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Rhinocerotidae (rhinoceros)

Mammalia - Perissodactyla - Rhinocerotidae

Synonyms
Synonymy list
YearName and author
1821Rhinocerotidae Gray p. 306
1850Rhinocerotidae Bonaparte p. 1
1858Rhinoceros crassus Leidy p. 28
1873Rhinoceros crassus Leidy p. 220
1873Rhinocerotidae Leidy p. 328
1873Rhinoceros oregonensis Marsh p. 410
1879Anchisodon tubifer Cope
1881Rhinoceridae Cope p. 393
1883Rhinocerotidae Flower p. 185
1889Rhinoceridae Cope p. 876
1890Aphelops crassus Osborn
1891Rhinocerotidae Flower and Lydekker p. 90
1899Aceratherium oregonensis Matthew
1902Anchisodon tubifer Hay p. 640
1902Rhinocerotidae Hay p. 642
1902Diceratherium oregonense Hay p. 644
1902Aphelops crassus Hay p. 645
1908Rhinocerotidae Lambe
1908Diceratherium oregonensis Loomis
1913Rhinocerotidae Scott p. 291
1916Rhinoceridae Stefani p. 11
1918Aphelops crassus Matthew
1960Rhinocerotidae Wood pp. 88-90
1966Rhinocerotidae Radinsky p. 631
1977Forstercooperia crudus Gabounia
1981Rhinocerotidae Thurmond and Jones p. 186
1988Rhinocerotidae Carroll
1989Rhinocerotidae Prothero and Schoch p. 535
1990Peraceras crassus Voorhies
1991Rhinocerotidae Nowak
1993Gigantamynodon akespensis Bayshashov
1996Rhinocerotidae Dashzeveg p. 8
1999Rhinocerotidae Holbrook p. 346
2000Rhinocerotidae Antoine et al. p. 250
2001Rhinocerotidae Sach and Heizmann p. 30
2002Rhinocerotidae Antoine et al.
2003Rhinocerotidae Antoine
2003Rhinocerotidae Antoine et al.
2003Rhinocerotidae Deng p. 142
2005Rhinocerotidae Geraads et al. p. 531
2005Rhinocerotidae Prothero
2006Rhinocerotidae Hulbert and Whitmore p. 18
2006Rhinocerotidae Symeonidis et al.
2010Rhinocerotidae Antoine et al.
2010Rhinocerotidae Becker et al.
2013Rhinocerotidae Böhme et al.
2014Rhinocerotidae Athanassiou et al.
2014Rhinocerotidae Khan et al.
2016Rhinocerotidae Lu et al.
2016Rhinocerotidae Pandolfi and Maiorino
2016Rhinocerotidae Wang et al.
2017Rhinocerotidae Geraads
2018Rhinocerotidae Tsoukala
2020Rhinocerotidae Lofgren et al.

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
Mammaliaformes
RankNameAuthor
classMammalia
Theriamorpha(Rowe 1993)
Theriiformes()
Trechnotheria
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Euungulata
Panperissodactyla
superorderPerissodactylamorpha
orderPerissodactyla()
Tapiromorpha(Haeckel 1873)
Ceratomorpha(Wood 1937)
superfamilyRhinocerotoidea
familyRhinocerotidae
familyRhinocerotidae

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. Rhinocerotidae Gray 1821 [rhinoceros]
show all | hide all
Subfm. †Aceratheriinae Dollo 1885
+
G. †Acerorhinus Kretzoi 1942
+
Acerorhinus fuguensis Deng 2000
Acerorhinus lufengensis Deng and Qi 2009
Acerorhinus neleus Athanassiou et al. 2014
Acerorhinus palaeosinensis Ringström 1924
Acerorhinus tsaidamensis Bohlin 1937
Acerorhinus zernowi Borissiak 1914
G. †Aphelops Owen 1845
+
Aphelops cristalatus Henshaw 1942
Aphelops longipes Leidy 1890
Aphelops malacorhinus Cope 1878
Aphelops megalodus Cope 1873
Aphelops mutilus Matthew 1924
+
Invalid names: Aphelops kimballensis Tanner 1967 [synonym], Aphelops longinaris Frye et al. 1956 [synonym]
Invalid names: Aphelops jemezanus Cope 1875 [nomen dubium]
G. †Chilotheridium Hooijer 1971
+
Chilotheridium pattersoni Hooijer 1971
Tr. †Chilotheriini Qiu et al. 1987
+
G. †Shansirhinus Kretzoi 1942
+
Shansirhinus brancoi Schlosser 1903
Shansirhinus ringstromi Kretzoi 1942
+
Invalid names: Acerorhinus cornutus Qiu and Yan 1982 [synonym], Chilotherium tianzhuensis Zheng 1982 [synonym], Chilotherium yunnanensis Tang et al. 1974 [synonym]
G. †Floridaceras Wood 1964
+
Floridaceras whitei Wood 1964
G. †Galushaceras Prothero 2005
+
Galushaceras levellorum Prothero 2005
G. †Hoploaceratherium Ginsburg and Heissig 1989
+
G. †Peraceras Cope 1880
+
Peraceras hessei Prothero and Manning 1987
Peraceras profectum Matthew 1899
+
Invalid names: Aphelops brachyodus Osborn 1904 [synonym], Aphelops ceratorhinus Douglass 1903 [synonym], Aphelops montanus Douglass 1908 [synonym], Diceratherium jamberi Tanner 1977 [synonym]
+
Invalid names: Peraceras troxelli Matthew 1918 [synonym]
G. †Plesiaceratherium Young 1937
+
Plesiaceratherium lumiarense Antunes and Ginsburg 1983
G. †Proaceratherium Ginsburg and Hugueney 1980
G. †Subchilotherium Heissig 1972
+
Subchilotherium intermedium Lydekker 1884
Tr. †Aceratheriini Dollo 1885
+
G. †Aceratherium Kaup 1834
+
Aceratherium incisivum Cuvier 1824
Aceratherium perimense Falconer and Cautley 1847
Invalid names: Aceratherium persistens Osborn 1904 [nomen dubium]
G. †Alicornops Ginsburg and Guerin 1979
+
Alicornops alfambrense Cerdeño and Alcalá 1989
Alicornops complanatum Heissig 1972
Alicornops simmorense Lartet 1851
G. †Chilotherium Ringström 1924
+
Chilotherium anderssoni Ringström 1924
Chilotherium haberi Ringström 1924
Chilotherium intermedium Heissig 1972
Chilotherium kiliasi Geraads and Koufos 1990
Chilotherium wimani Ringström 1924
G. †Persiatherium Pandolfi 2015
+
Persiatherium rodleri Pandolfi 2015
G. †Amphicaenopus Wood 1927
+
Amphicaenopus platycephalus Osborn and Wortman 1894
G. Atelodus Pomel 1853
+
Atelodus antiquitatis Blumenbach 1872
Subfm. †Diceratheriinae Dollo 1885
+
G. †Skinneroceras Prothero 2005
+
Skinneroceras manningi Prothero 2005
Subtr. †Elasmotheriina Bonaparte 1850
+
G. †Bugtirhinus Antoine and Welcomme 2000
+
Bugtirhinus praecursor Antoine and Welcomme 2000
G. †Caementodon Heissig 1972
+
Subg. †Caementodon (Caementodon) Heissig 1972
+
G. †Diceratherium Marsh 1875
+
+
Invalid names: Diceratherium cuspidatum Troxell 1921 [synonym], Diceratherium nanum Marsh 1875 [synonym]
Diceratherium armatum Marsh 1875
+
Invalid names: Aceratherium truquianum Cope 1879 [synonym], Diceratherium gregorii Peterson 1920 [synonym], Diceratherium lobatum Troxell 1921 [synonym]
Diceratherium minus Ringström 1924
Diceratherium niobrarense Peterson 1906
+
Invalid names: Aceratherium egregius Cook 1908 [synonym], Diceratherium petersoni Loomis 1908 [synonym], Epiaphelops virgasectus Cook 1912 [synonym]
Diceratherium niobrarensis Peterson 1906
Diceratherium persistens Osborn 1904
Diceratherium radtkei Prothero and Rasmussen 2008
+
Caenopus tridactylus avus Troxell 1921
Invalid names: Diceratherium avum Wood 1927 [synonym], Diceratherium proavitum Hatcher 1894 [synonym]
Invalid names: Epiaphelops Cook 1912 [synonym], Metacaenopus Cook 1908 [synonym]
Tr. †Elasmotheriini Dollo 1885
+
G. †Aegyrcitherium Antoine 1997
G. †Elasmotherium Fischer 1808
+
Elasmotherium caucasicum Borissiak 1914
Elasmotherium sibiricum Fischer 1809
G. †Gobitherium Kretzoi 1943
Subtr. †Menoceratina Prothero et al. 1986
+
G. †Menoceras Troxell 1921
+
Menoceras arikarense Barbour 1906
+
Invalid names: Aceratherium stigeri Loomis 1908 [synonym], Diceratherium aberrans Loomis 1908 [synonym], Diceratherium loomisi Cook 1912 [synonym], Diceratherium schiffi Loomis 1908 [synonym], Menoceras cooki Peterson 1906 [synonym]
Menoceras barbouri Wood 1964
+
Invalid names: Menoceras falkenbachi Tanner 1972 [synonym], Menoceras marslandensis Tanner 1969 [synonym], Moschoedestes delahoensis Stevens et al. 1969 [synonym]
Invalid names: Moschoedestes Stevens et al. 1969 [synonym]
G. †Parelasmotherium Killgus 1923
+
G. †Hispanotherium Crusafont and Villalta 1947
+
Subg. †Hispanotherium (Hispanotherium) Crusafont and Villalta 1947
Hispanotherium beonense Antoine 1997
Hispanotherium corcolense Antoine et al. 2002
Hispanotherium matritense de Prado 1864
+
Invalid names: Begertherium borissiaki Beliajeva 1971 [synonym], Caementodon tongxinensis Guan 1988 [synonym], Chilotherium quintanelensis Zbyszewsky 1952 [synonym], Hispanotherium alpani Sarac 1978 [synonym], Hispanotherium grimmi Heissig 1974 [synonym], Huaqingtherium qiui Guan 1993 [synonym]
Hispanotherium tungurense Cerdeño 1996
Invalid names: Begertherium [synonym], Tesselodon Yan 1979 [synonym]
G. †Huaqingtherium Huang and Yan 1983
+
G. †Iranotherium Ringström 1924
+
Iranotherium morgani Mecquenem 1908
G. †Kenyatherium Aguirre and Guerin 1974
+
Kenyatherium bishopi Aguirre and Guérin 1974
G. †Procoelodonta Matthew 1931
+
Subg. †Procoelodonta (Procoelodonta) Matthew 1931
G. †Sinotherium Ringstrom 1924
+
Sinotherium lagrelii Ringström 1923
G. †Gulfoceras Albright 1999
+
Gulfoceras westfalli Albright 1999
Subfm. †Iranotheriinae Kretzoi 1942
+
G. †Ninxiatherium Chen 1977
+
G. †Ougandatherium Guérin and Pickford 2003
+
Ougandatherium napakense Guérin and Pickford 2003
G. †Penetrigonias Tanner and Martin 1976
+
Penetrigonias dakotensis Peterson 1920
+
Invalid names: Penetrigonias hudsoni Tanner and Martin 1976 [synonym]
Penetrigonias sagittatus Russell 1982
Subfm. Rhinocerotinae Gray 1821
+
G. †Dromoceratherium Crusafont et al. 1955
+
Dromoceratherium mirallesi Crusafont et al. 1955
G. †Epiaceratherium Borissiak 1915
+
Epiaceratherium magnum Uhlig 1999
Epiaceratherium naduongense Böhme et al. 2013
G. †Guixia You 1977
G. †Mesaceratherium Keissig 1969
+
Mesaceratherium welcommi Antoine et al. 2010
G. †Molassitherium Becker et al. 2013
+
+
Invalid names: Diceratherium kuntneri Spillmann 1969 [synonym]
Molassitherium delemontense Becker et al. 2013
G. †Pleuroceros Roger 1898
+
Pleuroceros asphaltense Depéret and Douxami 1902
Pleuroceros blanfordi Lydekker 1884
Pleuroceros pleuroceros Duvernoy 1853
G. †Protaceratherium Abel 1910
+
Protaceratherium minutum Osborn 1900
+
G. Ceratotherium Gray 1868 [white rhino]
+
Ceratotherium cottoni Lydekker 1908 [northern white rhinoceros]
Ceratotherium neumayri Osborn 1900
+
Invalid names: Diceros pachygnathus Guéran 1980 [synonym]
Ceratotherium simum Burchell 1821 [white rhinoceros]
G. †Coelodonta Bronn 1831 [wooly rhino]
+
Coelodonta antiquitatis Blumenbach 1799
+
Invalid names: Coelodonta boiei Bronn 1831 [synonym], Rhinoceros tichorhinus Cuvier 1822 [synonym]
Coelodonta thibetana Deng et al. 2011
Coelodonta tologoijensis Beliajeva 1966
Invalid names: Tichorhinus Brandt 1849 [synonym]
G. Dicerorhinus Gloger 1841
+
Dicerorhinus cixianensis Chen and Wu 1976
Dicerorhinus sumatrensis Fischer 1814 [Sumatran rhino]
Invalid names: Didermocerus Brookes 1828 [synonym]
G. Diceros Gray 1821 [black rhino]
+
Diceros bicornis Linnaeus 1758 [black rhinoceros]
Subtr. †Dicerotina
+
G. †Paradiceros Hooijer 1968
+
Paradiceros mukirii Hooijer 1968
G. †Dihoplus Brandt 1878
+
Dihoplus kirchbergensis Jäger 1839
+
Invalid names: Rhinoceros mercki Kaup 1841 [synonym], Rhinoceros merckii Kaup 1841 [synonym]
Dihoplus megarhinus de Christol 1835
Dihoplus pikermiensis Toula 1906
Dihoplus ringstroemi Arambourg 1959
G. †Lartetotherium Ginsburg 1974
+
G. Rhinoceros Linnaeus 1758 [one-horned rhinoceros]
+
Rhinoceros deccanensis Foote 1874
Rhinoceros goldfussii Kaup 1834
Rhinoceros habereri Schlosser 1903
Rhinoceros iravadicus Lydekker 1876
Rhinoceros leptodon Kaup 1834
Rhinoceros leptorhinus Cuvier 1822
Rhinoceros minutus Cuvier 1830
Rhinoceros namadicus Falconer and Cautley 1847
Rhinoceros pacificus Leidy 1872
Rhinoceros palaeindicus Falconer and Cautley 1847
Rhinoceros philippinensis von Koenigswald 1956
Rhinoceros planidens Lydekker 1876
Rhinoceros platyrhinus Falconer and Cautley 1847
Rhinoceros plicidens Koken 1885
Rhinoceros sinensis Owen 1870
Rhinoceros sivalensis Falconer and Cautley 1847
Rhinoceros sondaicus Desmarest 1822 [Javan rhinoceros]
Rhinoceros unicornis Linnaeus 1758 [Indian rhinoceros]
Invalid names: Rhinoceros elatus Croizet and Jobert 1828 [nomen oblitum]
Subtr. †Rhinocerotina Gray 1825
+
G. †Gaindatherium Colbert 1934
+
Gaindatherium browni Colbert 1934
Gaindatherium vidali Heissig 1972
G. †Ronzotherium Aymard 1854
+
Ronzotherium filholi Osborn 1900
Ronzotherium orientale Dashzeveg 1991
Ronzotherium velaunum Aymard 1853
G. †Shennongtherium Huang and Yan 1983
+
Shennongtherium hypsodontus Huang and Yan 1983
G. †Stephanorhinus Kretzoi 1942
+
Stephanorhinus etruscus Falconer 1868
Stephanorhinus hemitoechus Falconer 1868
Stephanorhinus jeanvireti Guérin 1972
Stephanorhinus miguelcrusafonti Guerin and Santafe 1978
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Invalid names: Rhinoceros schleiermacheri samius Toula 1906 [synonym]
Invalid names: Rhinoceros schleiermacheri orientalis Schlosser 1921 [synonym]
Subtr. †Teleoceratina Hay 1902
+
G. †Aprotodon Forster-Cooper 1915
+
Aprotodon aralensis Beliajeva 1954
Aprotodon fatehjangense Pilgrim 1912
Aprotodon lanzhouensis Qiu and Xie 1997
Aprotodon smithwoodwardi Forster-Cooper 1915
G. †Brachydiceratherium Lavocat 1952
G. †Brachypodella Heissig 1973
G. †Brachypotherium Roger 1904
+
G. †Diaceratherium Dietrich 1931
+
Diaceratherium aginense Répelin 1917
Diaceratherium armatum Marsh 1875
Diaceratherium massiliae Ménouret and Guérin 2009
G. †Prosantorhinus Heissig 1973
+
Prosantorhinus douvillei Osborn 1902
G. †Teleoceras Hatcher 1894
+
Subg. †Teleoceras (Mesoceras) Cook 1930
Teleoceras aepysoma Short et al. 2019
Teleoceras americanum Yatkola and Tanner 1979
Teleoceras brachyrhinum Prothero 2005
Teleoceras felicis Freudenberg 1921
Teleoceras fossiger Cope 1878
+
Invalid names: Aceratherium acutum Marsh 1887 [synonym]
Teleoceras guymonense Prothero 2005
Teleoceras hicksi Cook 1927
+
Invalid names: Paraphelops rooksensis Lane 1927 [synonym], Peraceras ponderis Cook 1930 [synonym], Teleoceras ocotensis Dalquest and Mooser 1980 [synonym], Teleoceras schultzi Tanner 1975 [synonym], Teleoceras yumensis Cook 1930 [synonym]
Teleoceras major Hatcher 1894
Teleoceras medicornutum Osborn 1904
+
Invalid names: Aphelops planiceps Osborn 1904 [synonym], Teleoceras (Mesoceras) thomsoni Cook 1930 [synonym], Teleoceras thomsoni Cook 1930 [synonym]
Teleoceras meridianum Leidy 1865
+
Invalid names: Teleoceras minor Olcott 1909 [synonym]
Teleoceras proterum Leidy 1885
+
Invalid names: Eusyodon maximus Leidy 1886 [synonym]
Teleoceras pugnator Matsumoto 1921
Teleoceras snowi Fourtau 1918
Invalid names: Eusyodon Leidy 1886 [synonym], Paraphelops Lane 1927 [synonym]
Invalid names: Dicerorhinini [empty]
G. †Subhyracodon Wood 1927
+
Subhyracodon kewi Stock 1933
Subhyracodon mitis Cope 1875
+
Invalid names: Aceratherium copei Osborn 1898 [synonym], Aceratherium pumilum Cope 1885 [synonym], Caenopus pumilus Cope 1886 [synonym]
+
Caenopus trigonodus allus Troxell 1921
Invalid names: Aceratherium exiguum Lambe 1908 [synonym], Aceratherium trigonodum Osborn and Wortman 1894 [synonym], Hyracodon quadriplicatus Cope 1873 [synonym], Subhyracodon gidleyi Wood 1927 [synonym], Subhyracodon hesperius Leidy 1865 [synonym], Subhyracodon metalophum Wood 1927 [synonym]
Subhyracodon woodi Schlaikjer 1935
Invalid names: Anchisodon Cope 1879 [synonym], Caenopus Cope 1880 [synonym], Hyracodon planiceps Scott and Osborn 1887 [nomen dubium], Leptaceratherium Osborn 1898 [synonym]
G. †Teletaceras Hanson 1989
+
Teletaceras borissiaki Beliaeva 1959
Teletaceras radinskyi Hanson 1989
G. †Trigonias Lucas 1900
+
Trigonias osborni Lucas 1900
+
Invalid names: Caenopus premitis Gregory and Cook 1928 [synonym], Trigonias cooki Wood 1931 [synonym], Trigonias hypostylus Gregory and Cook 1928 [synonym], Trigonias paucidens Wood 1927 [synonym], Trigonias precopei Gregory and Cook 1928 [synonym], Trigonias preoccidentalis Gregory and Cook 1928 [synonym], Trigonias taylori Gregory and Cook 1928 [synonym], Trigonias tetradactylum Osborn 1923 [synonym]
Trigonias wellsi Wood 1927
+
Invalid names: Trigonias gregoryi Wood 1927 [synonym]
Invalid names: Procaenopus Figgins 1934 [synonym]
G. †Uintaceras Holbrook and Lucas 1997
+
Uintaceras radinskyi Holbrook and Lucas 1997
G. †Woodoceras Prothero 2005
+
Woodoceras brachyops Prothero 2005
G. †Zalabis
Invalid names: Anchisodon tubifer Cope 1879 [nomen dubium], Dicerorhininae Simpson 1945 [empty], Forstercooperia crudus Gabounia 1977 [nomen dubium], Gigantamynodon akespensis Bayshashov 1993 [nomen dubium], Rhinoceros crassus Leidy 1858 [nomen dubium], Rhinoceros oregonensis Marsh 1873 [nomen dubium], Teleoceratinae [empty]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: ground dwellingf
Diet: browserf
Reproduction: viviparoussubc
Comments: life habit based on Nowak 1999; diet is primitively browsing according to Prothero 1998f
Created: 2005-06-08 10:11:09
Modified: 2005-08-31 17:08:31
Source: f = family, subc = subclass, c = class, subp = subphylum
References: Carroll 1988, Nowak 1991, Lillegraven 1979, Ji et al. 2002, Hendy et al. 2009

Age range

Maximum range based only on fossils: base of the Bartonian to the top of the Holocene or 41.30000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 40.4 Ma

Collections (1912 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Middle Eocene48.6 - 37.2China (Yunnan) Rhinocerotidae indet. (36668)
Middle Eocene48.6 - 37.2USA (Oregon) Teletaceras radinskyi (16842)
Middle Eocene48.6 - 37.2China (Nei Mongol) Rhinocerotidae indet. (28786)
Sharamurunian48.6 - 37.2Russian Federation Eotrigonias borissiaki (192038)
Middle Eocene - Late/Upper Eocene48.6 - 33.9China (Nei Mongol) Rhinocerotidae indet. (180873)
Uintan46.2 - 40.4USA (Wyoming) Uintaceras radinskyi (16648)
Uintan46.2 - 40.4USA (Utah) Uintaceras radinskyi (16607)
Bartonian41.3 - 38.0Myanmar Teletaceras sp. (41393)
Bartonian41.3 - 38.0China (Henan) Rhinocerotidae indet. (177036)
Bartonian - Priabonian41.3 - 33.9Vietnam (Lang Son) Epiaceratherium naduongense (169420)
Bartonian - Rupelian41.3 - 28.1Mexico (Oaxaca) Subhyracodon sp., Trigonias sp. (166797)
Duchesnean40.4 - 37.2USA (Texas) Trigonias sp. (16835 16839)
Duchesnean40.4 - 37.2Canada (Saskatchewan) Rhinocerotidae indet. (16845)
Duchesnean40.4 - 37.2USA (Utah) Penetrigonias dakotensis (16847)
Duchesnean - Chadronian40.4 - 33.9USA (Montana) Rhinocerotidae indet. (16840)
Priabonian38.0 - 33.9USA (South Carolina) Subhyracodon mitis (206997)
Chadronian37.2 - 33.9Canada (Saskatchewan) Caenopus occidentalis, Caenopus pumilus (101961) Penetrigonias sagittatus (16979) Penetrigonias sagittatus, Trigonias osborni, Aceratherium exiguum (16919)
Chadronian37.2 - 33.9USA (Colorado) Aceratherium mite, Penetrigonias dakotensis, Trigonias osborni (16977) Caenopus premitis, Trigonias hypostylus, Trigonias osborni, Trigonias precopei, Trigonias preoccidentalis, Trigonias taylori (99642) Rhinocerotidae indet. (16968)
Chadronian37.2 - 33.9USA (South Dakota) Aceratherium trigonodum, Caenopus mitis, Trigonias gregoryi, Trigonias wellsi (17006) Caenopus mitis, Trigonias osborni (16938) Trigonias osborni (16936) Trigonias paucidens (17012) Trigonias sp., Caenopus mitis (16885) Trigonias wellsi (16937)
Chadronian37.2 - 33.9Mexico (Chihuahua) Rhinocerotidae indet. (17015)