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Hegetotheriidae
Taxonomy
Hegetotheriidae was named by Ameghino (1894). It was considered monophyletic by Croft and Anaya (2006).
It was corrected as Hegetotheridae by Sinclair (1908); it was reranked as the unranked clade Hegetotheriidae by Seoane et al. (2023).
It was assigned to Tipotheria by Ameghino (1897); to Hegetotheroidea by Carroll (1988); to Typotherioidea by Reguero and Prevosti (2010); to Notoungulata by Simpson and Minoprio (1949), Simpson et al. (1962), Croft and Anaya (2006), Kramarz et al. (2010) and Vera and Ercoli (2018); and to Typotheria by Ameghino (1894), Sinclair (1908), Simpson (1934), Billet (2011) and Seoane et al. (2023).
It was corrected as Hegetotheridae by Sinclair (1908); it was reranked as the unranked clade Hegetotheriidae by Seoane et al. (2023).
It was assigned to Tipotheria by Ameghino (1897); to Hegetotheroidea by Carroll (1988); to Typotherioidea by Reguero and Prevosti (2010); to Notoungulata by Simpson and Minoprio (1949), Simpson et al. (1962), Croft and Anaya (2006), Kramarz et al. (2010) and Vera and Ercoli (2018); and to Typotheria by Ameghino (1894), Sinclair (1908), Simpson (1934), Billet (2011) and Seoane et al. (2023).
Synonyms
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Synonymy list
| Year | Name and author |
|---|---|
| 1894 | Hegetotheriidae Ameghino |
| 1894 | Hegetotheridae Ameghino p. 19 |
| 1897 | Hegetotheridae Ameghino |
| 1897 | Propachyrucos crassus Ameghino p. 426 |
| 1898 | Propachyrucos robustus Roth p. 38 |
| 1901 | Propachyrucos aequilatus Ameghino p. 375 |
| 1908 | Hegetotheridae Sinclair |
| 1934 | Hegetotheriidae Simpson |
| 1949 | Hegetotheriidae Simpson and Minoprio |
| 1962 | Hegetotheriidae Simpson et al. |
| 1988 | Hegetotheriidae Carroll |
| 2006 | Hegetotheriidae Croft and Anaya pp. 393-396 |
| 2010 | Hegetotheriidae Kramarz et al. |
| 2010 | Hegetotheriidae Reguero and Prevosti |
| 2011 | Hegetotheriidae Billet |
| 2018 | Hegetotheriidae Vera and Ercoli |
| 2023 | Hegetotheriidae Seoane et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. †Hegetotheriidae Ameghino 1894
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Unr. †Hegetotheriinae Ameghino 1894
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G. †Hegetotherium Ameghino 1887
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†Hegetotherium cerdasensis Croft et al. 2016
†Hegetotherium mirabile Ameghino 1887
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Invalid names: Hegetotherium anceps Ameghino 1891 [synonym], Hegetotherium andinum Roth 1898 [synonym], Hegetotherium convexum Ameghino 1891 [synonym], Hegetotherium costatum Ameghino 1891 [synonym], Hegetotherium cuneatum Ameghino 1891 [synonym], Hegetotherium minum Ameghino 1894 [synonym], Hegetotherium strigatum Ameghino 1887 [synonym], Selatherium pachymorphum Ameghino 1894 [synonym], Selatherium remissum Ameghino 1894 [synonym]
†Hegetotherium novum Bordas 1939
Unr. †Hemihegetotheriomorpha Seoane et al. 2023
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G. †Hemihegetotherium Rovereto 1914
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†Hemihegetotherium achataleptum Rovereto 1914
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Invalid names: Hemihegetotherium lazai Zetti 1972 [objective synonym]
†Hemihegetotherium achathaleptum Rovereto 1914
†Hemihegetotherium affine Rovereto 1914
†Hemihegetotherium gracile Rovereto 1914
†Hemihegetotherium robustum Rovereto 1914
†Hemihegetotherium tantillum Vera 2019
†Hemihegetotherium torresi Cabrera and Kraglievich 1931
G. †Prohegetotherium Ameghino 1897
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†Prohegetotherium malalhuense Cerdeño and Reguero 2015
†Prohegetotherium schiaffinoi Kraglievich 1932
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Invalid names: Prohegetotherium carettei Minoprio 1947 [synonym]
†Prohegetotherium sculptum Ameghino 1897
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Invalid names: Prohegetotherium shumwayi Loomis 1914 [synonym]
Invalid names: Ethegotherium Simpson et al. 1962 [synonym]
Invalid names: Hemihegetotherium trilobus Croft and Anaya 2006 [nomen dubium]
Unr. †Pachyrukhinae Kraglievich 1934
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Unr. †Pachyrukhini Seoane et al. 2023
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G. †Pachyrukhos Ameghino 1885
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†Pachyrucos elongatus Moreno 1888
†Pachyrucos moyani Ameghino 1885
†Pachyrucos politus Ameghino 1902
†Pachyrucos teres Ameghino 1889
†Pachyrukhos moyani Ameghino 1885
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Invalid names: Pachyrucos absis Ameghino 1889 [synonym], Pachyrucos naevius Ameghino 1889 [synonym], Pachyrucos trivius Ameghino 1889 [synonym], Pachyrukhos teres Ameghino 1889 [synonym]
†Pachyrukhos ngenwinkul Solórzano et al. 2023
†Pachyrukhos politus Ameghino 1901
†Pachyrukhos tribius Ameghino 1889
Invalid names: Pedotherium Burmeister 1888 [synonym], Propachyrucos depressus Roth 1898 [nomen dubium], Propachyrucos medianus Roth 1898 [nomen dubium]
G. †Paedotherium Burmeister 1888
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†Paedotherium bonaerense Ameghino 1887
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Invalid names: Pachyrucos maximus Ameghino 1908 [synonym], Pachyrucos miramarensis Ameghino 1908 [synonym], Paedotherium imperforatum Kraglievich 1926 [synonym], Paedotherium insigne Burmeister 1888 [synonym], Paedotherium isolinense Castellanos 1936 [objective synonym]
†Paedotherium borrelloi Zetti 1972
†Paedotherium dolichognathus Zetti 1972
†Paedotherium ictus Ameghino 1889
†Paedotherium kakai Reguero et al. 2015
†Paedotherium typicum Ameghino 1887
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Invalid names: Paedotherium brusquitaense Ameghino 1908 [synonym], Paedotherium chapadmalense Ameghino 1908 [synonym], Paedotherium marplatense Ameghino 1908 [synonym]
Invalid names: Raulringueletia Zett 1972 [synonym]
G. †Tremacyllus Ameghino 1891
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†Tremacyllus impressus Ameghino 1888
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Invalid names: Tremacyllus chapalmalensis Ameghino 1908 [synonym], Tremacyllus diminutus Ameghino 1888 [synonym], Tremacyllus intermedius Rovereto 1914 [synonym], Tremacyllus novus Ameghino 1908 [synonym]
†Tremacyllus subdiminutus Rovereto 1914
G. †Prosotherium Ameghino 1897
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†Prosotherium garzoni Ameghino 1897
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Invalid names: Propachyrucos ameghinorum Simpson 1945 [synonym], Propachyrucos simpsoni Chaffee 1952 [synonym], Prosotherium quartum Ameghino 1901 [synonym], Prosotherium robustum Ameghino 1897 [synonym], Prosotherium triangulidens Ameghino 1897 [synonym]
G. †Paedsotherium Burmeister 1888
G. †Pseudohegetotherium Cabrera and Kraglievich 1931
G. †Selatherium Ameghino 1894
Invalid names: Propachyrucos aequilatus Ameghino 1901 [nomen dubium], Propachyrucos crassus Ameghino 1897 [nomen dubium], Propachyrucos robustus Roth 1898 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: s = species, o = order, subc = subclass, c = class, subp = subphylum | |||||
| References: Carroll 1988, Ji et al. 2002, Lillegraven 1979, Hendy et al. 2009, Solórzano et al. 2023, MacFadden et al. 1996, Vizcaíno et al. 2010 | |||||
Collections (181 total)
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Early/Lower Oligocene | Argentina (Chubut) | Hegetotheriidae indet. (186572) | |
| Rupelian | Argentina (Chubut) | Prohegetotherium sp. (176201) | |
| Deseadan | Argentina (Corrientes) | Prohegetotherium sp. (181625 181627) | |
| Deseadan | Bolivia (La Paz) | Prohegetotherium sp. (133615) Sallatherium altiplanense, Prohegetotherium schiaffinoi (13081) | |
| Deseadan | Argentina (Santa Cruz) | Hegetotherium sp. (50074) | |
| Deseadan | Uruguay | Hegetotherium sp., Pseudohegetotherium palmirense (176487) | |
| Deseadan | Argentina (Chubut) | Prohegetotherium sculptum (49852 176490 176647) Prohegetotherium sculptum, Propachyrucos ameghinorum, Medistylus dorsatus (146358) Prohegetotherium sculptum, Propachyrucos crassus, Prosotherium garzoni, Prosotherium triangulidens, Phanophilus dorsatus, Prohegetotherium shumwayi, Medistylus dorsatus, Hegetotheriopsis sulcatus (14211) Propachyrucos simpsoni, Prosotherium garzoni (49851) Propachyrucos sp., Prohegetotherium sculptum (176465) | |
| Deseadan | Argentina (Entre Rios) | Prohegetotherium sp. (181629) | |
| Deseadan | Argentina (Mendoza) | Prohegetotherium sp., Prosotherium sp., Hegetotheriinae indet., Prohegetotherium schiaffinoi, Prohegetotherium malalhuense, Hegetotheriopsis sulcatus, Hegetotheriinae indet., Propachyrucos smithwoodwardi (176466) | |
| Deseadan | Peru (Moquegua) | Hegetotheriidae indet. (59940) | |
| Deseadan | Uruguay (Canelones) | Propachyrucos schiaffinoi (176488) | |
| Deseadan | Bolivia (None) | Prohegetotherium sp. (13276) | |
| Deseadan | Bolivia | Sallatherium altiplanense, Prohegetotherium schiaffinoi (95688) | |
| Late/Upper Oligocene - Early/Lower Miocene | Bolivia | Prohegetotherium schiaffinoi (225573) | |
| Colhuehuapian | Argentina (Chubut) | Hegetotheriidae indet. (197191) Hegetotheriopsis sulcatus, Hegetotherium novum (145897) Pachyrucos indet. (176441) Pachyrukhos politus (141368) Pachyrukhos sp., Hegetotheriidae indet. (232404) Tegehotherium burmeisteri (232372) | |
| Colhuehuapian | Argentina (Neuquén) | Hegetotheriidae indet., Hegetotheriopsis sulcatus (89027) Hegetotherium sp. (89030) | |
| Colhuehuapian | Argentina (Mendoza) | Hegetotherium sp. (189519) | |
| Colhuehuapian | Argentina (Río Negro) | Hegetotherium sp., Hegetotheriopsis sulcatus (89093) | |
| Colhuehuapian - Burdigalian | Argentina (Chubut) | Pachyrukhos sp. (204204) | |
| Burdigalian | Chile (Aisén) | Hegetotherium sp. (71843) | |
| Burdigalian | Chile (Bío Bío) | Hegetotheriidae indet. (140103 140104) | |
| Burdigalian | Argentina (Chubut) | Pachyrukhos sp. (176205) | |
| Burdigalian - Langhian | Argentina (Santa Cruz) | Hegetotherium mirabile, Pachyrukhos moyani (197691) | |
| Burdigalian - Langhian | Bolivia | Hegetotheriidae indet. (91435) | |
| Santacrucian | Argentina (Santa Cruz) | Hegetotherium mirabile (27313 142533 176600 176602 176603 176641 206234 206242) Hegetotherium mirabile, Pachyrukhos moyani (176589 176590 210638) Hegetotherium sp. (176571 176578 210154) Hegetotherium sp., Pachyrukhos moyanoi, Hegetotherium mirabile (210153) Pachyrukhos moyani (87314 210637) Pachyrukhos sp. (176569 176570 176587) Pachyrukhos sp., Hegetotherium mirabile (176588) | |
| Santacrucian | Chile | Hegetotherium mirabile (103804) | |
| Santacrucian | Argentina (Mendoza) | Prohegetotherium carettei (176472) | |
| Santacrucian | Argentina (San Juan) | Hegetotheriinae indet., Pachyrukhinae indet. (197900) Hegetotherium mirabile (197899) | |
| Santacrucian - Friasian | Chile (Bío Bío) | Hegetotherium sp. (140107) | |
| Santacrucian - Laventan | Chile (Bío Bío) | Hegetotheriinae indet., Pachyrukhinae indet., Paedotherium minor (140105) | |
| Friasian | Chile (Aisén) | Pachyrukhos sp. (139845) | |
| Friasian | Chile | Pachyrukhos sp. (176666) | |
| Langhian | Argentina (Rio Negro) | Hemihegetotherium tantillum (198569) | |
| Langhian | Bolivia (Potosí) | Hegetotherium sp. (68402) | |
| Langhian - Serravallian | Argentina (Chubut) | Pachyrukhos moyani (215594 215595) | |
| Colloncuran | Argentina (Río Negro) | Hegetotherium sp., Pachyrukhos sp. (140691) | |
| Colloncuran | Bolivia (Oruro) | Hegetotheriidae indet. (68477) | |
| Serravallian - Tortonian | Argentina (La Pampa) | Paedotherium sp., Hemihegetotherium torresi (231228) | |
| Laventan | Colombia (Huila) | Hegetotheriidae indet. (133441) | |
| Laventan | Bolivia (Tarija) | Hegetotheriidae indet. (38071) | |
| Tortonian | Bolivia (La Paz) | H. sp. (90023) | |
| Tortonian | Argentina (Chubut) | Hemihegetotherium tantillum (201907) Tremacyllus sp. (201906) | |
| Tortonian | Chile (Parinacota) | Hegetotherium cerdasensis (144471) | |
| Late/Upper Miocene | Argentina (Salta) | Paedotherium kakai (192105) Paedotherium minor, Paedotherium kakai (197784) | |
| Late/Upper Miocene | Venezuela (Falcon) | Hemihegetotherium indet. (146449) | |
| Late/Upper Miocene | Bolivia (La Paz) | Pseudohegetotherium sp. (68796) | |
| Late/Upper Miocene - Early/Lower Pliocene | Argentina (Catamarca) | Tremacyllus incipiens (232594) | |
| Chasicoan | Argentina (La Pampa) | Paedotherium minor (206264) | |
| Chasicoan | Argentina (Buenos Aires) | Hegetotheriinae indet., Paedotherium minor, Hemihegetotherium torresi (79180) | |
| Chasicoan - Huayquerian | Argentina (La Pampa) | Paedotherium minor, Tremacyllus impressus (204494) | |
| Huayquerian | Argentina (La Pampa) | Hemihegetotherium achataleptum (204360 211562 211564) Hemihegetotherium sp. (13778) Hemihegetotherium sp., Paedotherium minor (230751) Paedotherium minor (192135 204361 206262) Paedotherium minor, Hemihegetotherium achataleptum (192166 206260 211563) Paedotherium minor, Hemihegetotherium torresi, Hemihegetotherium achataleptum (204363) Paedotherium minor, Tremacyllus impressus (151500) Paedotherium minor, Tremacyllus impressus, Hemihegetotherium achataleptum (152061 206031) Paedotherium minor, Tremacyllus impressus, Hemihegetotherium achataleptum, Hemihegetotherium lazai (87198) Paedotherium minor, Tremacyllus impressus, Hemihegetotherium torresi, Hemihegetotherium achataleptum (152063) Paedotherium sp. (151499) Paedotherium sp., Paedotherium minor, Tremacyllus impressus, Hemihegetotherium achataleptum (140925 151498) Tremacyllus sp., Paedotherium sp., Paedotherium minor, Tremacyllus impressus, Hemihegetotherium torresi (204598) | |
| Huayquerian | Argentina (Mendoza) | Paedotherium sp., Tremacyllus impressus, Hemihegetotherium achathaleptum (231327) Paedotherium typicum, Tremacyllus impressus (200444) | |
| Huayquerian | Argentina (San Juan) | Hemihegetotherium achataleptum (198513) | |
| Huayquerian | Bolivia (Chuquisaca) | Pseudohegetotherium sp. (68792) | |
| Huayquerian | Argentina (La Rioja) | Paedotherium minor (151506) Paedotherium minor, Tremacyllus sp., Hemihegetotherium torresi (55596) | |
| Huayquerian | Argentina (Buenos Aires) | Paedotherium bonaerense (61300 61304 61306 198868) Paedotherium minor (198270 198689 206032) Paedotherium minor, Tremacyllus impressus (61292 61299) Paedotherium sp. (198862) | |
| Huayquerian | Argentina (Catamarca) | Paedotherium sp. (157794) Tremacyllus incipiens (198547 206028) | |
| Messinian | Argentina (Catamarca) | Tremacyllus incipiens (198820) | |
| Messinian | Argentina (La Pampa) | Paedotherium minor (140932) Paedotherium sp., Paedotherium minor, Tremacyllus impressus (152067) Tremacyllus sp., Paedotherium minor, Tremacyllus impressus (141939) | |
| Messinian - Zanclean | Argentina (Mendoza) | Pachyrukhinae indet. (230923) Tremacyllus impressus (200118) | |
| Messinian - Zanclean | Argentina (Catamarca) | Tremacyllus incipiens (204710) | |
| Early/Lower Pliocene | Argentina (Buenos Aires) | Paedotherium sp. (191764 191767 191768 191769 191770 191773) | |
| Zanclean | Argentina (Buenos Aires) | Paedotherium bonaerense (191775) Paedotherium bonaerense, Tremacyllus impressus (191776) Paedotherium sp. (191774) | |
| Zanclean | Argentina (Catamarca) | Tremacyllus latifrons, Tremacyllus incipiens (198549) | |
| Montehermosan | Argentina (Mendoza) | Tremacyllus sp., Pachyrukhinae indet. (189520) | |
| Montehermosan | Argentina (Buenos Aires) | Paedotherium bonaerense (198684 198685 198686 198687) Paedotherium insigne (145757) Paedotherium sp. (152064 152086) Paedotherium sp., Paedotherium bonaerense (152062) Paedotherium sp., Paedotherium bonaerense, Tremacyllus impressus (152087) Paedotherium sp., Paedotherium typicum, Paedotherium bonaerense, Tremacyllus impressus (152081 152084 152085 152088) Paedotherium typicum, Paedotherium bonaerense, Tremacyllus impressus (198863) | |
| Montehermosan | Argentina (Catamarca) | Hemihegetotherium sp., Hemihegetotherium robustum (176757) | |
| Montehermosan - Chapadmalalan | Argentina (Catamarca) | Tremacyllus latifrons, Hemihegetotherium robustum (157795) | |
| Montehermosan - Chapadmalalan | Argentina (Córdoba) | Paedotherium bonaerense (197904) | |
| Chapadmalalan | Argentina (Córdoba) | Tremacyllus impressus (202212) | |
| Chapadmalalan | Argentina (Buenos Aires) | Paedotherium chapadmalense (198844) Paedotherium insigne (198699) Paedotherium typicum (202340) | |
| Chapadmalalan | Argentina (San Luis) | Paedotherium sp., Paedotherium typicum, Paedotherium insigne, Tremacyllus impressus (152073) | |
| Late/Upper Pliocene | Argentina (Córdoba) | Paedotherium sp., Paedotherium isolinense (211573) | |
| Late/Upper Pliocene | Argentina (Cordoba) | Tremacyllus impressus (191133) | |
| Late/Upper Pliocene | Argentina (Entre Rios) | Paedotherium typicum (79631 207682) | |
| Late/Upper Pliocene | Argentina (Buenos Aires) | Paedotherium sp. (191760 191763 191771 199160) | |
| Piacenzian | Argentina (Córdoba) | Tremacyllus impressus (225604) | |
| Early/Lower Pleistocene | Argentina (Buenos Aires) | Tremacyllus sp., Paedotherium sp. (71304) | |
| Lujanian | Argentina (Entre Ríos) | Paedotherium typicum (212840) | |
| Lujanian | Argentina (Buenos Aires) | Paedotherium sp., Paedotherium typicum, Paedotherium insigne, Tremacyllus impressus (202096) |