0 - 33.9Egypt (Fayum) Basilosaurus cetoides (175726) Basilosaurus isis, Dorudon atrox (13402) Basilosaurus isis, Dorudon atrox, Cynthiacetus maxwelli (13403) Basilosaurus isis, Prozeuglodon atrox, Zeuglodon intermedius (83938) Basilosaurus isis, Prozeuglodon sp. (13103) Dorudon atrox (124991) Masracetus markgrafi (153755) Saghacetus osiris, Prozeuglodon stromeri, Zeuglodon osiris (74740) Zeuglodon sp. (74738 74739) Zeuglodon sp., Saghacetus osiris (74737)
Priabonian38.0 - 33.9Ukraine (Kiev) Basilosaurus sp. (183530) Zeuglodon paulsonii (48026)
Priabonian38.0 - 33.9USA (Mississippi) Basilosauridae indet. (132666 132667 132844) Basilosaurus cetoides (32926 45627 45628 45629 45630 45631 45632 45633 45634 55689 55868 135787) Cynthiacetus maxwelli (6802) Zeuglodon sp. (84058 132820 132822 132823 132824 132825) Zygorhiza kochii (45639 55869) Zygorhiza sp. (132843)
Priabonian38.0 - 33.9Ukraine Platyosphys paulsonii (48027) Zeuglodon paulsonii (48028)
Priabonian38.0 - 33.9Netherlands Basilotritus sp. (162500)
Priabonian38.0 - 33.9Senegal Dorudontinae indet. (32990)
Priabonian38.0 - 33.9USA (Florida) Archaeoceti indet. (32918) Basilosauridae indet. (32920 132611 132702 132817) Basilosaurus brachyspondylus (57123) Basilosaurus cetoides (57122) Cynthiacetus maxwelli (60448 60449 60450) Zygorhiza sp., zeuglodont sp. (32916)
Priabonian38.0 - 33.9USA (South Carolina) Basilosauridae indet. (72046) Basilosauridae indet., Chrysocetus healyorum (122725) Basilosaurus sp., Archaeoceti indet., Zygorhiza sp. (32908) Dorudon serratus (13405 188994)
Priabonian38.0 - 33.9Western Sahara Basilosauridae indet., Basilosaurus sp., Dorudon sp. (96686) Basilosaurus isis (159110) Basilosaurus isis, Dorudon atrox (159112 159114) Basilosaurus isis, Dorudon atrox, Stromerius sp. (159115) Basilosaurus sp., Dorudon sp., Basilosauridae indet. (96688) Dorudon atrox, Stromerius sp. (159106) Dorudon sp. (159111) Saghacetus sp., Dorudon atrox (159113)
Priabonian38.0 - 33.9United Kingdom (England) Balaenoptera juddi (56937) Zygorhiza sp. (184754)
Priabonian38.0 - 33.9New Zealand (South Island) Dorudontinae indet. (87886)
Priabonian38.0 - 33.9Egypt Ancalecetus simonsi (28977) Basilosaurus cetoides (123919) Basilosaurus isis (79677 79678 79681 137568) Basilosaurus isis, Dorudon atrox (13102) Basilosaurus isis, Dorudon atrox, Masracetus sp. (96952) Dorudon atrox (79676 79680 79682 124990 124992 124993 124994 124999 125000 125001 125002 125004 125005 125006 125008 125010 125013 125015 125016 125017 125019 125020 125021 125022 125023 125025 125026 125028) Masracetus markgrafi (96953) Saghacetus osiris (49919) Stromerius nidensis (79699) Zeuglodon isis (49924 65974 79679) Zeuglodon osiris (47947 79700) Zeuglodon zitteli (79698)
Priabonian38.0 - 33.9Antarctica (Seymour Island) Dorudontinae indet. (100190)
Kaiatan - Runangan37.2 - 34.3New Zealand (Canterbury) Basilosauridae indet. (162502)
Late/Upper Eocene37.2 - 33.9Egypt Zeuglodon sensitivus, Dorudon osiris (62468) Zeuglodon sp. (40516)
Late/Upper Eocene37.2 - 33.9New Zealand (Canterbury) Zygorhiza sp., Dorudontinae indet. (45041)
Late/Upper Eocene37.2 - 33.9Romania Berardiopsis miocaenus (65135)