PBDB Taxon

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Eoblattodea

Insecta

Taxonomy

Eoblattodea was named by Laurentiaux (1959).

It was reranked as the unranked clade Eoblattodea by Li (2019).

It was assigned to Blattodea by Laurentiaux-Vieira and Laurentiaux (1980); and to Holopandictyoptera by Li (2019).

Subtaxa

Blattinariae, Ensiferoblattidae, Euthyrrhaphoidea, Latindioidea, Mylacroidea, Necymylacroidea, Phoberoblattoidea, Proceroblatta, Spiloblattinidae

Synonyms

Mylacridina was named by Anisyutkin (2007).

It was synonymized subjectively with Eoblattodea by Li (2019).

It was assigned to Blattodea by Anisyutkin (2007).
Palaeoblattariae was named by Scudder (1879) [possibly 1878 or earlier]. It is not extant.

It was reranked as the suborder Palaeoblattina by Durden (1969); it was reranked as the unranked clade Palaeoblattariae by Kluge (2010); it was synonymized subjectively with Eoblattodea by Li (2019).

It was assigned to Orthoptera by Scudder (1886); to Orthopteroidea by Scudder (1893); to Dictyoptera by Durden (1969); and to Holopandictyoptera by Kluge (2010).

Synonymy list

Year	Name and author
1879	Palaeoblattariae Scudder
1886	Palaeoblattariae Scudder p. 210
1893	Palaeoblattariae Scudder
1959	Eoblattodea Laurentiaux
1969	Palaeoblattina Durden p. 1171
1980	Eoblattodea Laurentiaux-Vieira and Laurentiaux p. 416
2007	Mylacridina Anisyutkin
2010	Palaeoblattariae Kluge
2019	Eoblattodea Li

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
superclass	Hexapoda	Latreille 1825
class	Insecta

Rank	Name	Author
—	Dicondylia
—	Paranotalia
subclass	Pterygota	()
—	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
—	Polyneoptera	Martynov 1938
—	Holopandictyoptera
—	Eoblattodea	()
—	Eoblattodea	()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. †Eoblattodea Laurentiaux 1959

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Fm. †Ensiferoblattidae Li and Huang 2023

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G. †Ensiferoblatta Li and Huang 2023

†Ensiferoblatta oecanthoides Li and Huang 2023

G. †Proceroblatta Li and Huang 2023

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†Proceroblatta colossea Li and Huang 2023

Fm. †Spiloblattinidae Handlirsch 1906

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G. †Ametroblatta Handlirsch 1906

†Ametroblatta strigosa Scudder 1890

G. †Arrhythmoblatta Handlirsch 1906

†Arrhythmoblatta detecta Scudder 1895

†Arrhythmoblatta scudderiana Handlirsch 1906

G. †Atactoblatta Handlirsch 1906

†Atactoblatta anomala Handlirsch 1906

G. †Dicladoblatta Handlirsch 1906

†Dicladoblatta defossa Scudder 1895

†Dicladoblatta marginata Scudder 1885

G. †Doryblatta Handlirsch 1906

†Doryblatta longipennis Handlirsch 1906

G. †Kinneyblatta Schneider et al. 2021

†Kinneyblatta huberi Schneider et al. 2021

G. †Olethroblatta Handlirsch 1906

†Olethroblatta americana Handlirsch 1906

†Olethroblatta lineolata Cockerell 1927

†Olethroblatta minuta Guthorl 1930

†Olethroblatta tomiensis Becker-Migdisova 1961

Subfm. †Permoblattininae Vishnyakova 1993

G. †Permoblattina Tillyard 1937

†Permoblattina curta Sellards 1908

Invalid names: Etoblattina pecta Sellards 1908 [synonym]

†Permoblattina permiana Sellards 1908

G. †Pycnoblattina Sellards 1908

G. †Sysciolata Guthorl 1933

†Sysciolata vorkutensis Vishnyakova 1993

G. †Spiloblattina Scudder 1885

†Spiloblattina allegheniensis Carpenter 1934

†Spiloblattina homigtalensis Schneider and Werneburg 1993

†Spiloblattina lawrenceana Handlirsch 1906

Invalid names: Spiloblattina curvata Sellards 1908 [synonym], Spiloblattina lineata Sellards 1908 [synonym], Sysciophlebia sellardsi Handlirsch 1906 [synonym]

†Spiloblattina odernheimensis Schneider 1982

†Spiloblattina pygmaea Meunier 1904

Invalid names: Sysciophlebia lawrenceana Handlirsch 1906 [synonym]

†Spiloblattina sperbersbachensis Schneider and Werneburg 1993

†Spiloblattina steinbachi Guthörl 1934

†Spiloblattina variegata Scudder 1890

Invalid names: Etoblattina exsensa Scudder 1895 [synonym], Etoblattina fasciata Scudder 1890 [synonym], Etoblattina funesta Scudder 1895 [synonym], Etoblattina gracilenta Scudder 1895 [synonym], Etoblattina hastata Scudder 1895 [synonym], Etoblattina hustoni Scudder 1890 [synonym], Etoblattina maledicta Scudder 1895 [synonym], Etoblattina marginata Scudder 1890 [synonym], Etoblattina ramosa Scudder 1895 [synonym], Etoblattina stipata Scudder 1890 [synonym], Etoblattina tenuis Scudder 1890 [synonym], Etoblattina willsiana Scudder 1895 [synonym], Gerablattina apicalis Scudder 1895 [synonym], Poroblattina longinqua Scudder 1895 [synonym]

†Spiloblattina weissigensis Geinitz 1873

Invalid names: Spiloblattina gardineri Scudder 1885 [synonym], Spiloblattina guttata Scudder 1885 [synonym], Spiloblattina perforata Handlirsch 1906 [synonym], Spiloblattina triassica Scudder 1885 [synonym], Sysciophlebia fenestrata Handlirsch 1906 [synonym]

†Spiloblattinidae gardinerana Handlirsch 1906

†Spiloblattinidae zinkeniana Schlechtendal 1906

G. †Syscioblatta Handlirsch 1906

†Syscioblatta allegheniensis Carpenter 1934

†Syscioblatta anomala Handlirsch 1906

†Syscioblatta dohrni Scudder 1879

Invalid names: Sysciophlebia steinbachi Guthörl 1934 [synonym]

†Syscioblatta euryptera Vishnyakova 1993

†Syscioblatta kaierkanensis Vishnyakova 1993

†Syscioblatta minor Handlirsch 1906

†Syscioblatta misera Handlirsch 1906

†Syscioblatta obscura Handlirsch 1906

†Syscioblatta pennsylvanica Carpenter 1934

†Syscioblatta slusiensis Schneider and Werneburg 1993

†Syscioblatta steubenvilleana Handlirsch 1906

†Syscioblatta teberdensis Vishnyakova 1993

G. †Sysciophlebia Handlirsch 1906

†Sysciophlebia acutipennis Handlirsch 1906

†Sysciophlebia adumbrata Handlirsch 1906

†Sysciophlebia affinis Handlirsch 1906

†Sysciophlebia alligans Schneider 1982

†Sysciophlebia angusta Langiaux and Parriat 1975

†Sysciophlebia arcuata Sellards 1904

†Sysciophlebia balteata Scudder 1879

Invalid names: Etoblattina occulta Scudder 1895 [synonym], Gerablattina cassvici Scudder 1895 [synonym], Gerablattina diversinervis Scudder 1895 [synonym], Sysciophlebia diversipennis Scudder 1895 [synonym], Sysciophlebia frankei Schlechtendal 1906 [synonym]

†Sysciophlebia benedicta Scudder 1895

†Sysciophlebia euglyptica Germar 1851

Invalid names: Dicladoblatta limbata Schlechtendal 1906 [synonym], Gerablattina producta Scudder 1879 [synonym], Gerablattina weissiana Goldenberg 1869 [synonym], Sysciophlebia agilis Schlechtendal 1906 [synonym], Sysciophlebia deperdita Schlechtendal 1906 [synonym], Sysciophlebia elegantissima Schlechtendal 1906 [synonym], Sysciophlebia elongata Schlechtendal 1906 [synonym], Sysciophlebia huysseni Schlechtendal 1906 [synonym], Sysciophlebia ignota Schlechtendal 1906 [synonym], Sysciophlebia laspeyresiana Schlechtendal 1906 [synonym], Sysciophlebia martiusana Schlechtendal 1906 [synonym], Sysciophlebia schlechtendali Handlirsch 1920 [synonym]

†Sysciophlebia grata Schneider 1982

†Sysciophlebia hercynica Scharf 1924

†Sysciophlebia hybrida Handlirsch 1906

†Sysciophlebia ilfeldensis Schlechtendal 1906

†Sysciophlebia intermedia Goldenberg 1877

†Sysciophlebia invisa Scudder 1895

†Sysciophlebia kovacsae Riek 1976

†Sysciophlebia nana Handlirsch 1906

†Sysciophlebia obtusa Handlirsch 1906

†Sysciophlebia oligoneura Handlirsch 1906

†Sysciophlebia patiens Scudder 1895

†Sysciophlebia picta Handlirsch 1906

†Sysciophlebia praepilata Meunier 1921

Invalid names: Sysciophlebia expansa Bolton 1925 [synonym]

†Sysciophlebia recidiva Scudder 1895

†Sysciophlebia rotundata Handlirsch 1906

†Sysciophlebia rubida Schneider 1982

†Sysciophlebia schucherti Handlirsch 1906

†Sysciophlebia scudderi Handlirsch 1906

†Sysciophlebia stulta Schlechtendal 1906

†Sysciophlebia subtilis Schlechtendal 1906

Invalid names: Berlichia wettinensis Schlechtendal 1906 [synonym], Sysciophlebia angustipennis Schlechtendal 1906 [synonym]

†Sysciophlebia variegata Scudder 1890

†Sysciophlebia whitei Handlirsch 1906

Subfm. †Vivablattinae Vršanský and Mostovsky 2000

G. †Vivablatta Vršanský and Mostovsky 2000

†Vivablatta orenburgensis Vishnyakova 1993

†Vivablatta pectinacea Vishnyakova 1993

Invalid names: Kargalia Vishnyakova 1993 [replaced]

Invalid names: Kargaliinae Vishnyakova 1993 [replaced]

G. †Zavjaloblatta Becker-Migdisova 1961

†Zavjaloblatta rotundata Becker-Migdisova 1961

Invalid names: Anthracoblattina triassica Scudder 1890 [nomen dubium], Anthracoblattina wagneri Kliver 1886 [nomen dubium], Etoblattina aperta Scudder 1895 [nomen dubium], Spiloblattina laxa Sellards 1908 [nomen dubium]

Invalid names: Mylacridina Anisyutkin 2007 [synonym], Palaeoblattariae Scudder 1879 [synonym]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Composition:

chitinc

Environment:

terrestrialc

Locomotion:

actively mobilep

Created:

2017-04-17 09:56:18

Modified:

2017-04-17 09:56:18

Source: c = class, p = phylum

References: Bush and Bambach 2015, Kiessling 2004

Age range

Maximum range based only on fossils: base of the Westphalian C to the top of the Early/Lower Cenomanian or 314.60000 to 93.50000 Ma
Minimum age of oldest fossil (stem group age): 309.8 Ma

Collections (110 total)

Time interval	Ma	Country or state	Original ID and collection number
Moscovian	315.2 - 307.0	USA (Maryland)	Olethroblatta lineolata, Ambloblatta bassleri (124496)
Westphalian C	314.6 - 309.8	Canada (Nova Scotia)	Discoblatta coalburnensis (216824) Grypoblattina phaseolus (216810) Intermylacris intermedia (216808 216811) Intermylacris sp. (216818) Linemylacris mclellanensis (216807)
Westphalian C - Westphalian D	314.6 - 307.5	USA (Rhode Island)	Etoblattina scholfieldi (112704)
Westphalian C - Westphalian D	314.6 - 307.5	Canada (Nova Scotia)	Archimylacris acadica (155668)
Westphalian C - Westphalian D	314.6 - 307.5	USA (Pennsylvania)	Olethroblatta americana (114456) Pteridomylacris paradoxa (125764)
Westphalian D	309.8 - 307.5	USA (Pennsylvania)	Phthinomylacris pauper (140162) Schizoblatta pennsylvanica (124504)
Westphalian D	309.8 - 307.5	Canada (Nova Scotia)	Xenoblattidae indet. (216842)
Westphalian D	309.8 - 307.5	France	Grypoblattina chandesrisi (155432) Grypoblattina phaseolus, Grypoblattina robineti (155431)
Stephanian	307.5 - 300.4	Germany (Rheinland-Pfalz)	Blattina weissiana (122603)
Stephanian	307.5 - 300.4	Germany (Saarland)	Blattina intermedia, Blattina wemmetsweileriensis (122600) Fulgorina klieveri (138356)
Stephanian	307.5 - 300.4	Germany	Etoblattina pygmaea (125762) Sysciophlebia sp. (138004) Sysciophlebia steinbachi (155778)
Stephanian	307.5 - 300.4	USA (Ohio)	Etoblattina maledicta, Etoblattina benedicta, Schizoblatta alutacea, Etoblattina fossa, Sysciophlebia whitei, Sysciophlebia scudderi, Sysciophlebia hybrida, Sysciophlebia affinis, Etoblattina ramosa, Etoblattina variegata, Sysciophlebia schucherti, Sysciophlebia picta, Sysciophlebia adumbrata, Etoblattina funesta, Sysciophlebia rotundata, Sysciophlebia nana, Sysciophlebia obtusa, Sysciophlebia acutipennis, Etoblattina hastata, Etoblattina fasciata, Etoblattina marginata, Gerablattina apicalis, Etoblattina tenuis, Etoblattina willsiana, Etoblattina exsensa, Syscioblatta obscura, Syscioblatta anomala, Syscioblatta minor, Etoblattina hustoni, Etoblattina gracilenta, Syscioblatta steubenvilleana, Syscioblatta misera, Etoblattina strigosa, Poroblattina longinqua, Atactoblatta anomala, Doryblatta longipennis, Etoblattina stipata (123812) Gerablattina balteata (122605)
Kasimovian	307.0 - 303.7	USA (Kansas)	Blattinariae indet., Gerablattina arcuata, Spiloblattina lineata, Spiloblattina curvata, Spiloblattina laxa, Sysciophlebia lawrenceana, Sysciophlebia sellardsii, Spiloblattina maledicta (113003) Spiloblattina maledicta (147045)
Kasimovian - Gzhelian	307.0 - 298.9	Russian Federation	Syscioblatta teberdensis (126062)
Gzhelian	303.7 - 298.9	Russian Federation	Zavjaloblatta rotundata (124583)
Gzhelian	303.7 - 298.9	France	Etoblattina praepilata (121979) Sysciophlebia expansa (124916)
Gzhelian	303.7 - 298.9	Spain	Sysciophlebia sp. (227128)
Stephanian B	301.5 - 301.0	Morocco	Spiloblattina pygmaea (155288 155290 155291) Sysciophlebia grata (155293)
Stephanian B	301.5 - 301.0	France	Sysciophlebia angusta (172760)
Stephanian B	301.5 - 301.0	Czech Republic	Sysciophlebia grata (156567)
Stephanian C	301.0 - 300.4	Germany	Berlichia wettinensis (172340)
Stephanian C	301.0 - 300.4	Czech Republic (Hradec Králové)	Spiloblattina lawrenceana, Sysciophlebia rubida (156568)
Stephanian C	301.0 - 300.4	Germany (Saxony-Anhalt)	Blattina euglyptica (153702) Spiloblattinidae indet., Blattina euglyptica, Etoblattina dohrnii, Gerablattina producta, Berlichia wettinensis, Sysciophlebia elongata, Sysciophlebia angustipennis, Sysciophlebia deperdita, Sysciophlebia agilis, Sysciophlebia elegantissima, Sysciophlebia stulta, Sysciophlebia ignota, Dicladoblatta subtilis, Dicladoblatta limbata, Spiloblattinidae zinkeniana (122578) Syscioblatta dohrni, Sysciophlebia subtilis (125736) Sysciophlebia huysseni, Sysciophlebia agilis, Sysciophlebia oligoneura, Sysciophlebia laspeyresiana, Syscioblatta dohrni, Anthracoblattina wagneri, Sysciophlebia sp. 3, Sysciophlebia sp. 2 (122579) Sysciophlebia sp., Sysciophlebia martiusana (125761)
Stephanian C	301.0 - 300.4	USA (Pennsylvania)	Syscioblatta allegheniensis (144528) Syscioblatta pennsylvanica (155383)
Asselian	298.9 - 295.5	Germany (Rhineland-Palatinate)	Sysciophlebia sp. B (156570 156571)
Asselian	298.9 - 295.5	USA (Colorado)	Spiloblattina gardineri, Spiloblattina triassica, Spiloblattina guttata, Spiloblattina marginata, Sysciophlebia fenestrata, Anthracoblattina triassica, Spiloblattinidae gardinerana, Spiloblattina perforata (106345)
Asselian	298.9 - 295.5	Czech Republic	Spiloblattina weissigensis (114141) Sysciophlebia alligans (133680 133681 156569)
Asselian	298.9 - 295.5	Germany	Spiloblattina odernheimensis (156564 156565) Sysciophlebia balteata (192084)
Asselian	298.9 - 295.5	USA (West Virginia)	Gerablattina balteata, Etoblattina sagittaria, Etoblattina mediana, Etoblattina ovata, Etoblattina debilis, Etoblattina patiens, Etoblattina mucronata, Etoblattina detecta, Etoblattina invisa, Etoblattina occulta, Etoblattina defossa, Etoblattina recidiva, Gerablattina diversinervis, Gerablattina cassvici, Gerablattina permanenta, Gerablattina lata, Gerablattina rotundata, Anthracoblattina virginiensis, Poroblattina fossa, Poroblattina complexinervis, Sysciophlebia balteata, Arrhythmoblatta scudderiana, Etoblattina fasciata (122604)
Asselian	298.9 - 295.5	Germany (Thüringer Wald)	Spiloblattina weissigensis (128478) Sysciophlebia balteata (167765 167768)
Asselian	298.9 - 295.5	Germany (Thuringia)	Sysciophlebia hercynica (155777) Sysciophlebia ilfeldensis (128321)
Asselian	298.9 - 295.5	Austria	Sysciophlebia sp 1, Sysciophlebia sp 2, Sysciophlebia sp 3 (146278)
Asselian - Sakmarian	298.9 - 290.1	Russian Federation (Krasnoyarsk)	Syscioblatta euryptera (126061) Syscioblatta kaierkanensis (126063)
Asselian - Sakmarian	298.9 - 290.1	Germany (Rhineland-Palatinate)	Olethroblatta minuta (122606)
Asselian - Sakmarian	298.9 - 290.1	Germany	Sysciophlebia frankei, Sysciophlebia balteata (127934)
Sakmarian	293.52 - 290.1	Germany (Thüringer Wald)	Sysciophlebia balteata (167766) Sysciophlebia balteata, Spiloblattina weissigensis (160734) Sysciophlebia balteata, Syscioblatta slusiensis (167767)
Sakmarian	293.52 - 290.1	Germany	Blattina weissigensis, Blattina mahri (122580) Spiloblattina odernheimensis (156566)
Artinskian	290.1 - 279.3	Russian Federation	Olethroblatta tomiensis (124594)
Artinskian	290.1 - 279.3	USA (Kansas)	Etoblattina curta, Etoblattina pecta, Etoblattina permiana (21979) Spiloblattinidae indet., Permoblattina curta, Pycnoblattina sp., Permoblattina permiana (111219)
Artinskian	290.1 - 279.3	USA (Oklahoma)	Permoblattina curta (104826)
Kungurian	279.3 - 272.3	Russian Federation (Komi Republic)	Sysciolata vorkutensis (126064)
Kungurian	279.3 - 272.3	Russian Federation	Spiloblattinidae indet. (102504)
Kungurian	279.3 - 272.3	South Africa (Gauteng)	Sysciophlebia kovacsae (122863)
Roadian	272.3 - 268.8	Russian Federation (Kirov)	Kitjakia longifurcata (105058)
Capitanian	264.28 - 259.51	Russian Federation (Orenburg)	Kargalia orenburgensis, Kargalia pectinacea (104057)
Triassic	251.902 - 201.4	China (Xinjiang)	Triassoblatta tudupinensis (146283)
Bithynian	247.2 - 242.0	Germany (Bavaria)	Triassoblatta sp. (115969)
Anisian - Norian	247.2 - 208.5	Australia (Tasmania)	Triassoblatta reducta (151871) Triassoblatta tasmanica (150065)
Ladinian	242.0 - 237.0	Brazil (Rio Grande do Sul)	Triassoblatta cargnini (117269)
Longobardian	242.0 - 235.0	Argentina (Mendoza)	Triassoblatta sp. (115104)
Longobardian	242.0 - 235.0	China (Shaanxi)	Triassoblatta phyllopteris (123778)
Carnian	237.0 - 228.0	South Africa (Kwazulu-Natal)	Triassoblatta natalensis (23002)
Carnian	237.0 - 228.0	Japan	Triassoblatta sp. A, Triassoblatta sp. B, Triassoblatta bella, Triassoblatta tenuicubiti, Triassoblatta rotundipenna (155677) Triassoblatta sp. C, Triassoblatta okafujii (155678)
Carnian	237.0 - 228.0	China (Xinjiang)	Triassoblatta tudupinensis, Triassoblatta hyptis (156325)
Carnian	237.0 - 228.0	South Africa	Rhipidoblattopsis natalensis (151912)
Carnian	237.0 - 228.0	Argentina (La Rioja)	Triassoblatta argentina (123045) Triassoblatta sp. (122987)
Carnian	237.0 - 228.0	South Africa (Eastern Cape)	Triassoblatta robusta (23142 23317) Triassoblatta simplex (115196)
Late/Upper Triassic	237.0 - 201.4	South Africa (Kwazulu-Natal)	Triassoblatta sp. (23003)
Norian	228.0 - 208.5	Australia (Queensland)	Triassoblatta jonesi, Triassoblatta denmeadi (156626) Triassoblatta ovata, Triassoblatta dehiscens (122248) Triassoblatta triquestra (122247) Triassoblatta triquestra, Triassoblatta ovata, Triassoblatta grandis, Triassoblatta obtusa, Triassoblatta intramedia, Triassoblatta jonesi (122246) Triassoblatta typica, Triassoblatta insignita, Triassoblatta intermedia (111558)
Norian - Rhaetian	228.0 - 201.3	China (Nei Mongol)	Triassoblatta sp. D (155680)
Rhaetian	208.5 - 201.3	China (Guangdong)	Triassoblatta opistodictyna, Triassoblatta tegmina (156321)
Rhaetian - Hettangian	208.5 - 199.3	Russian Federation (Sverdlovsk)	Triassoblatta uralica (117987)
Callovian - Oxfordian	166.1 - 157.3	China (Liaoning)	Triassoblatta fusiformis (113475)
Callovian - Oxfordian	166.1 - 157.3	China (Hebei)	Triassoblatta damiaoliangensis (113480)
Early/Lower Cenomanian	99.6 - 93.5	Myanmar	Ensiferoblatta oecanthoides, Proceroblatta colossea (161726)