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Systemodon tapirinus

Mammalia - Perissodactyla

Taxonomy
Orohippus tapirinus was named by Cope (1875). Its type specimen is USNM 1064, a mandible (right mandible fragment with P3–M3), and it is a 3D body fossil.

It was recombined as Hyracotherium tapirinus by Cope (1877), Matthew (1899), Troxell (1922) and Radinsky (1963); it was recombined as Hyracotherium tapirinum by Wortman (1896) and MacFadden (1998); it was recombined as Eohippus tapirinus by Hay (1899), Hay (1902) and Peterson (1919); it was considered a nomen dubium by Granger (1908); it was recombined as Systemodon tapirinus by Cope (1882), Cope (1884), Froehlich (1999) and Froehlich (2002).

Synonyms
Synonymy list
YearName and author
1875Orohippus tapirinus Cope p. 20
1877Hyracotherium tapirinus Cope
1882Systemodon tapirinus Cope p. 183
1884Systemodon tapirinus Cope
1896Hyracotherium cristatum Wortman
1896Hyracotherium tapirinum Wortman
1896Hyracotherium cristatum Wortman p. 96 fig. 6
1899Eohippus tapirinus Hay
1899Hyracotherium tapirinus Matthew
1902Eohippus cristatus Hay p. 609
1902Eohippus tapirinus Hay p. 610
1908Eohippus cristatus Granger
1919Eohippus tapirinus Peterson p. 103
1922Hyracotherium tapirinus Troxell
1930Eohippus cristatus Hay
1940Eohippus cristatus Stirton
1963Hyracotherium tapirinus Radinsky
1991Hyracotherium cristatum Gingerich
1998Hyracotherium cristatum MacFadden p. 542
1998Hyracotherium tapirinum MacFadden p. 542
1999Systemodon tapirinus Froehlich
2002Systemodon tapirinus Froehlich p. 183–184

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
RankNameAuthor
Mammaliaformes
classMammalia
Theriamorpha(Rowe 1993)
Theriiformes()
Trechnotheria
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
Laurasiatheria
Scrotifera
Euungulata
Panperissodactyla
superorderPerissodactylamorpha
orderPerissodactyla()
Tapiromorpha(Haeckel 1873)
genusSystemodon
speciestapirinus()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Systemodon tapirinus Cope 1875
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Invalid names: Hyracotherium cristatum Wortman 1896 [synonym]
Diagnosis
ReferenceDiagnosis
D. J. Froehlich 2002This taxon has no identified unambiguous autapomorphies but it can be diagnosed by the characters it shares with Cymbalophus. The synapomorphies include the following characters: upper molar parastyle small (Character 38, 2 to 1), M3 smaller than M2, M3 larger than M2 (Character 67, 2 to 1), P4 entoconid small and variable (Character 75, 2 to 1), and M3 hypoconulid connection forming a triple junction with the prehypoconulid crista (Character 105, 0 to 1). The last of these characters is only found in these two taxa and strongly supports their monophyly. Cymbalophus can be distinguished from Systemodon only on size and geography.