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Heterocaryonidae

Trilobita - Harpetida - Heterocaryonidae

Taxonomy
Heterocaryonidae was named by Hupé (1953). Its type is Heterocaryon.

It was synonymized subjectively with Entomaspididae by Ludvigsen (1982).

It was assigned to Harpetida by Beech and Lamsdell (2021).

Subtaxa

Synonymy list
YearName and author
1953Heterocaryonidae Hupé
2021Heterocaryonidae Beech and Lamsdell p. 2220

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
RankNameAuthor
phylumArthropodaLatreille 1829
subphylumArtiopoda(Hou and Bergstrom)
classTrilobitaWalch 1771
orderHarpetida(Whittington 1959)
familyHeterocaryonidaeHupé 1953
familyHeterocaryonidaeHupé 1953

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Fm. †Heterocaryonidae Hupé 1953
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G. †Bowmania Walcott 1925
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Bowmania americana Walcott 1884
G. †Conococheaguea Rasetti 1959
G. †Heterocaryon Raymond 1937
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Heterocaryon tuberculatum Rasetti 1944
Diagnosis
ReferenceDiagnosis
J. D. Beech and J. C. Lamsdell 2021Angle of cephalic curvature greater than 90°. Yoked librigenae, but lacking true bilamellar fringe. Facial sutures directed outward and forward. Small eye lobes, diverging posteriorly, with eye ridges anterolaterally directed. High cephalon convexity and equilateral glabellar lateral margins. Highly convex genae. Lacking alae. Narrow, rounded genal prolongations (i.e. spines). Pygidium of four to eight segments.
Measurements
No measurements are available
Composition: low Mg calcitec
Composition 2: chitinc
Ontogeny: molting, addition of partsc
Environment: marinec
Locomotion: fast-movingc
Life habit: low-level epifaunalc
Diet: carnivorec
Created: 2004-02-29 09:07:31
Modified: 2016-06-02 12:44:28
Source: c = class
Reference: Aberhan et al. 2004

Age range: base of the Trempealeauan to the top of the Sunwaptan or 492.50000 to 487.50000 Ma

Collections (29 total)


Time interval Ma Country or state Original ID and collection number
Merioneth497.0 - 485.4USA (Vermont) Bowmania americana (6871) Heterocaryon platystigma (8976)
Sunwaptan493.5 - 487.5USA (New York) Bowmania pennsylvanica (257 85185)
Sunwaptan493.5 - 487.5Canada (Alberta) Heterocaryon vargum (54096 81298 81301) Heterocaryon vargum, Conococheaguea ludvigseni (54094 81297)
Sunwaptan493.5 - 487.5Canada (Newfoundland and Labrador) Heterocaryon platystigma (81270 81285) Heterocaryon platystigma, Heterocaryon tuberculatum (81269)
Trempealeauan492.5 - 485.4Canada (Quebec) Heterocaryon tuberculatum (224136 224138 224139)
Trempealeauan492.5 - 485.4Canada (Newfoundland and Labrador) Heterocaryon sp. (105858)
Trempealeauan492.5 - 485.4USA (Texas) Bowmania americana (233745) Bowmania pennsylvanica (219651) Bowmania pennsylvanica, Bowmania americana (219652) Bowmania sagitta (233747) Bowmania sp. (216018)
Trempealeauan492.5 - 485.4Canada (Yukon) Bowmania americana (55285 55310 55311 55313)
Trempealeauan492.5 - 485.4USA (Wyoming) Bowmania pennsylvania (9775)
Trempealeauan492.5 - 485.4USA (Nevada) Bowmania americana (233789) Heterocaryon indet. (50831)
Trempealeauan492.5 - 485.4USA (Utah) Bowmania sp. (259)