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Archaeocyatha (archaeocyath sponge)

Archaeocyatha

Taxonomy

Archaeocyatha was named by Vologdin (1937).

It was reranked as the class Archaeocyatha by Rowland (2001) and Debrenne et al. (2015).

It was assigned to Porifera by Rowland (2001) and Debrenne et al. (2015).

Subtaxa

Adaecyathus, Ajacicyathida, Archaeocyathida, Butovia, Capsulocyathida, Cornutocyathus, Irregulares, Kazachstanicyathida, Monocyathida, Putapacyathida, Regulares, Rhizacyathus

Synonyms

Butovia is a genus. It is not extant.

It was considered a nomen dubium by Debrenne et al. (2015).

It was assigned to Cribricyathea by Sepkoski (2002).
Rhizacyathus was named by Bedford (1939).

It was considered a nomen nudum by Debrenne et al. (2015).

It was assigned to Irregulares by Sokolov et al. (1974).
Adaecyathus was named by Zhuravlev et al. (1983).

It was considered a nomen nudum by Debrenne et al. (2015).

It was assigned to Archaeocyatha by Zhuravlev et al. (1983).
Regulares was named by Vologdin (1937). It was considered polyphyletic by Debrenne et al. (2015).

It was considered an invalid subgroup of Archaeocyatha by Debrenne et al. (2015).

It was assigned to Porifera by Sepkoski (2002); and to Archaeocyatha by Bambach et al. (2007).
Irregulares was named by Vologdin (1937).

It was considered an invalid subgroup of Archaeocyatha by Debrenne et al. (2015).

It was assigned to Porifera by Sepkoski (2002); and to Archaeocyatha by Bambach et al. (2007).
Cornutocyathus was named by Zhuravleva et al. (1997).

It was considered a nomen nudum by Debrenne et al. (2015).

It was assigned to Irregulares by Zhuravleva et al. (1997).

Synonymy list

Year	Name and author
1937	Irregulares Vologdin
1937	Regulares Vologdin
1939	Rhizacyathus Bedford
1974	Rhizacyathus Sokolov et al.
1983	Adaecyathus Zhuravlev et al.
1997	Cornutocyathus Zhuravleva et al. p. 166
2001	Archaeocyatha Rowland p. 1065
2002	Butovia Sepkoski
2002	Irregulares Sepkoski
2002	Regulares Sepkoski
2007	Irregulares Bambach et al. p. S6
2007	Regulares Bambach et al. p. S6
2015	Archaeocyatha Debrenne et al. p. 1

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Rank	Name	Author
kingdom	Animalia	()
phylum	Porifera	Grant 1836

Rank	Name	Author
class	Archaeocyatha	(Vologdin 1937)
class	Archaeocyatha	(Vologdin 1937)

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Cl. †Archaeocyatha Vologdin 1937 [archaeocyath sponge]

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Or. †Ajacicyathida Bedford and Bedford 1939

Subor. †Ajacicyathina Bedford and Bedford 1939

Superfm. †Annulocyathoidea Krasnopeeva 1953

Fm. †Annulocyathidae Krasnopeeva 1953

G. †Annulocyathella Vologdin 1962

G. †Annulocyathus Vologdin 1937

G. †Annulofungia Krasnopeeva 1955

Invalid names: Kiyafungia Zhuravleva et al. 1997 [synonym]

G. †Hemithalamocyathus

Fm. †Gagarinicyathidae Debrenne et al. 1989

G. †Gagarinicyathus

Fm. †Jakutocarinidae Debrenne et al. 1989

G. †Jakutocarinus Zhuravleva 1960

G. †Kosticyathus Debrenne and Zhuravlev 2000

G. †Kruseicyathus Zhuravleva et al. 1997

G. †Rossocyathella

G. †Russocyathus

Fm. †Tumulifungiidae Rozanov 1989

G. †Sclerocyathus

G. †Subtumulocyathellus Osadchaya et al. 1979

Invalid names: Arturocyathus Debrenne et al. 1990 [synonym]

G. †Tologoicyathus Voronin 1988

G. †Tumulifungia Zhuraleva 1968

Superfm. †Bronchocyathoidea Bedford and Bedford 1936

Fm. †Ajacicyathidae Bedford and Bedford 1939

G. †Ajacicyathus Bedford and Bedford 1939

G. †Davidicyathus Debrenne et al. 1990

G. †Dentatocyathus Okuneva 1972

G. †Iljinicyathus Zhuravleva 1972

G. †Kisasacyathus Konyushkov 1972

Invalid names: Prethmophyllum [synonym]

G. †Nochoroicyathus Zhuravleva 1951

Invalid names: Ajacicyathellus Debrenne 1958 [synonym], Aldanocyathus Voronin 1979 [synonym], Howellicyathus Vologdin 1962 [synonym]

G. †Orbicyathellus Osadchaya 1973

G. †Orbicyathus Vologdin 1937

G. †Robustocyathellus Konyushkov 1972

G. †Rotundocyathus Vologdin 1960

G. †Sibirecyathus Vologdin 1937

G. †Stapicyathus Debrenne 1964

Invalid names: Nochoroicyathellus Osadchaya et al. 1979 [synonym]

G. †Turgorocyathus Skorlotova 2013

Fm. †Bipallicyathidae Debrenne et al. 1989

G. †Bipallicyathus

G. †Heckericyathus Zhuravleva 1960

Fm. †Bronchocyathidae Bedford and Bedford 1936

G. †Compositocyathus

G. †Conannulofungia

G. †Cyathocricus

G. †Cyclocyathella

G. †Denaecyathus

G. †Gordonicyathus Zhuravleva 1959

Invalid names: Sichotecyathus Okuneva and Repina 1973 [synonym]

G. †Gordonifungia

G. †Morenicyathus

Invalid names: Denaecyathellus Osadchaya et al. 1979 [synonym], Kellericyathus [synonym]

G. †Pseudotennericyathellus Osadchaya et al. 1979

G. †Sagacyathus Kruse 1982

G. †Stillicidocyathus

G. †Svetlanocyathus

G. †Taylorcyathus

Invalid names: Tersicyathus [synonym], Thalamocyathellus Osadchaya et al. 1979 [synonym]

G. †Taylorfungia Perejon 1989

G. †Thalamocyathus

Invalid names: Bronchocyathus Bedford and Bedford 1936 [synonym], Thalamopectinus Debrenne 1973 [synonym]

G. †Trininaecyathus

Fm. †Densocyathidae Vologdin 1937

G. †Angustocyathus Skorlotova 2013

G. †Cadniacyathus

G. †Dailycyathus

Invalid names: Joanaecyathus Gravestock 1984 [synonym]

G. †Deceptioncyathus Gravestock 1984

G. †Densocyathus

G. †Khirgisocyathus Voronin 1988

G. †Leptosocyathellus Osadchaya et al. 1979

G. †Leptosocyathus

Invalid names: Halysicyathus [synonym]

G. †Natalijaecyathus Kotel'nikov 1995

G. †Rectannulus Debrenne 1977

G. †Tennericyathus

Invalid names: Raropectinus Debrenne and Rozanov 1983 [synonym]

Fm. †Ethmocyathidae Debrenne 1969

G. †Afiacyathus Voronin 1962

G. †Baikalocyathus

Invalid names: Baikalopectinus Gravestock 1984 [synonym]

G. †Carpicyathus

G. †Degeletticyathus

G. †Ethmocyathus

G. †Frinalicyathus

Invalid names: Pseudodegeletticyathellus Osadchaya et al. 1979 [synonym]

G. †Gnaltacyathus

G. †Hyptocyathus

G. †Inessocyathellus

G. †Inessocyathus

Invalid names: Rowanpectinus Gravestock 1984 [synonym], Voroninicyathus [synonym]

G. †Mackenziecyathus

Invalid names: Ussuricyathus Okuneva and Repina 1973 [synonym]

G. †Rasetticyathus

G. †Terraecyathus Zhuravleva 1974

G. †Ussuricyathellus Voronin 1988

G. †Zonacyathellus Zhuravleva 1974

Fm. †Sajanocyathidae Vologdin 1956

G. †Chakassicyathus Zhuravleva and Elkina 1974

G. †Emucyathus Kruse and Moreno-Eiris 2013

G. †Formosocyathus

G. †Irinaecyathus

Invalid names: Kandatocyathus Osadchaya et al. 1979 [synonym]

G. †Kiwicyathus Debrenne and Kruse 1986

G. †Palmericyathus Handfield 1971

G. †Sajanocyathus

G. †Siderocyathus Debrenne et al. 1990

G. †Zonacyathus

Invalid names: Diplocyathellus [synonym]

Superfm. †Erbocyathoidea Vologdin and Zhuravleva 1956

Fm. †Erbocyathidae Volgdin and Zhuravleva 1956

G. †Ladaecyathus

G. †Milaecyathus Debrenne and Zhuravlev 2000

G. †Pluralicyathus Okulitch 1950

Fm. †Peregrinicyathidae Zhuravleva 1967

G. †Peregrinicyathus

Fm. †Tegerocyathidae Krasnopeeva 1972

G. †Krasnopeevaecyathus Rozanov 1964

G. †Tegerocyathus

Invalid names: Alexandricyathus Osadchaya et al. 1979 [synonym]

Fm. †Vologdinocyathidae Yaroshevich 1957

G. †Gumbycyathus

G. †Inacyathella

G. †Kordecyathus

G. †Sanarkophyllum

G. †Syringocyathus

G. †Vologdinocyathus

Invalid names: Larecyathus Osadchaya et al. 1979 [synonym], Tegerocyathella Konyushkov 1967 [synonym]

Superfm. †Ethmophylloidea Okulitch 1937

Fm. †Carinacyathidae Krasnopeeva 1953

G. †Carinacyathus

Invalid names: Porocyathus Zhuravleva 1960 [synonym]

G. †Hupecyathellus

G. †Porocyathellus Bojarinov and Osadchaja 1979

G. †Vologdinocyathellus

Fm. †Ethmophyllidae Okulitch 1937

G. †Angaricyathus Zhuravleva 1965

G. †Aulocricus

G. †Cordilleracyathus Handfield 1971

G. †Dupliporocyathus

G. †Ethmophyllum Meek 1868

G. †Kolbicyathus

G. †Parethmophyllum

G. †Squamosocyathus

G. †Stephenicyathus Voronova et al. 1987

Fm. †Fallocyathidae Rozanov 1969

G. †Fallocyathus

G. †Sekwicyathus Handfield 1971

G. †Yukonocyathus Handfield 1971

Fm. †Gloriosocyathidae Rozanov 1969

G. †Gandinocyathus Debrenne and Debrenne 2007

G. †Gloriosocyathus

G. †Nalivkinicyathus

Fm. †Kijacyathidae Zhuravleva 1964

G. †Aporosocyathus Kruse 1978

G. †Fansycyathus

G. †Flexanulus

G. †Kijacyathus

G. †Protocyathus Ford 1878

G. †Qinlingocyathus Yang and Yuan 2012

G. †Yudjaicyathus

Superfm. †Lenocyathoidea Zhuravleva 1956

Fm. †Japhanicyathidae Rozanov 1989

G. †Japhanicyathus

Fm. †Lenocyathidae Zhuravleva 1956

G. †Lenocyathus

Fm. †Torosocyathidae Debrenne et al. 2002

G. †Torosocyathella Kotel'nikov 1995

G. †Torosocyathus Kashina 1972

Invalid names: Rewardocyathus Rozanov 1973 [synonym]

Superfm. †Pretiosocyathoidea Rozanov 1969

Fm. †Pretiosocyathidae Rozanov 1969

G. †Jangudacyathus

G. †Loculicyathopsis Zhuravleva et al. 1997

G. †Pretiosocyathus Rozanov 1966

Invalid names: Pretiosocyathellus Osadchaya et al. 1979 [synonym]

Fm. †Robertocyathidae Rozanov 1969

G. †Mattajacyathus

G. †Robertocyathus Rozanov 1969

G. †Urcyathella

Superfm. †Sigmocyathoidea Krasnopeeva 1953

Fm. †Sigmocyathidae Krasnopeeva 1953

G. †Didymocyathus

G. †Sigmocyathus

Fm. †Wrighticyathidae Kruse 1978

G. †Wrighticyathus

Superfm. †Tercyathoidea Vologdin 1939

Fm. †Botomocyathidae Zhuravleva 1955

G. †Botomocyathus

G. †Clathrithalamus Debrenne and Kruse 1986

Fm. †Olgaecyathidae Borodina 1974

G. †Olgaecyathus

Fm. †Piamaecyathellidae Rozanov 1974

G. †Piamaecyathellus

Fm. †Tercyathidae Vologdin 1939

G. †Clathricyathellus

G. †Clathricyathus

G. †Tercyathellus

G. †Tercyathus

Invalid names: Piamaecyathus [synonym]

Superfm. †Tumulocyathoidea Krasnopeeva 1953

Fm. †Geocyathidae Debrenne 1964

G. †Geocyathus

Invalid names: Jakutocyathus [synonym]

Fm. †Konjuschkovicyathidae Debrenne and Zhuravlev 2000

G. †Konjuschkovicyathus Debrenne and Zhuravlev 2000

Fm. †Sanarkocyathidae Hill 1972

G. †Neokolbicyathus Konayaeva 1997

G. †Ringifungia Korshunov 1969

G. †Sanarkocyathus

Fm. †Tumulocyathidae Krasnopeeva 1953

G. †Isiticyathus

G. †Kotuyicyathellus Osadchaya et al. 1979

Invalid names: Borocyathus [synonym]

G. †Plicocyathus Vologdin 1960

Invalid names: Torosocyathellus Osadchaya et al. 1979 [synonym], Tumulocyathellus Zhuravleva 1960 [synonym]

G. †Tumulocyathus

Invalid names: Kotuyicyathus Zhuravleva 1960 [synonym], Tumulocyathoides Zhuravleva et al. 1997 [synonym]

G. †Botocyathus

Subor. †Dokidocyathina Vologdin 1957

Superfm. †Dokidocyathoidea Bedford and Bedford 1936

Fm. †Cordobicyathidae Perejón 1975

G. †Cordobicyathus Perejón 1975

Fm. †Dokidocyathellidae Debrenne 1964

G. †Dokidocyathella Zhuravleva 1960

G. †Incurvocyathus Rozanov and Missarzhevsky 1966

Fm. †Dokidocyathidae Bedford and Bedford 1936

G. †Dokidocyathus Taylor 1910

Superfm. †Kaltatocyathoidea Kerner et al. 2011

Fm. †Kaltatocyathidae Rovanov 1964

G. †Kaltatocyathus Zhuravleva et al. 1964

Invalid names: Aroonacyathus Gravestock 1984 [synonym]

Superfm. †Kidrjasocyathoidea Rozanov 1964

Fm. †Kidrjasocyathidae Rozanov 1964

G. †Kidrjasocyathus Rozanov 1960

Superfm. †Kymbecyathoidea Debrenne et al. 1990

Fm. †Kymbecyathidae Debrenne et al. 1990

G. †Kymbecyathus Debrenne and Kruse 1986

G. †Orbiasterocyathus Zhuravleva 1964

G. †Urcyathus

Superfm. †Papillocyathoidea Rozanov 1989

Fm. †Papillocyathidae Rozanov 1989

G. †Papillocyathus Zhuravleva et al. 1964

Superfm. †Soanicyathoidea Rozanov 1964

Fm. †Soanicyathidae Rozanov 1964

G. †Batschykicyathus Zhuravlev 1983

G. †Subtilocyathus Vologdin 1960

Fm. †Zhuravlevaecyathidae Rozanov 1989

G. †Zhuravlevaecyathus Rozanov 1964

Subor. †Erismacoscinina Debrenne et al. 1989

Superfm. †Coscinoptyctoidea Debrenne et al. 1989

Fm. †Coscinoptyctidae Debrenne et al. 1989

G. †Coscinoptycta

Fm. †Geyericoscinidae Debrenne and Zhuravlev 2000

G. †Geyericoscinus Debrenne and Zhuravlev 2000

Fm. †Jebileticoscinidae Debrenne et al. 1989

G. †Irhoudicoscinus Debrenne 1977

G. †Jebileticoscinus

Invalid names: Pachycoscinus [synonym]

Superfm. †Ethmocoscinoidea Zhuravleva 1957

Fm. †Ethmocoscinidae Zhuravleva 1957

G. †Ethmocoscinus

Fm. †Tumulocoscinidae Zhuravleva 1960

G. †Asterotumulus

G. †Orbicoscinus

G. †Retetumulus

G. †Tumulocoscinus

Superfm. †Kasyricyathoidea Zhuravleva 1961

Fm. †Agyrekocyathidae Konyushkov 1967

G. †Agyrekocyathus Konyushkov 1967

Invalid names: Mennericyathus [synonym]

Fm. †Kasyricyathidae Zhuravleva 1961

G. †Kasyricyathus

Fm. †Membranacyathidae Debrenne et al. 2002

G. †Membranacyathus

Fm. †Xestecyathidae Debrenne et al. 1989

G. †Xestecyathus

Superfm. †Mootwingeecyathoidea Kruse 1982

Fm. †Mootwingeecyathidae Kruse 1982

G. †Mootwingeecyathus

Superfm. †Polycoscinoidea Debrenne 1964

Fm. †Anaptyctocyathidae Debrenne 1970

G. †Anaptyctocyathus Debrenne 1969

Fm. †Polycoscinidae Debrenne 1964

G. †Lunulacyathus

G. †Polycoscinus

Invalid names: Erugatocyathus [synonym]

Fm. †Veronicacyathidae Debrenne et al. 2002

G. †Veronicacyathus

Invalid names: Bractocyathus [synonym]

Fm. †Zonacoscinidae Debrenne 1971

G. †Orienticyathus

G. †Zonacoscinus

Superfm. †Porocoscinoidea Debrenne 1964

Fm. †Porocoscinidae Debrenne 1964

G. †Geniculicyathus Debrenne 1960

G. †Porocoscinus

G. †Tubicoscinus

Fm. †Rozanovicyathidae Korshunov 1969

G. †Rozanovicyathus

Fm. †Tatijanaecyathidae Korshunov 1976

G. †Muchattocyathus

Invalid names: Tatijanaecyathus Korshunov 1976 [synonym]

G. †Schumnyicyathus

Superfm. †Salairocyathoidea Zhuravleva 1956

Fm. †Asterocyathidae Vologdin 1956

G. †Antoniocoscinus Zhuravlev 1988

G. †Asterocyathus Vologdin 1940

G. †Erismacoscinus Debrenne 1958

Invalid names: Churanocyathus [synonym]

G. †Ichnusocyathus

G. †Retecoscinus

G. †Rozanovicoscinus

Fm. †Crassicoscinidae Debrenne et al. 1988

G. †Crassicoscinus

G. †Crucicyathus Gravestock 1984

G. †Dentatocoscinus

Fm. †Rudanulidae Debrenne et al. 1989

G. †Pilodicoscinus Debrenne and Jiang 1989

G. †Rudanulus

G. †Yhecyathus Belyaeva and Kesin 1995

Fm. †Salairocyathidae Zhuravleva 1956

G. †Kotuyicoscinus

G. †Polystillicidocyathus Debrenne 1959

G. †Salairocyathus

Superfm. †Sigmocoscinoidea Bedford and Bedford 1939

Fm. †Sigmocoscinidae Bedford and Bedford 1939

G. †Sigmocoscinus

G. †Statanulocyathus

Fm. †Sylviacoscinidae Debrenne et al. 1989

G. †Sylviacoscinus

G. †Kidjasocyathus Rozanov 1960

G. †Procyathellus

G. †Ptretiosocyathus Rozanov 1966

G. †Stephanicyathus Voronova et al. 1987

Invalid names: Archaeocyathellus Ford 1873 [nomen dubium], Nevadacyathus Okulitch 1943 [nomen dubium], Somphocyathus Taylor 1910 [nomen dubium], Tabulacyathus [nomen dubium]

Or. †Archaeocyathida Okulitch 1935

Subor. †Anthomorphina Okulitch 1935

Superfm. †Anthomorphoidea Okulitch 1935

Fm. †Anthomorphidae Okulitch 1935

G. †Anthomorpha Bornemann 1884

G. †Tollicyathus

Fm. †Shiveligocyathidae Fonin 1983

G. †Shiveligocyathus

Invalid names: Voznesenskicyathus [synonym]

Subor. †Archaeocyathina Okulitch 1935

Superfm. †Archaeocyathoidea Hinde 1889

Fm. †Archaeocyathidae Taylor 1910

G. †Araneocyathus Vologdin 1937

G. †Archaeocyathus Billings 1861

Invalid names: Batenevia Krasnopeeva 1961 [synonym], Claruscyathus Vologdin 1932 [synonym], Flindersicyathus Bedford and Bedford 1937 [synonym], Retecyathus Vologdin 1932 [synonym], Sanxiacyathus Yuan and Zhang 1977 [synonym], Syringsella Krasnopeeva 1961 [synonym], Vadimocyathus Osadchaya et al. 1979 [synonym]

G. †Arrythmocricus

G. †Pycnoidocyathus Taylor 1910

Invalid names: Archaeofungia Taylor 1910 [synonym]

G. †Sigmofungia

Invalid names: Palmericyathellus Debrenne 1970 [synonym]

Fm. †Archaeopharetridae Bedford and Bedford 1936

G. †Archaeopharetra

Invalid names: Dictyocoscinus Bedford and Bedford 1936 [synonym], Hawkercyathus Gravestock 1984 [synonym]

G. †Dictyosycon

G. †Markocyathus

G. †Protopharetra Bornemann 1884

G. †Spirocyathella Vologdin 1939

Invalid names: Aruntacyathus Kruse and West 1980 [synonym]

Fm. †Archaeosyconidae Zhuravleva 1954

G. †Archaeosycon

Invalid names: Pustulacyathellus Debrenne and Gangloff 1987 [synonym]

Superfm. †Beltanacyathoidea Debrenne 1974

Fm. †Beltanacyathidae Debrenne 1974

G. †Beltanacyathus

Invalid names: Bayleicyathus Gravestock 1984 [synonym], Fridaycyathus Gravestock 1984 [synonym]

Fm. †Maiandrocyathidae Debrenne 1974

G. †Ataxiocyathus

G. †Maiandrocyathus

Superfm. †Dictyocyathoidea Taylor 1910

Fm. †Claruscoscinidae Debrenne and Zhuravlev 1992

G. †Claruscoscinus Handfield 1971

Invalid names: Arisacyathus Osadchaya et al. 1979 [synonym]

G. †Fenestrocyathus Handfield 1971

G. †Landercyathus Debrenne et al. 1990

G. †Stevocyathus

Fm. †Dictyocyathidae Taylor 1910

G. †Bottonaecyathus

G. †Cellicyathus

G. †Chouberticyathus

G. †Dictyocyathus

Invalid names: Prismocyathus [synonym]

G. †Graphoscyphia Debrenne 1974

Invalid names: Graphoscyphus [objective synonym]

G. †Molybdocyathus Debrenne et al. 1990

G. †Paracoscinus

G. †Retilamina Debrenne and James 1981

Fm. †Pycnoidocoscinidae Debrenne 1974

G. †Pycnoidocoscinus Bedford and Bedford 1936

G. †Pycnoidoscinus Bedford and Bedford 1936

Superfm. †Metacyathoidea Bedford and Bedford 1934

Fm. †Copleicyathidae Bedford and Bedford 1937

G. †Agastrocyathus Debrenne 1964

G. †Copleicyathus

G. †Gabrielsocyathus

G. †Metacyathellus Debrenne and Zhuravlev 1990

G. †Spinosocyathus Zhuravleva 1960

G. †Spirillicyathella Bedford and Bedford 1937

G. †Spirillicyathus Bedford and Bedford 1937

G. †Tabulacyathellus

Fm. †Jugalicyathidae Gravestock 1984

G. †Alaskacoscinus

G. †Jugalicyathus Gravestock 1984

Fm. †Metacyathidae Bedford and Bedford 1934

G. †Changicyathus

G. †Metaldetes

Invalid names: Metacyathus Bedford and Bedford 1934 [synonym], Metafungia Bedford 1934 [synonym]

Invalid names: Sigmofungiidae Bedford and Bedford 1936 [empty]

Superfm. †Naimarkcyathoidea Wrona and Zhuravlev 1996

Fm. †Naimarkcyathidae Wrona and Zhuravlev 1996

G. †Naimarkcyathus Wrona and Zhuravlev 1996

Superfm. †Tabellaecyathoidea Fonin 1963

Fm. †Tabellaecyathidae Fonin 1963

G. †Taeniaecyathellus

Invalid names: Tabellaecyathus Fonin 1963 [synonym]

Superfm. †Warriootacyathoidea Debrenne and Zhuravlev 1992

Fm. †Warriootacyathidae Debrenne and Zhuravlev 1992

G. †Warriootacyathus Gravestock 1984

G. †Cambrocyathells

G. †Dendrocyathus Okulitch and Roots 1947

Subor. †Dictyofavina Debrenne 1991

Superfm. †Gatagacyathoidea Debrenne and Zhuravlev 1992

Fm. †Gatagacyathidae Debrenne and Zhuravlev 1992

G. †Gatagacyathus

Superfm. †Keriocyathoidea Debrenne and Gangloff 1992

Fm. †Keriocyathidae Debrenne and Gangloff 1992

G. †Keriocyathus Debrenne et al. 1990

Superfm. †Usloncyathoidea Fonin 1966

Fm. †Usloncyathidae Fonin 1966

G. †Kechikacyathus

G. †Usloncyathus

Invalid names: Dictyofavus Gravestock 1984 [synonym], Nostrocyathus [synonym]

G. †Zunyicyathus

Subor. †Loculicyathina Zhuravleva 1954

Superfm. †Chankacyathoidea Yakovlev 1959

Fm. †Chankacyathidae Yakovlev 1959

G. †Chankacyathus

Fm. †Tchojacyathidae Debrenne and Zhuravlev 1992

G. †Tchojacyathus

Superfm. †Loculicyathoidea Zhuravleva 1954

Fm. †Eremitacyathidae Debrenne 1992

G. †Eremitacyathus

Fm. †Loculicyathidae Zhuravleva 1954

G. †Ardrossacyathus

Invalid names: Egiinocyathus Fonin 1983 [synonym]

G. †Cambrocyathellus

Invalid names: Robustocyathus [synonym]

G. †Loculicyathus

G. †Mikhnocyathus Maslov 1957

Invalid names: Mikhnocyathus Maslov 1957 [objective synonym]

G. †Neoloculicyathus

G. †Okulitchicyathus

Invalid names: Alconeracyathus [synonym]

G. †Paranacyathus

Invalid names: Ramuscyathus Voronin et al. 1982 [synonym]

Superfm. †Sakhacyathoidea Debrenne and Zhuravlev 1990

Fm. †Sakhacyathidae Debrenne and Zhuravlev 1990

G. †Sakhacyathus

G. †Metyacyathellus

G. †Retillamina Debrenne and James 1981

Subor. †Syringocnemina Okulitch 1935

Superfm. †Auliscocyathoidea Debrenne and Zhuravlev 1992

Fm. †Auliscocyathidae Debrenne and Zhuravlev 1992

G. †Auliscocyathus

Superfm. †Fragilicyathoidea Belyaeva 1975

Fm. †Fragilicyathidae Belyaeva 1975

G. †Fragilicyathus

Superfm. †Kruseicnemoidea Debrenne and Zhuravlev 1990

Fm. †Kruseicnemidae Debrenne and Zhuravlev 1990

G. †Kruseicnema

Superfm. †Syringocnemoidea Taylor 1910

Fm. †Syringocnemidae Taylor 1910

G. †Pseudosyringocnema

G. †Syringocnema

G. †Syringothalamus Debrenne et al. 1990

G. †Williamicyathus

Fm. †Tuvacnemidae Debrenne and Zhuravlev 1990

G. †Tuvacnema

G. †Warrriootacyathus Gravestock 1984

Invalid names: Sphinctocyathus Zhuravleva 1962 [nomen dubium]

Or. †Capsulocyathida Zhuravleva 1964

Subor. †Capsulocyathina Zhuravelva 1964

Fm. †Cryptoporocyathidae Zhuravleva 1960

G. †Capsulocyathus

G. †Complicatocyathus Yaroshevich 1990

G. †Cryptoporocyathus Korshunov and Zhuravleva 1967

G. †Gerbicanicyathus Belyaeva 1969

G. †Mirandocyathus Belyaeva 1974

G. †Polythalamia Debrenne and Wood 1990

†Polythalamia americana Debrenne and Wood 1990

Fm. †Fransuasaecyathidae Debrenne 1964

G. †Fransuasaecyathus

G. †Yukonensis Ozdikmen 2009

†Yukonensis yukonensis Handfield 1967

Invalid names: Acanthopyrgus Handfield 1967 [replaced]

Fm. †Tubericyathidae Debrenne et al. 1989

G. †Tubericyathus

Fm. †Tylocyathidae Zhuravlev 1988

G. †Korshunovicyathus Zhuravlev 1988

G. †Tylocyathus

Fm. †Uralocyathellidae Zhuravleva 1964

G. †Rhabdolynthus

Invalid names: Uralocyathella Zhuravleva 1960 [synonym]

Invalid names: Thylocyathidae Zhuravlev 1988 [empty]

Subor. †Coscinocyathina Zhuravleva 1955

Superfm. †Alataucyathoidea Zhuravleva 1955

Fm. †Alataucyathidae Zhuravleva 1955

G. †Alataucyathus

Superfm. †Calyptocoscinoidea Debrenne 1964

Fm. †Calyptocoscinidae Debrenne 1964

G. †Calyptocoscinus

Fm. †Tomocyathidae Debrenne et al. 1989

G. †Coscinocyathella

Invalid names: Tomocyathus [synonym]

Superfm. †Clathricoscinoidea Rozanov 1964

Fm. †Clathricoscinidae Rozanov 1964

G. †Clathricoscinus

Fm. †Lanicyathidae Debrenne et al. 1989

G. †Lanicyathus

Superfm. †Coscinocyathoidea Taylor 1910

Fm. †Coscinocyathellidae Zhuravleva 1956

G. †Coscinocyathellus

Fm. †Coscinocyathidae Taylor 1910

G. †Coscinocyathus Bornemann 1884

G. †Flossocyathus Skorlotova 2013

Fm. †Mawsonicoscinidae Debrenne and Kruse 1986

G. †Mawsonicoscinus Debrenne and Kruse 1986

G. †Cyptoporocyathus Korshunov and Zhuravleva 1967

Or. †Kazachstanicyathida Konushkov 1967

Subor. †Altaicyathina Debrenne 1991

Fm. †Altaicyathidae Debrenne and Zhuravlev 1992

G. †Altaicyathus

Invalid names: Abakanicyathus Konyushkov 1964 [synonym]

Subor. †Kazachstanicyathina Konushkov 1967

Fm. †Korovinellidae Khalfina 1960

G. †Bicoscinus

G. †Korovinella

Or. †Monocyathida Okulitch 1935

G. †Arimasia Runnegar et al. 2024

†Arimasia germsi Runnegar et al. 2024

Fm. †Favilynthidae Debrenne 1989

G. †Favilynthus Debrenne and Zhuravleva 1974

G. †Robertiolynthus Zhuravleva 1987

†Robertiolynthus handfieldi Zhuravlev 1987

Invalid names: Veolynthus Zhuravleva et al. 1997 [synonym]

Fm. †Globosocyathidae Okuneva 1969

G. †Melkanicyathus Belyaeva 1969

G. †Propriolynthus Okuneva 1967

†Propriolynthus crassus Okuneva and Repina 1973

†Propriolynthus vologdini Yakovlev 1956

Invalid names: Globosocyathus Okuneva 1969 [synonym], Tumuloglobosus Okuneva and Repina 1973 [synonym]

Fm. †Monocyathidae Bedford and Bedford 1934

G. †Archaeolynthus Taylor 1910

Invalid names: Monocyathus Bedford 1934 [synonym], Rhabdocyathella [synonym]

G. †Kyarocyathus Kruse 1982

Fm. †Palaeoconulariidae Chudinova 1959

G. †Butakovicyathus Zhuravleva 1980

G. †Palaeoconularia Chudinova 1959

Fm. †Sajanolynthidae Rozanov 1989

G. †Sajanolynthus Vologdin and Kashina 1972

Fm. †Tumuliolynthidae Rozanov 1966

G. †Tumuliolynthus Zhuravleva 1963

Or. †Putapacyathida Vologdin 1961

Superfm. †Alphacyathoidea Bedford and Bedford 1939

Fm. †Alphacyathidae Bedford and Bedford 1939

G. †Alphacyathus Bedford and Bedford 1939

G. †Aptocyathus Vologdin 1937

†Aptocyathus prima Zhuravleva et al. 1964

Invalid names: Aptocyathella Zhuravleva et al. 1964 [synonym], Galinaecyathus [synonym]

Superfm. †Chabakovicyathoidea Rozanov 2002

Fm. †Chabakovicyathidae Rozanov 2002

G. †Chabakovicyathus Konyushkov 1964

Superfm. †Hupecyathoidea Debrenne et al. 1990

Fm. †Hupecyathidae Debrenne et al. 1990

G. †Hupecyathus Debrenne 1964

Superfm. †Putapacyathoidea Bedford and Bedford 1936

Fm. †Putapacyathidae Bedford and Bedford 1936

G. †Putapacyathus

Invalid names: Adaecyathus Zhuravlev et al. 1983 [nomen nudum], Butovia [nomen dubium], Cornutocyathus Zhuravleva et al. 1997 [nomen nudum], Irregulares Vologdin 1937 [invalid subgroup], Regulares Vologdin 1937 [invalid subgroup], Rhizacyathus Bedford 1939 [nomen nudum]

Diagnosis

Reference	Diagnosis
F. Debrenne et al. 2015	Skeleton nonspiculate, calcareous cup of microgranular microstructure and (with few exceptions) original magnesium calcite composition. Cup generally of archaeocya- than architecture with one or two porous walls bounding inner or central cavity respectively; porous to aporose septa, pseudosepta, taeniae, pseudotaeniae, pseudotaenial network, dicty- onal network, syringes, and/or tabulae may form in intervallum; a minority are of chae- tetid (intervallum with calicles) or thalamid architecture (cup consisting of successive chambers). Solitary or modular. Secondary calcareous skeleton may be present.

Measurements

No measurements are available

Locomotion:

stationaryp

Attached:

yesp

Life habit:

intermediate-level epifaunalp

Diet:

suspension feederp

Vision:

blindp

Comments:

LIFE HISTORY: Erect or surficial, non-motile attached, suspension feeder according to Bambach et al (2007); placed in intermediate-level tier according to Bottjer and Ausich (1986).p

Created:

2009-05-09 19:07:07

Modified:

2009-10-06 09:36:25

Source: p = phylum

References: Kiessling 2004, Bambach et al. 2007

Age range

Maximum range based only on fossils: base of the Ediacaran to the top of the Ashgill or 635.00000 to 443.70000 Ma
Minimum age of oldest fossil (stem group age): 541.0 Ma

Collections (413 total)

Time interval	Ma	Country or state	Original ID and collection number
Oldest occurrences
Ediacaran	635.0 - 541.0	India	Archaeocyatha indet. (10741)
Ediacaran	635.0 - 541.0	Namibia	Arimasia germsi (237049 237209)
Early/Lower Cambrian	538.8 - 509.0	USA (Nevada)	Archaeocyatha indet. (204025)
Early/Lower Cambrian	538.8 - 509.0	Falkland Islands (Malvinas)	Favilynthus contractus, Tumuliolynthus irregularis, Anaptyctocyathus sp., Erismacoscinus sp., Ethmocyathus lineatus, Flexanulus oosthuizeni, Polycoscinus scuratus, Polycoscinus papillatus, Thalamocyathus trachealis, Thalamocyathus sp., Pycnoidocyathus sp., Spirocyathella cooperi (182205) Thalamocyathus trachealis, Mawsonicoscinus sigmoides, Syringocnema sp. (182203)
Early/Lower Cambrian	538.8 - 509.0	USA (California)	Archaeocyatha indet. (226261)
Waucoban	538.8 - 509.0	USA (Washington)	Archaeocyatha indet. (264)
Tommotian	525.5 - 521.0	Russian Federation (Siberia)	Archaeolynthus polaris, Dokidocyathus regularis, Nochoroicyathus similis, Okulitchicyathus disciformis, Aldanocyathus anabarensis, Aldanocyathus virgatus, Aldanocyathus tkatchenkovi, Cryptoporocyathus junicanensis, Robustocyathus robustus (50321) Archaeolynthus polaris, Nochoroicyathus similis, Okulitchicyathus tkatchenkovi, Aldanocyathus anabarensis, Aldanocyathus virgatus, Aldanocyathus tkatchenkovi, Cryptoporocyathus junicanensis (50320) Cryptoporocyathus junicanensis, Aldanocyathus sunnaginicus (50319) Nochoroicyathus sp., Okulitchicyathus disciformis, Aldanocyathus anabarensis, Cryptoporocyathus junkanensis, Aldanocyathus sunnaginicus (50315)
Tommotian	525.5 - 521.0	Russian Federation (Kemorovo)	Archaeolynthus sp., Tumuliolynthus sp., Sibirecyathus sp., Capsulocyathus sp., Dictyocyathus yavorskii, Dictyocyathus sp. (56740)
Tommotian	525.5 - 521.0	Russian Federation	Archaeocyathida indet. (80846) Dictyocyathus translucidus, Paranacyathus tuberculatus, Cambrocyathus tschuranicus, Robustocyathus robustus (90090) Nochoroicyathus virgatus, Nochoroicyathus dragunovi, Nochoroicyathus igarcaensis, Nochoroicyathus sunnaginicus (90089) Paranacyathus tuberculatus, Robustocyathus robustus (90091) Tumuliolynthus primigenius, Arturocyathus varlamovi (10080)
Tommotian	525.5 - 521.0	Russian Federation (Yakutiya)	Ajacicyathus gigantoporus, Ajacicyathus sp., Dokidocyathus sp., Lenocyathus lenaicus, Nochoroicyathus grandis, Coscinocyathus sp., Coscinocyathus rojkovi, Cryptoporocyathus melnikovi (80951) Ajacicyathus subturbidus, Ajacicyathus centesimus, Ajacicyathus tkatchenkoi, Nochoroicyathus grandis, Retecoscinus proximus, Robustocyathus orbus, Robustocyathus apertus, Robustocyathus novus, Robustocyathus tetracyclicus, Taylorcyathus eximius, Coscinocyathus rojkovi, Monocyathus polaris, Rhizacyathus sp., Nevadacyathus sp., Cryptoporocyathus melnikovi, Cryptoporocyathus junicanensis (80950) Ajacicyathus tkatchenkoi, Ajacicyathus anabarensis, Nochoroicyathus mirabilis, Nochoroicyathus grandis, Retecoscinus proximus, Robustocyathus tetracyclicus, Robustocyathus apertus, Coscinocyathus rojkovi, Dictyocyathus translucidus, Okulitchicyathus discoformis, Paranacyathus tuberculatus, Cambrocyathellus tschuranicus, Sphinctocyathus oimuranicus, Monocyathus polaris, Cryptoporocyathus junicanensis, Robustocyathus robustus (80952) Archaeolynthus polaris, Ajacicyathus anabarensis, Nochoroicyathus mirabilis, Nochoroicyathus vulgarus, Nochoroicyathus grandis, Dictyocyathus translucidus, Loculicyathus sp., Paranacyathus subartus, Protopharetra polymorpha, Cambrocyathellus tschuranicus, Kotuyicyathus kotuyikensis, Sphinctocyathus sp., Robustocyathus robustus (96400) Archaeolynthus polaris, Ajacicyathus tkatschenkoi, Dokidocyathus regularis, Nochoroicyathus mirabilis, Nochoroicyathus vulgaris, Coscinocyathus rojkovi, Dictyocyathus translucidus, Okulitchicyathus disciformis, Paranacyathus tuberculatus, Cambrocyathellus tschuranicus (96251 96391 96394) Archaeolynthus polaris, Ajacicyathus virgatus, Nochoroicyathus vulgarus, Retecoscinus retetabulae, Coscinocyathus rojkovi, Dictyocyathus translucidus, Okulitchicyathus disciformis, Paranacyathus sp., Cambrocyathellus sp., Ajacicyathus sunnaginicus (10041) Archaeolynthus polaris, Cambrocyathellus tschuranicus (80866) Archaeolynthus polaris, Dictyocyathus translucidus, Okulitchicyathus discoformis, Cambrocyathellus tschuranicus (80804) Archaeolynthus polaris, Dictyocyathus translucidus, Okulitchicyathus discoformis, Cambrocyathellus tschuranicus, Cryptoporocyathus junicanensis (80796 80809 80812) Archaeolynthus polaris, Dictyocyathus translucidus, Okulitchicyathus discoformis, Cryptoporocyathus junicanensis (80815) Archaeolynthus polaris, Dokidocyathus regularis, Erismacoscinus rojkovi, Nochoroicyathus tkatschenkoi, Nochoroicyathus vulgaris, Nochoroicyathus virgatus, Nochoroicyathus anabarensis, Nochoroicyathus aldanicus, Nochoroicyathus prosper, Nochoroicyathus mirabilis, Rotundocyathus spinosus, Sclerocyathus australis, Dictyocyathus translucidus, Okulitchicyathus discoformis, Sakhacyathus subartus, Spinosocyathus maslennikovae, Cambrocyathellus proximus, Cambrocyathellus tschuranicus, Cryptoporocyathus junicanensis (80794) Archaeolynthus polaris, Dokidocyathus regularis, Erismacoscinus rojkovi, Nochoroicyathus vulgaris, Nochoroicyathus tkatschenkoi, Nochoroicyathus mirabilis, Nochoroicyathus prosper, Rotundocyathus spinosus, Dictyocyathus translucidus, Okulitchicyathus discoformis, Sakhacyathus subartus, Cambrocyathellus tschuranicus (80802) Archaeolynthus polaris, Erismacoscinus rojkovi, Nochoroicyathus mirabilis, Okulitchicyathus discoformis (80816) Archaeolynthus polaris, Erismacoscinus rojkovi, Nochoroicyathus mirabilis, Okulitchicyathus discoformis, Cambrocyathellus tschuranicus (80800 80807 80810) Archaeolynthus polaris, Okulitchicyathus discoformis, Cambrocyathellus tschuranicus (80813) Archaeolynthus polaris, Tumuliolynthus primigenius, Dokidocyathus regularis, Erismacoscinus rojkovi, Nochoroicyathus virgatus, Nochoroicyathus vulgaris, Nochoroicyathus tkatschenkoi, Nochoroicyathus aldanicus, Nochoroicyathus mirabilis, Nochoroicyathus prosper, Nochoroicyathus ridiculus, Nochoroicyathus transitorius, Rotundocyathus spinosus, Sclerocyathus australis, Dictyocyathus translucidus, Okulitchicyathus discoformis, Sakhacyathus subartus, Spinosocyathus maslennikovae, Cambrocyathellus tschuranicus, Cambrocyathellus proximus, Cryptoporocyathus junicanensis (80808) Archaeolynthus polaris, Tumuliolynthus primigenius, Erismacoscinus rojkovi, Nochoroicyathus tkatschenkoi, Nochoroicyathus anabarensis, Nochoroicyathus aldanicus, Nochoroicyathus mirabilis, Nochoroicyathus ridiculus, Nochoroicyathus pseudocultatus, Nochoroicyathus turbidus, Retecoscinus sakhaensis, Dictyocyathus translucidus, Okulitchicyathus discoformis, Cryptoporocyathus junicanensis (80814) Archaeolynthus polaris, Tumuliolynthus primigenius, Nochoroicyathus tkatschenkoi, Nochoroicyathus anabarensis, Nochoroicyathus aldanicus, Rotundocyathus spinosus, Dictyocyathus translucidus, Okulitchicyathus discoformis, Sakhacyathus subartus, Cambrocyathellus tschuranicus, Cryptoporocyathus junicanensis (80811) Archaeolynthus sp. (56858) Archaeolynthus sp., Ajacicyathus tkatchenkoi, Nochoroicyathus mirabilis, Robustocyathus sp., Dictyocyathus translucidus, Okulitchicyathus discoformis, Rhizacyathus sp., Cryptoporocyathus junicanensis (80953) Nochoroicyathus sp., Cryptoporocyathus junicanensis (80867) Nochoroicyathus virgatus, Nochoroicyathus dragunovi, Nochoroicyathus igarcaensis, Cryptaporocyathus junicanensis, Nochoroicyathus sunnaginicus (90088)
Tommotian	525.5 - 521.0	Russian Federation (Evenki)	Archaeolynthus polaris, Nochoroicyathus occultatus, Nochoroicyathus vulgaris, Robustocyathus moori, Aldanocyathus tkatschenkoi, Robustocyathus robustus (80873) Archaeolynthus polaris, Nochoroicyathus vulgaris, Retecoscinus retetabulae, Robustocyathus moori, Aldanocyathus fabrefactus, Aldanocyathus anabarensis, Aldanocyathus belvederi, Robustocyathus robustus (80869) Archaeolynthus polaris, Tumuliolynthus sp., Nochoroicyathus aldanicus, Robustocyathus moori, Coscinocyathus sp., Aldanocyathus sp., Robustocyathus robustus (80868)
Tommotian	525.5 - 521.0	Russian Federation (Yakutia)	Archaeolynthus polaris, Ajacicyathus tkatschenkoi, Ajacicyathus anabarensis, Ajacicyathus cautus, Ajacicyathus turbidus, Nochoroicyathus ridiculus, Robustocyathus moorei, Robustocyathus dotatus, Tennericyathus kotuyikensis, Capsulocyathus sp., Coscinocyathus rojkovi, Dictyocyathus translucidus, Paranacyathus subartus, Tumulocyathellus sp., Kotuyicyathus kotuyikensis (96393) Archaeolynthus polaris, Ajacicyathus tkatschenkoi, Ajacicyathus anabarensis, Ajacicyathus turbidus, Nochoroicyathus mirabilis, Nochoroicyathus supervacuus, Nochoroicyathus ridiculus, Retecoscinus retetabulae, Tumulocyathus sp., Coscinocyathus rojkovi, Dictyocyathus translucidus, Okulitchicyathus disciformis, Paranacyathus subartus, Robustocyathus robustus (10067) Archaeolynthus polaris, Ajacicyathus tkatschenkoi, Dokidocyathus regularis, Nochoroicyathus mirabilis, Coscinocyathus rojkovi, Cryptaporocyathus junicanensis, Robustocyathus robustus (45856) Archaeolynthus polaris, Ajacicyathus virgathus, Ajacicyathus tkatschenkoi, Dokidocyathus sp., Nochoroicyathus aldanicus, Robustocyathus belvederi, Cambrocyathus tschuranicus, Ajacicyathus sunnaginicus (45847) Archaeolynthus polaris, Ajacicyathus virgatus, Ajacicyathus tkatschenkoi, Nochoroicyathus vulgarus, Robustocyathus belvederi, Dictyocyathus translucidus, Okulitchicyathus disciformis, Paranacyathus tuberculatus, Cambrocyathellus tschuranicus, Ajacicyathus sunnaginicus, Robustocyathus robustus (96245) Coscinocyathus rojkovi, Dictyocyathus translucidus, Robustocyathus robustus (45850)
Tommotian	525.5 - 521.0	Russian Federation (Sakha)	Archaeolynthus polaris, Ajacicyathus tkatschenkoi, Ajacicyathus anabarensis, Nochoroicyathus aldanicus, Nochoroicyathus mirabilis, Coscinocyathus rojkovi, Dictyocyathus translucidus, Okulitchicyathus disciformis, Cryptoporocyathus junicanensis (232888) Archaeolynthus polaris, Ajacicyathus virgatus, Dokidocyathus sp., Nochoroicyathus aldanicus, Nochoroicyathus vulgaris, Okulitchicyathus disciformis, Cryptoporocyathus junicanensis, Ajacicyathus sunnaginicus (232887)
Tommotian	525.5 - 521.0	Russian Federation (Krasnoyarskiy Kray)	Archaeolynthus polaris, Nochoroicyathus tkatschenkoi, Nochoroicyathus anabarensis, Nochoroicyathus mirabilis, Nochoroicyathus occulathus, Retecoscinus retetabulae, Tumulocyathus kotuylkensis, Coscinocyathus rojkovi, Dictyocyathus translucidus, Paranacyathus tuberculatus, Protopharetra sp., Cambrocyathellus tschuranicus, Robustocyathus robustus (80841)
Tommotian	525.5 - 521.0	Russian Federation (Y)	Ajacicyathus anabarensis, Retecoscinus retetabulae, Coscinocyathus rojkovi, Dictyocyathus translucidus, Okulitchicyathus disciformis, Cambrocyathellus sp., Kotuyicyathus sp., Ramuscyathus proximus, Robustocyathus robustus (10093)
Atdabanian	521.0 - 516.0	Morocco	Afiacyathus compositus, Afiacyathus simplex, Erismacoscinus rojkovi, Erismacoscinus marocanus, Mennericyathus echinus, Retecoscinus zhuravlevae, Tumulocoscinus equiporus, Agastrocyathus gregarius, Neoloculicyathus abadiei (80906) Afiacyathus compositus, Afiacyathus tabulatus, Afiacyathus utriculoporus, Afiacyathus pruvosti, Erismacoscinus diourii, Erismacoscinus marocanus, Nochoroicyathus crassus, Nochoroicyathus infundibulus, Rotundocyathus hupei, Agastrocyathus gregarius, Neoloculicyathus abadiei, Protopharetra gemmata, Protopharetra grandicaveata, Geniculicyathus varius (232981) Afiacyathus compositus, Ajacicyathellus sp., Erismacoscinus primus, Nochoroicyathus crassus, Agastrocyathus gregarius, Protopharetra circula (80885) Afiacyathus compositus, Erismacoscinus fasciola, Nochoroicyathus sp., Retecoscinus minutus, Dictyocyathus stipatus, Neoloculicyathus magnus, Protopharetra circula (80893) Afiacyathus compositus, Erismacoscinus marocanus, Erismacoscinus primus, Mennericyathus echinus, Nochoroicyathus crassus, Retecoscinus minutus, Rotundocyathus sp., Agastrocyathus gregarius, Neoloculicyathus abadiei, Protopharetra circula, Protopharetra gemmata (80899) Afiacyathus compositus, Erismacoscinus primus, Agastrocyathus gregarius, Archaeopharetra sp., Neoloculicyathus magnus, Protopharetra taissensis, Protopharetra grandicaveata, Protopharetra circula, Tumulocyathidae indet. (80882) Afiacyathus compositus, Erismacoscinus primus, Rotundocyathus sp., Agastrocyathus gregarius, Neoloculicyathus magnus (80881) Afiacyathus compositus, Erismacoscinus primus, Tumulifungia marocana, Archaeopharetra sp., Neoloculicyathus magnus, Protopharetra grandicaveata (80891) Afiacyathus compositus, Nochoroicyathus crassus, Agastrocyathus gregarius, Neoloculicyathus abadiei, Protopharetra grandicaveata, Protopharetra taissensis (80907) Afiacyathus compositus, Protopharetra circula (80896) Afiacyathus compositus, Protopharetra taissensis (80880) Afiacyathus compositus, Tumulifungia marocana, Agastrocyathus gregarius (80886) Afiacyathus pruvosti, Erismacoscinus primus, Erismacoscinus fasciola, Rotundocyathus sp., Tumulifungia marocana, Agastrocyathus gregarius, Dictyocyathus stipatus, Neoloculicyathus magnus, Protopharetra circula (80892) Afiacyathus sp., Protopharetra taissensis (80883) Agastrocyathus gregarius, Neoloculicyathus abadiei, Protopharetra gemmata (80905) Erismacoscinus marocanus, Mennericyathus echinus, Nochoroicyathus crassus, Retecoscinus minutus, Tumulocoscinus equiporus, Agastrocyathus chouberti, Neoloculicyathus abadiei, Protopharetra stipata, Protopharetra gemmata (80900) Erismacoscinus primus, Erismacoscinus fasciola, Nochoroicyathus crassus, Retecoscinus minutus, Tumulifungia marocana, Agastrocyathus gregarius, Neoloculicyathus magnus, Protopharetra taissensis, Protopharetra circula (80898) Erismacoscinus primus, Erismacoscinus fasciola, Rotundocyathus sp., Neoloculicyathus magnus (80884) Erismacoscinus primus, Erismacoscinus fasciola, Tumulifungia marocana, Neoloculicyathus magnus, Protopharetra taissensis, Protopharetra grandicaveata, Protopharetra circula (80895) Erismacoscinus primus, Erismacoscinus sp. (80878) Mennericyathus asper (80908) Protopharetra taissensis (80879) Protopharetra taissensis, Protopharetra grandicaveata, Protopharetra circula (80897) Tumulifungia marocana, Neoloculicyathus magnus (80894)
Atdabanian	521.0 - 516.0	Spain (Extremadura)	Erismacoscinus sp., Mennericyathus sp., Nochoroicyathus sp., Rasetticyathus sp., Rotundocyathus sp., Taylorcyathus sp., Alconeracyathus sp., Chouberticyathus sp., Protopharetra sp. (81040) Erismacoscinus sp., Nochoroicyathus sp., Taylorcyathus sp., Coscinocyathus sp. (81039)
Atdabanian	521.0 - 516.0	Germany (Brandenburg)	Afiacyathus paracompositus, Afiacyathus sp., Erismacoscinus primus, Erismacoscinus sp., Erismacoscinus tainius, Coscinocyathus sp., Dictyocyathus stipatus, Protopharetra gemmata (24191) Afiacyathus paracompositus, Erismacoscinus sp., Coscinocyathus germanicus, Protopharetra gemmata (24192) Erismacoscinus tainius, Dictyocyathus stipatus, Protopharetra dissuta, Protopharetra gemmata (24189) Erismacoscinus tainius, Dictyocyathus stipatus, Protopharetra gemmata, Protopharetra dissuta (24190)
Atdabanian	521.0 - 516.0	Russian Federation (Siberia)	Archaeocyatha indet. (10011)
Atdabanian	521.0 - 516.0	China (Sichuan)	Dailycyathus sp., Erismacoscinus zunyiensis, Nochoroicyathus sp., Metacyathellus lepidus, Archaeocyatha indet. (224601)