PBDB Taxon

Basic info	Taxonomic history	Classification	Included Taxa
Morphology	Ecology and taphonomy	External Literature Search	Age range and collections

Mandibulata

Taxonomy

Mandibulata was named as a subphylum.

It was reranked as the unranked clade Mandibulata by Ortega-Hernández (2016).

It was assigned to Arthropoda by Shoshani et al. (1989), Nielsen (2012); and to Deuteropoda by Ortega-Hernández (2016).

Subtaxa

Acheronauta, Aquilonifer, Balhuticaris, Cambropachycope, Hymenocarina, Martinssonia, Musacaris, Oelandocaris, Pancrustacea

Synonymy list

Year	Name and author
1989	Mandibulata Shoshani et al. p. 435
2012	Mandibulata Nielsen p. 244 figs. Fig. 44.2
2016	Mandibulata Ortega-Hernández

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Protostomia
—	Ecdysozoa	Aguinaldo et al. 1997

Rank	Name	Author
—	Panarthropoda
phylum	Arthropoda	Latreille 1829
—	Deuteropoda
—	Mandibulata
—	Mandibulata

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Unr. †Mandibulata

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G. †Acheronauta Pulsipher et al. 2022

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†Acheronauta stimulapis Pulsipher et al. 2022

G. †Aquilonifer Briggs et al. 2016

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†Aquilonifer spinosus Briggs et al. 2016

G. †Balhuticaris Izquierdo-López and Caron 2022

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†Balhuticaris voltae Izquierdo-López and Caron 2022

G. †Cambropachycope Waloszek and Mueller 1990

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†Cambropachycope clarksoni Waloszek and Mueller 1990

Or. †Hymenocarina Clarke 1900

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Fm. †Perspicarididae Briggs 1978

G. †Perspicaris Briggs 1977

†Perspicaris dilatus Robison and Richards 1981

Fm. †Pseudoarctolepidae Brooks and Caster 1956

G. †Pseudoarctolepis

†Pseudoarctolepis sharpi Brooks and Caster 1956

G. †Martinssonia Mueller and Waloszek 1986

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†Martinssonia elongata Mueller and Waloszek 1986

G. †Musacaris Haug et al. 2010

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†Musacaris gerdgeyeri Haug et al. 2010

G. †Oelandocaris Müller 1983

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†Oelandocaris oelandica Müller 1983

Unr. Pancrustacea Zrzavý 1997

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Unr. Altocrustacea Regier et al. 2010

Supercl. Allotriocarida Oakley et al. 2013

Unr. Athalassocarida Lozano-Fernandez et al. 2019

Cl. Branchiopoda Latreille 1817 [branchiopod]

Unr. Labiocarida Schwentner et al. 2017

Cl. Cephalocarida Sanders 1955

Or. †Brachypoda Birshteyn 1960

G. †Rehbachiella Mueller 1983

†Rehbachiella kinnekullensis Mueller 1983

G. †Bredocaris Mueller 1983

†Bredocaris admirabilis Mueller 1983

Unr. †Hymenostraca Rolfe 1969

G. †Branchiocaris Briggs 1976

†Branchiocaris malongensis Luo et al. 2008

†Branchiocaris pretiosa Resser 1929

†Branchiocaris xundianensis Luo et al. 2008

†Branchiocaris yiliangensis Luo et al. 2008

†Branchiocaris yunnanensis Hou 1987

G. †Ercaicunia Luo et al. 1999

†Ercaicunia multinodosa Luo et al. 1999

G. †Hymenocaris Salter 1853

†Hymenocaris dubia Resser and Howell 1938

Supercl. Multicrustacea Regier et al. 2010

Infracl. †Halicyna Gall and Grauvogel 1967

Or. †Cyclida Glaessner 1928

Cl. Hexanauplia Oakley et al. 2013

Subcl. Copepoda Milne-Edwards,Milne-Edwards 1840 [copepod]

Cl. Malacostraca Latreille 1802

G. †Amphipeltis Salter 1863

Subcl. Eumalacostraca Grobben 1892

G. †Naranda Muenster 1842

Subcl. Phyllocarida Packard 1879

Invalid names: Belotelsonidea Schram 1981 [empty], Waterstonellidea [empty]

G. †Nereocaris Legg et al. 2012

†Nereocaris exilis Legg et al. 2012

Supercl. Oligostraca Zrzavý et al. 1997

Cl. Ichthyostraca Zrzavý et al. 1997

Subcl. Branchiura Thorell 1864

Subcl. Pentastomida Diesing 1836

Invalid names: Mystacocarida Pennak and Zinn 1943 [empty]

Cl. Ostracoda Latreille 1802 [ostracod]

G. †Acanthomeridion Hou et al. 1989

G. †Acrossula

G. †Anabarochilina

G. †Anfedatia

G. †Aristaluta

G. †Aurilia

G. †Australopsis

G. †Bennviaspis

Or. †Beyrichiocopa Pokorný 1954

G. †Biaurina

G. †Borodiella

G. †Boucekites

G. †Brachycytheropteron

G. †Brasacypris Krömmelbein 1965

G. †Bullaluta

Fm. †Buregiidae Polenova 1953

G. †Bursulella Jones 1961

G. †Cambraechmina

G. †Carnarvonia

G. †Cedocamia

G. †Craspedopyxion

G. †Crescenticythere

G. Cyclocypris Brady and Norman 1889

G. Cyprinotus Brady 1886

G. †Cytherina

G. Cytheropsis Sars 1866

G. †Damaoella

G. †Dhrumaella Manivit and Vaslet 1987

G. †Dorispina

G. †Duibianella

G. †Eucytheridea Bronstein 1930

G. †Fengtaiella

G. †Flemingia

G. †Flemingopsis

G. †Galeopsis Hinz-Schallreuter 1993

G. †Gelidicella

G. †Gerdalia Belousova 1961

G. †Gerubiella

G. †Gladioscutum

G. Glossicythereis Benson 1974

G. †Hameaschmidtella

G. †Hemiaechminoides

G. Heterocypris Claus 1893

G. †Huanghuaella

G. †Hupehella

G. †Ilmenoindivisia

G. †Infracytheropteron

G. †Ivaria

G. †Jixinglingella

G. †Koilocythere Deltel 1963

G. †Kuiperiana Bassiouni 1962

G. †Kuresaaria

G. †Liagnshanella

Or. †Limbatulocopida Abushik 1990

G. †Lublinella

G. †Ludvigsenites

G. †Maghrebeis

G. †Majiashanella

G. †Mannosocmia

G. †Marginotimorites

G. †Microcosmia

G. †Moorea

Subcl. Myodocopa Sars 1866

Superor. Myodocopomorpha Kozur 1972

G. †Navicularina

G. Neoatlanticythere Benson 1974

G. †Netzkaina

G. †Oepikaluta

Subor. †Paleocopa Henningsmoen 1953

G. †Paleocythere

G. †Paleodarwinula Molostovskaya 1990

G. †Parashergoldopsis

G. †Paratsunyiella

G. †Phasoia

G. †Plesidielymella

Subcl. Podocopa Sars 1865

Superor. Podocopamorpha Kozur 1972

Superor. Podocopomorpha Sars 1865

G. †Ponthocythere Dubowsky 1939

G. †Prasuchonella Molostovskaya 1980

G. †Pseudonodellina

G. †Pseudoperissocytheridea

G. Puriana Coryell and Fields 1953

Invalid names: Favella Coryell and Fields 1937 [synonym]

G. †Reginacypris

G. †Renicytherura Gruendel 1981

G. †Reticulocambria

G. †Reversoparaparchites

G. †Rockallia Whatley et al. 1978

G. †Schallreuterina

G. †Septadella Smith and Stubblefield 1933

G. †Shergoldopsis

G. †Sigillium

G. †Spytihnevites

G. †Suchonellina Spizharsky 1937

G. †Supratoarcina

G. †Svealuta

G. †Tatariella

G. †Tethysia Donze 1975

G. †Tetrasulcata

G. †Torusilina

G. †Trapezisylthere

G. †Utsatia

G. †Uvonhactia

G. †Venzavella

G. †Vitella

Superfm. †Volganelloidea Mandelstam 1956

G. †Yaxianella

G. †Yulinella

G. †Zerqacythere Basha 1980

Invalid names: Beichuania Xie 1989 [nomen nudum], Beyrichiida Pokorny 1953 [empty], Cuselina Ammon 1910 [nomen nudum], Darwinulacea Brady and Norman 1889 [empty], Golcondella [nomen dubium]

G. †Tokummia Aria and Caron 2017

†Tokummia katalepsis Aria and Caron 2017

Invalid names: Hymenocarididae Haeckel 1896 [empty]

G. †Skara Mueller 1983

†Skara anulata Mueller 1983

G. †Wujicaris Zhang et al. 2010

†Wujicaris muelleri Zhang et al. 2010

G. †Yicaris Zhang et al. 2007

†Yicaris dianensis Zhang et al. 2007

Invalid names: Crustacea Brünnich 1772 [invalid subgroup]

Diagnosis

No diagnoses are available

Measurements

No measurements are available

Locomotion:

actively mobilep

Created:

2004-02-29 09:38:49

Modified:

2004-02-29 11:38:49

Source: p = phylum

Reference: Kiessling 2004

Age range

Maximum range based only on fossils: base of the Ediacaran to the top of the Holocene or 635.00000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 538.8 Ma

Collections (17177 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Ediacaran	635.0 - 538.8	Mexico	Platella sp. (10662)
Meishucunian	530.7 - 521.0	China (Shaanxi)	Ostracoda indet. (6851)
Atdabanian	521.0 - 516.0	China (Yunnan)	Branchiocaris yunnanensis (90163 90166 90494) Canadaspis laevigata (96403 96405 96416) Chuandianella ovata (86847 90126 90129 90149 90150 90151 90152 90161 90162) Chuandianella ovata, Branchiocaris yunnanensis (86845 86846 90138 90164) Chuandianella ovata, Canadaspis laevigata (90165 96402) Chuandianella ovata, Canadaspis laevigata, Branchiocaris yunnanensis (86851) Chuandianella ovata, Ercaicunia multinodosa (90137) Waptia sp. (86931) Waptia sp., Branchiocaris sp. (86929) Waptia sp., Branchiocaris sp., Acanthomeridion sp. (86930) Waptia sp., Canadaspis sp., Acanthomeridion sp. (86933) Waptia sp., Chuandianella ovata, Branchiocaris yunnanensis, Ercaicunia multinodosa (68083)
Atdabanian	521.0 - 516.0	Australia (South Australia)	Phasoia spicata (155233)
Atdabanian	521.0 - 516.0	Germany (Brandenburg)	Diplostraca indet. (24190)
Nangaoian	521.0 - 514.5	China (Yunnan)	Waptia ovata (132328) Waptia ovata, Branchiocaris yunnanensis (132329 194140 194162 194163) Waptia ovata, Branchiocaris yunnanensis, Ercaicunia multinodosa (194141 194150 194153) Waptia ovata, Canadaspis laevigata, Branchiocaris yunnanensis (132327 194160 194161) Waptia ovata, Canadaspis laevigata, Branchiocaris yunnanensis, Ercaicunia multinodosa (132330 194143 194155)
Nangaoian	521.0 - 514.5	China (Hubai)	Branchiocaris sp. (219687)
Nangaoian	521.0 - 514.5	China (Guizhou)	Perspicaris sp. (128423)
Branchian	516.2 - 506.5	United Kingdom (England)	Hesslandona n. sp. B (79939)
Botomian	516.0 - 512.1	China (Yunnan)	Branchiocaris sp. (90353)
Botomian - Lenian	516.0 - 508.2	Russian Federation (Transbaikalia)	Ostracoda indet. (10007)
Botomian - Lenian	516.0 - 508.2	Russian Federation	Duibianella sp. (10018)
Dyeran	515.3 - 509.7	USA (Pennsylvania)	Crustacea indet. (226194) Roddyia typa, Crustacea indet., Hymenocaris dubia (226192) Roddyia typa, Hymenocaris sp. (192)
Dyeran	515.3 - 509.7	USA (California)	Ostracoda indet. (219693)
Dyeran	515.3 - 509.7	USA (Nevada)	Canadaspis sp., Perspicaris sp. (10560)
Dyeran	515.3 - 509.7	Canada (Newfoundland and Labrador)	Leperditicopa sp. (9900)
Dyeran	515.3 - 509.7	Canada (Northwest Territories)	Branchiopoda indet. (6850)
Duyunian	514.5 - 506.5	China (Guizhou)	Waptia sp., Canadaspis sp. (237148)
Duyunian	514.5 - 506.5	China (Hubei)	Cycloconchoides venustus (201164)
Delamaran	509.7 - 505.3	Canada (British Columbia)	Canadaspis perfecta (84420) Canadaspis perfecta, Perspicaris recondita (84437) Waptia fieldensis (84430 84431) Waptia fieldensis, Canadaspis perfecta (84416 84417 84434 84436) Waptia fieldensis, Canadaspis perfecta, Branchiocaris pretiosa (84412) Waptia fieldensis, Canadaspis perfecta, Perspicaris dictynna (84421 84422 84424 84425 84428) Waptia fieldensis, Canadaspis perfecta, Perspicaris dictynna, Perspicaris recondita (84426) Waptia fieldensis, Canadaspis perfecta, Perspicaris recondita (84414 84415 84418 84419 84429 84432 84433 84435) Waptia fieldensis, Canadaspis perfecta, Perspicaris recondita, Perspicaris dictynna (84423 84427) Waptia sp., Canadaspis perfecta, Branchiocaris sp., Nereocaris exilis, Perspicaris sp. (121959)