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Borophagus hilli

Mammalia - Carnivora - Canidae

Taxonomy
Osteoborus hilli was named by Johnston (1939). Its type specimen is Panhandle Plains Historical Museum No. 1558, a partial skull, and it is a 3D body fossil. Its type locality is Axtel, which is in a Hemphillian terrestrial horizon in Texas.

It was synonymized subjectively with Borophagus direptor by Kurten and Anderson (1980); it was recombined as Borophagus hilli by Wang et al. (1999), Wang et al. (2008), Tseng and Geisler (2016) and Albright et al. (2019).

Synonyms
Synonymy list
YearName and author
1939Osteoborus hilli Johnston p. 895 figs. 1 - 4
1944Osteoborus progressus Hibbard
1956Osteoborus crassapineatus Olsen
1979Borophagus crassapineatus Richey
1999Borophagus hilli Wang et al.
2008Borophagus hilli Wang et al.
2016Borophagus hilli Tseng and Geisler p. 1
2019Borophagus hilli Albright et al. p. 172

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Life
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
RankNameAuthor
Mammaliaformes
classMammalia
Theriamorpha(Rowe 1993)
Theriiformes()
Trechnotheria
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Ferae()
CarnivoramorphaWyss and Flynn 1993
CarnivoraformesFlynn et al.
orderCarnivora
familyCanidae
subfamilyBorophaginae
genusBorophagus
specieshilli()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Borophagus hilli Johnston 1939
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Invalid names: Osteoborus crassapineatus Olsen 1956 [synonym], Osteoborus progressus Hibbard 1944 [synonym]
Diagnosis
ReferenceDiagnosis
C. S. Johnston 1939Osteoborus hilli is considerably larger than the 0. cyonoides from the Middle Pliocene beds of Hemphill County, Texas. There is considerable variation within the latter spe- cies as shown in numerous upper and lower jaws in the collec- tions of the "\Vest Texas State College. In general massiveness the lower jaw of 0. hilli more closely approaches Borophagus diversidens, the type of which was recently redescribed by Van- derHoof (1936). In 0. hilli J3, C. P \ and M1 are powerfully developed. It is not possible to tell from the alveolus whether P 1 was double or single rooted. P 2 and P 3 are double rooted, however, and are so crowded that they are placed somewhat transversally in the jaw. These teeth are small and poorly developed with a very low central cusp. The parastyle on P4 is present, but vestigial. There is no anterior internal cusp on this tooth, though the root at this point is well developed. There is no trace of a cingulum on either P4 or M1 as is true in 0. cyonoides. In the following tables of dental measure- ments 0. hilli is shown in comparison to 0. cyonoides and B.diversidens. The figures for the two latter species are maxima taken from a number of specimens. The material representing B. diversidens was collected in the Cita Canyon Type Locality, referred to in a preliminary report by Johnston (1938).
Comparison of 0. hilli with 0. cyonoides and B. diversidens. Measurements are expressed in millimeters.