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Mesocoleoptera
Taxonomy
Mesocoleoptera was named by Boudinot et al. (2022). It is extant.
It was assigned to Coleoptera by Boudinot et al. (2022).
It was assigned to Coleoptera by Boudinot et al. (2022).
Synonymy list
| Year | Name and author |
|---|---|
| 2022 | Mesocoleoptera Boudinot et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. Mesocoleoptera Boudinot et al. 2022
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Unr. Metacoleoptera Boudinot et al. 2022
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Fm. †Ademosynidae Ponomarenko 1968
+
G. †Ademosyne Handlirsch 1906
+
†Ademosyne adunca Dunstan 1923
†Ademosyne arcucciae Martins-Neto and Gallego 2006
†Ademosyne australiensis Tillyard 1916
†Ademosyne bacca Ponomarenko 1969
†Ademosyne brevis Dunstan 1923
†Ademosyne cameroni Tillyard 1916
†Ademosyne congener Tillyard 1916
†Ademosyne curvata Dunstan 1923
†Ademosyne elliptica Ponomarenko 1969
†Ademosyne elongatus Martins-Neto and Gallego 2006
†Ademosyne hexacostata Martins-Neto and Gallego 2006
†Ademosyne intermedia Dunstan 1923
†Ademosyne kirghizica Ponomarenko 1969
†Ademosyne lata Dunstan 1923
†Ademosyne llantenesensis Brauckmann et al. 2010
†Ademosyne major Handlirsch 1906
†Ademosyne olliffi Handlirsch 1906
†Ademosyne parva Dunstan 1923
†Ademosyne polyzetete Ponomarenko 2011
†Ademosyne prisca Riek 1974
†Ademosyne punctata Tillyard 1916
†Ademosyne punctuada Martins-Neto and Gallego 2006
†Ademosyne ramocostata Dunstan 1923
†Ademosyne reducta Riek 1976
†Ademosyne rosenfeldi Brauckmann et al. 2010
†Ademosyne rugulosa Dunstan 1923
†Ademosyne sibirica Rohdendorf 1961
†Ademosyne speciosa Riek 1974
†Ademosyne umutu Martins-Neto and Gallego 2009
†Ademosyne vittamargina Dunstan 1923
†Ademosyne wianamattensis Tillyard 1918
G. †Dolichosyne Ponomarenko 1969
+
†Dolichosyne confragosa Ponomarenko 1969
†Dolichosyne rostrata Ponomarenko 1969
†Dolichosyne sulcata Ponomarenko 1969
Subor. Adephaga Clairville 1806
+
G. †Amphoxyne Bode 1953
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†Amphoxyne darbiensis Ponomarenko 1986
†Amphoxyne minuta Bode 1953
+
Invalid names: Coilotrachelus lineatus Bode 1953 [synonym], Cricotrachelites rotundatus Bode 1953 [synonym], Loxoncus procerus Bode 1953 [synonym]
Invalid names: Coilotrachelus Bode 1953 [synonym], Cricotrachelites Bode 1953 [synonym], Loxostelidotus Bode 1953 [synonym]
Fm. Carabidae Latreille 1802 [ground beetle]
+
G. †Antemnacrassa Gomez Pallerola 1979
+
†Antemnacrassa albomaculata Nikolajev 2004
†Antemnacrassa albosulcata Nikolajev 2004
†Antemnacrassa geminata Nikolajev 2004
†Antemnacrassa incerta Ponomarenko 1990
†Antemnacrassa kazakhstanica Nikolajev 2007
†Antemnacrassa latitibialis Nikolajev 1992
†Antemnacrassa maculata Nikolajev 1992
†Antemnacrassa magna Nikolajev 2004
†Antemnacrassa monreali Gomez Pallerola 1979
†Antemnacrassa nebulosa Nikolajev 2004
†Antemnacrassa nigrimontana Nikolajev 1992
†Antemnacrassa picturata Nikritin 1977
†Antemnacrassa punctata Ponomarenko 1986
†Antemnacrassa sulcata Nikritin 1977
†Antemnacrassa vitimensis Nikritin 1977
G. Apotomopterus Hope 1838
G. Bembidium Latreille 1802
+
Subg. Bembidium (Campa) Motschulsky 1844
Subg. Bembidium (Testidium) Motschulsky 1864
†Bembidium absolutum Giebel 1856
†Bembidium cyaneomicans Piton 1940
†Bembidium exoletum Scudder 1876
†Bembidium expletum Scudder 1900
†Bembidium florissantensis Wickham 1913
†Bembidium fragmentum Scudder 1890
†Bembidium glaciatum Scudder 1890
†Bembidium haywardi Scudder 1900
†Bembidium infernum Heer 1856
†Bembidium laevigatum Förster 1891
†Bembidium obductum Scudder 1900
†Bembidium praeteritum Scudder 1900
†Bembidium saportanum Oustalet 1874
†Bembidium tumulorum Scudder 1900
†Bembidium vanum Scudder 1900
†Bembidium vestigium Scudder 1900
Subfm. Carabinae Latreille 1802
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G. †Atrirabus Hong and Wang 1990
Tr. Cychrini Perty 1830
Tr. Pamborini Hope 1838
G. †Carabites Ponomarenko 1977
+
†Carabites caraboides Ponomarenko 1989
†Carabites nigriventris Ponomarenko 1986
G. †Carabites Heer 1852
+
†Carabites anthracinus Heer 1852
†Carabites arapahoensis Cockerell 1918
†Carabites capellinii Cecconi 1892
†Carabites creta Ponomarenko 1977
†Carabites cuneatus Cockerell 1920
†Carabites dubius Geinitz 1894
†Carabites ellipticus Haupt 1956
†Carabites eocenicus Cockerell 1921
†Carabites exanimus Scudder 1892
†Carabites exilis Heer 1862
†Carabites feildenianus Heer 1878
†Carabites gardneri Cockerell 1920
†Carabites geinitzi Handlirsch 1906
†Carabites harringtoni Cockerell 1936
†Carabites islandicus Heer 1868
†Carabites kincaidi Cockerell 1908
†Carabites laesicollis Heer 1862
†Carabites parallelus Cockerell 1920
†Carabites peracutus Cockerell 1920
†Carabites rugosus Meunier 1898
†Carabites russelli Cockerell 1928
†Carabites schueli Cockerell 1926
†Carabites scoticus Cockerell 1921
†Carabites vitimensis Ponomarenko 1977
G. †Carabopteron Martynov 1926
+
†Carabopteron punctatolineatum Martynov 1926
†Carabopteron rugosum Ponomarenko 1985
Unr. Celeripedes Latreille 1802
+
Invalid names: Barbati Latreille 1802 [empty], Crepitantes Latreille 1802 [empty], Longipalpati Latreille 1802 [empty], Melanchlaeni Latreille 1802 [empty], Metallici Latreille 1802 [empty]
Subfm. †Conjunctiinae Ponomarenko 1977
+
G. †Conjunctia Ponomarenko 1977
G. †Mesorabus Ponomarenko 1977
G. †Fonsecacarabus Martins-Neto and Mendes 2002
+
†Fonsecacarabus placidus Martins-Neto and Mendes 2002
G. †Hadrocephalus Handlirsch 1906
+
†Hadrocephalus anglicus Handlirsch 1906
†Hadrocephalus liasinus Handlirsch 1906
†Hadrocephalus minor Handlirsch 1906
Subfm. Harpalinae Bonelli 1810
+
Tr. Abacetini Chaudoir 1873
G. †Aethocarabus Ren 1995
Tr. Chlaeniini Brullé 1834
Tr. Galeritini Kirby 1825
Tr. Harpalini Bonelli 1810
Tr. Helluonini Hope 1838
Tr. Lebiini Bonelli 1810
Tr. Licinini Bonelli 1810
Tr. Panagaeini Bonelli 1810
Tr. Pseudomorphini Hope 1838
Tr. Pterostichini Bonelli 1810
G. †Sinis Heer 1862
Tr. Sphodrini Laporte 1834
Tr. Zabrini Bonelli 1810
Tr. Zuphiini Bonelli 1810
Invalid names: Dryptini Bonelli 1810 [empty], Licininae Bonelli 1810 [synonym]
Subfm. Loricerinae Bonelli 1810
+
Tr. Loricerini Bonelli 1810
Invalid names: Loriceritae Bonelli 1810 [empty]
Subfm. Nebriinae Laporte 1834
+
G. †Ledouxnebria Deuve 1998
Tr. Nebriini Laporte 1834
Tr. Notiophilini Motschulsky 1850
Tr. Opisthiini Dupuis 1912
Tr. Pelophilini Kavanaugh 1996
G. †Nebrorabus Ponomarenko 1989
+
†Nebrorabus baculum Ponomarenko 1989
†Nebrorabus capitatus Ponomarenko 1989
†Nebrorabus nebrioides Ponomarenko 1989
Subfm. Paussinae Latreille 1806 [ant nest beetle]
+
G. †Cerapterites Wasmann 1926
Tr. Paussini Latreille 1806
G. †Paussoides Motschulsky 1856
G. †Paussopsis Cockerell 1911
G. †Pleurarthropterus Wasmann 1929
Invalid names: Ozaenidae Hope 1838 [synonym], Ozaenini [empty]
G. †Procarabites Handlirsch 1906
Subfm. †Protorabinae Ponomarenko 1977
+
G. †Cordorabus Ponomarenko 1977
G. †Cretorabus Ponomarenko 1977
G. †Lirabus Hong 1992
G. †Lithorabus Ponomarenko 1977
G. †Magnirabus Hong and Wang 1990
G. †Ovrabites Ponomarenko 1977
G. †Penecupes Ren 1995
G. †Protorabus Ponomarenko 1977
Subfm. Scaritinae Bonelli 1810
+
Tr. Clivinini Rafinesque 1815
Tr. Dyschiriini Kolbe 1880
G. †Dyschiriomimus Yablokov-Khnzorian 1960
Tr. †Palaeoaxinidiini McKay 1991
Tr. Scaritini Bonelli 1810
Invalid names: Promecognathinae Le Conte 1853 [synonym]
G. Stereocerus Kirby 1837
+
Stereocerus haematopus Dejean 1831
Stereocerus rubripes Motschulsky 1860
Subfm. Trechinae Bonelli 1810
+
Tr. Patrobini Kirby 1837
G. †Trechinites Heer 1862
Unr. Trechitae Bonelli 1810
Invalid names: Bembidion succini Giebel 1856 [nomen dubium], Patrobinae Kirby 1837 [synonym]
Invalid names: Cratacanthus florissantensis Wickham 1917 [nomen dubium], Fossores Latreille 1802 [empty], Loricera exita Scudder 1900 [nomen dubium], Rhysodinae Laporte 1840 [empty], Siagoninae Bonelli 1813 [empty], Trechus fractus Wickham 1912 [nomen dubium]
Infraor. Carabiformia Lameere 1900
+
Superfm. Caraboidea Latreille 1802
+
G. †Beipiaocarabus Hong 1983
G. †Carabilarva Ponomarenko 1985
G. †Eurycoleus Hong 1982
G. †Glottocoleus Hong 1982
G. †Glyptocoleus Hong 1982
G. †Grypocoleus Hong 1982
G. †Jiangxicoleus Hong 1988
G. †Leptocoleus Hong 1982
G. †Mesocoleus Hong 1982
G. †Obesofemoria Hong 1982
G. †Phyllocoleus Hong 1982
G. †Pleurocoleus Hong 1982
G. †Procalosoma Meunier 1895
G. †Vago Hong 1982
G. †Yumenocoleus Hong 1982
Invalid names: Unda chonggezhuangensis Wang 1998 [nomen dubium], Unda pandurata Ren 1995 [nomen dubium]
Fm. Cicindelidae Latreille 1802 [tiger beetle]
+
G. †Hujia Song et al. 2022
Tr. Manticorini Laporte 1834
+
G. Omus Eschscholtz 1829
Invalid names: Amblycheilini Csiki 1903 [synonym]
Tr. Megacephalini Laporte 1834
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G. Tetracha Hope 1838
Invalid names: Megacephalina Laporte 1834 [empty]
Invalid names: Oxycheilini Thomson 1857 [empty]
Fm. †Colymbotethidae Ponomarenko 1993
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Fm. †Coptoclavidae Ponomarenko 1961
+
G. †Amblycephalonius Bode 1953
+
†Amblycephalonius tenuistriatus Bode 1953
+
Invalid names: Barycephalus nudatus Bode 1953 [synonym], Tricyrtus tenuistriatus Bode 1953 [synonym]
Invalid names: Barocephalus Carpenter 1986 [synonym], Barycephalus Bode 1953 [synonym], Tricyrtus Bode 1953 [synonym]
Subfm. †Charonoscaphinae Ponomarenko 1977
+
G. †Charonoscapha Ponomarenko 1977
G. †Charonoscaphidia Ponomarenko 1977
Subfm. †Coptoclavinae Ponomarenko 1961
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G. †Bolbonectes Ponomarenko 1987
G. †Coptoclava Ping 1928
G. †Hoyaclava Soriano et al. 2007
G. †Megacoptoclava Ponomarenko and Martínez-Delclòs 2000
Subfm. †Coptoclaviscinae Soriano et al. 2007
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G. †Coptoclavella Ponomarenko 1980
G. †Coptoclavisca Ponomarenko 1987
G. †Holcoptera Handlirsch 1906
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Invalid names: Holcoelytrum Handlirsch 1906 [synonym], Stargelytron Ponomarenko 2015 [synonym]
Subfm. †Timarchopsinae Wang et al. 2010
+
G. †Daohugounectes Wang et al. 2009
G. †Protonectes Prokin and Ponomarenko 2013
Invalid names: Necronectinae Ponomarenko 1977 [replaced]
Fm. Dytiscidae Leach 1815 [predaceous diving beetle]
+
Subfm. Copelatinae Van den Branden 1885
+
G. Copelatus Erichson 1832
Invalid names: Copelatini Van den Branden 1885 [empty]
G. †Cretodytes Ponomarenko 1977
+
†Cretodytes incertus Prokin et al. 2013
†Cretodytes latipes Ponomarenko 1977
Subfm. Dytiscinae Leach 1815
+
Tr. Aciliini Thomson 1867
Tr. †Ambarticini Yang et al. 2019
Tr. Dytiscini Leach 1815
Tr. Eretini Crotch 1873
Tr. Hydaticini Sharp 1880
Subfm. Hydroporinae Aubé 1836
+
Tr. Bidessini Sharp 1880
G. †Glessaria Koch and Berendt 1854
Tr. Hydroporini Aubé 1836
Tr. Hygrotini Portevin 1929
Tr. Hyphydrini Gistel 1848
Tr. Methlini Van den Branden 1885
G. †Procoelambus Theobald 1937
Tr. †Schistomerini Palmer 1957
Tr. Vatellini Sharp 1880
Subfm. Laccophilinae Gistel 1848
+
G. Japanolaccophilus Sato 1972
Tr. Laccophilini Gistel 1848
Invalid names: Palaeogyrininae von Schlechtendal 1894 [synonym]
Subfm. †Liadytiscinae Prokin and Ren 2010
+
G. †Liadroporus Prokin and Ren 2010
Tr. †Liadytiscini Prokin and Ren 2010
Tr. †Mesoderini Prokin et al. 2013
G. †Palaeodytes Ponomarenko 1987
+
†Palaeodytes baissiensis Prokin et al. 2013
†Palaeodytes gutta Ponomarenko 1987
†Palaeodytes incompleta Ponomarenko et al. 2005
†Palaeodytes sibiricus Ponomarenko 1987
Invalid names: Aubehydrinae Guignot 1942 [empty], Necticus palustris Aymard 1856 [nomen nudum]
Fm. Gyrinidae Latreille 1810 [whirligig beetle]
+
Subfm. Gyrininae Latreille 1810
+
G. †Chimerogyrus Gustafson et al. 2020
Tr. Gyrinini Latreille 1810
Tr. Orectochilini Kolbe 1880
G. †Protogyrininus Hatch 1927
+
†Protogyrininus sculpturatus Mjöberg 1905
+
Invalid names: Protogyrininus confinis Hatch 1927 [synonym]
G. †Tunguskagyrus Yan et al. 2018
+
†Tunguskagyrus planus Yan et al. 2018
†Tunguskagyrus yani Ponomarenko 2021
Invalid names: Spanglerogyrinae Folkerts 1979 [empty]
Subfm. Haliplidae Aubé 1836 [crawling water beetle]
+
G. †Cretihaliplus Ren et al. 1995
+
†Cretihaliplus chifengensis Ren et al. 1995
†Cretihaliplus sidaojingensis Ren et al. 1995
G. Haliplus Latreille 1802
+
†Haliplus cretaceus Prokin and Ponomarenko 2013
Haliplus obliquus Fabricius 1787
Haliplus ruficollis De Geer 1774
G. Peltodytes Regimbart 1878
Fm. Hygrobiidae Regimbart 1878
+
G. Hygrobia Latreille 1804
+
†Hygrobia cretzschmari von Heyden and von Heyden 1866
Invalid names: Paelobius Schoenherr 1808 [synonym]
Invalid names: Paelobiidae Erichson 1837 [synonym]
Fm. †Liadytidae Ponomarenko 1977
+
G. †Liadytes Ponomarenko 1963
+
†Liadytes aspidytoides Prokin et al. 2013
†Liadytes avus Ponomarenko 1963
†Liadytes crassus Ponomarenko 1977
†Liadytes dajensis Ponomarenko 1987
†Liadytes longus Ponomarenko 1977
†Liadytes major Ponomarenko 1985
G. †Memptus Handlirsch 1906
+
†Memptus ganglbaueri Ponomarenko 1985
†Memptus guttula Ponomarenko 1985
†Memptus handlirschi Ponomarenko 1985
Fm. †Parahygrobiidae Ponomarenko 1977
+
Fm. Trachypachidae Thomson 1857 [false ground beetle]
+
Subfm. †Eodromeinae Ponomarenko 1977
+
G. †Ademosynoides Dunstan 1923
G. †Coreoeicos Bode 1953
G. †Dolichorabus Ponomarenko 1985
G. †Eodromeus Ponomarenko 1977
G. †Karadromeus Ponomarenko 1977
G. †Karatoma Ponomarenko 1977
G. †Liassodites Kirejtshuk and Ansorge 2022
G. †Necronectulus Ponomarenko 1977
G. †Permunda Ponomarenko and Volkov 2013
G. †Petrodromeus Ponomarenko and Volkov 2013
G. †Platycoxa Ponomarenko 1977
G. †Prosynactus Bode 1953
+
Invalid names: Critotrachelus Carpenter 1986 [synonym], Cyclotrachelus Bode 1953 [synonym], Dysmorphus Bode 1953 [synonym], Euenarthrus Bode 1953 [synonym], Leptynticus Bode 1953 [synonym], Liassocarabites Bode 1953 [synonym], Megelytrites Bode 1953 [synonym], Mesalocistus Bode 1953 [synonym], Rhabdotus Bode 1953 [synonym], Tetragonotrachelus Bode 1953 [synonym], Trapezotrachelus Bode 1953 [synonym], Trigonocephalites Bode 1953 [synonym]
G. †Psacodromeus Ponomarenko 1977
G. †Sinodromeus Wang et al. 2012
G. †Sogdodromeus Ponomarenko 1977
G. †Tauredon Handlirsch 1910
G. †Xishanocarabus Hong 1984
G. †Eodromeites Ponomarenko 1985
+
†Eodromeites minimus Ponomarenko 1985
†Eodromeites udensis Ponomarenko 1985
Invalid names: Karadromeus xiangfanggouensis Hong 1995 [nomen dubium]
Fm. †Triaplidae Ponomarenko 1977
+
G. †Catinius Ponomarenko 1968
+
†Catinius artus Zhang 1997
†Catinius ovatus Zhang 1997
†Catinius pelta Ponomarenko 1968
G. †Tomiaplus Ponomarenko 2021
+
†Tomiaplus minimus Ponomarenko 2021
†Tomiaplus sibiricus Volkov 2013
G. †Triaplus Ponomarenko 1977
+
†Triaplus laticoxa Ponomarenko 1977
†Triaplus macroplatus Ponomarenko 1977
G. †Triassocatinius Ponomarenko 1969
+
†Triassocatinius brachynotus Ponomarenko 1969
†Triassocatinius glabratus Ponomarenko 1969
Invalid names: Catiniidae Ponomarenko 1968 [replaced]
Invalid names: Amphizoidae Leconte 1853 [empty], Dytiscoidea Leach 1815 [invalid subgroup], Pseudomorphinae Hope 1838 [empty], Tritarsusidae Hong 2002 [invalid subgroup]
Subor. Archostemata Kolbe 1908
+
G. †Argentinocupes Martins-Neto and Gallego 2006
+
†Argentinocupes abdalai Martins-Neto and Gallego 2006
†Argentinocupes pulcher Martins-Neto and Gallego 2006
†Argentinocupes sara Martins-Neto and Gallego 2009
Superfm. †Asiocoleoidea Rohdendorf 1961
+
Fm. †Tricoleidae Ponomarenko 1969
+
G. †Triassocupes Hong 1985
G. †Tricoleodes Ponomarenko 1969
G. †Tricoleus Ponomarenko 1969
G. †Willcoxia Dunstan 1923
G. †Brachilatus Strelnikova and Yan 2023
+
†Brachilatus caducus Ponomarenko 1969
†Brachilatus dundulaensis Ponomarenko 1994
†Brachilatus foersteri Ponomarenko 1971
+
Invalid names: Notocupes reticulatus Ponomarenko 1968 [replaced]
†Brachilatus longicoxa Soriano and Delclos 2006
†Brachilatus nigrimonticola Ponomarenko 1968
†Brachilatus oculatus Soriano and Delclos 2006
†Brachilatus premeris Lee et al. 2022
†Brachilatus spinosus Li and Cai 2023
†Brachilatus viridis Soriano and Delclos 2006
G. †Conexicoxa Lin 1986
+
†Conexicoxa brachycephala Ponomarenko 1994
†Conexicoxa crassa Ponomarenko 1969
†Conexicoxa denticollis Jiang et al. 2020
†Conexicoxa epicharis Tan et al. 2005
†Conexicoxa eumeura Tan et al. 2006
†Conexicoxa exigua Ponomarenko 1994
†Conexicoxa homora Lin 1986
†Conexicoxa jurassica Hong 1983
†Conexicoxa khetanensis Ponomarenko 1993
†Conexicoxa kirghizica Ponomarenko 1969
†Conexicoxa longicollis Ponomarenko 1994
†Conexicoxa martinclosas Soriano and Delclos 2006
†Conexicoxa porrecta Tan et al. 2006
†Conexicoxa psilata Tan et al. 2005
†Conexicoxa siniestri Soriano and Delclos 2006
†Conexicoxa stabilis Tan et al. 2005
†Conexicoxa tripartita Oppenheim 1888
Superfm. Cupedoidea Laporte 1836
+
Fm. Cupedidae Laporte 1836 [reticulated beetle]
+
G. †Cainocups Kirejtshuk et al. 2016
Subfm. Cupedinae Lacordaire 1857
Subfm. Priacminae Crowson 1962
Invalid names: Mesocupedinae Ponomarenko 1969 [empty]
Fm. Micromalthidae Barber 1913 [telephone pole beetle]
+
G. †Archaeomalthus Yan et al. 2020
G. †Cretomalthus Kirejtshuk and Azar 2008
G. Micromalthus LeConte 1878
G. †Protomalthus Tihelka et al. 2020
Fm. Ommatidae Sharp and Muir 1912
+
G. †Asiania Lee et al. 2023
G. †Clessidromma Jarzembowski et al. 2017
G. †Kirejtomma Li and Cai 2021
G. †Limnomma Li and Cai 2021
G. †Miniomma Li et al. 2020
Subfm. Tetraphalerinae Crowson 1962
Invalid names: Crowsoniellidae Iablokoff-Khnzorian 1983 [empty], Magnocoleidae Hong 1998 [invalid subgroup]
G. †Notocupes Ponomarenko 1964
+
†Lupicupes trachylaenus Ren 1995
†Notocupes alienus Tan and Ren 2006
†Notocupes baojiatunensis Hong 1992
†Notocupes caudatus Ponomarenko 1966
†Notocupes cyclodontus Tan et al. 2006
†Notocupes daohugouensis Li and Cai 2023
†Notocupes diazromerali Soriano and Delclos 2006
†Notocupes elegans Ponomarenko 1994
†Notocupes excellens Ponomarenko 1966
†Notocupes kezuoensis Hong 1987
†Notocupes khasurtyiensis Strelnikova 2019
†Notocupes laetus Lin 1976
†Notocupes laiyangensis Hong and Wang 1990
†Notocupes lapidarius Ponomarenko 1968
†Notocupes latus Ponomarenko 1969
†Notocupes lini Ponomarenko et al. 2012
†Notocupes ludongensis Wang and Liu 1996
†Notocupes mongolicus Ponomarenko 1994
†Notocupes neli Tihelka et al. 2019
†Notocupes ohmkuhnlei Jarzembowski et al. 2019
†Notocupes picturatus Ponomarenko 1964
†Notocupes pingi Ponomarenko and Ren 2010
†Notocupes pulcher Ponomarenko 1968
†Notocupes reticulatus Oppenheim 1888
†Notocupes robustus Li and Cai 2023
†Notocupes rudis Tan et al. 2005
†Notocupes shiluoensis Hong 1984
†Notocupes tuanwangensis Hong and Wang 1990
†Notocupes validus Lin 1976
Invalid names: Amblomma Tan et al. 2005 [synonym], Chengdecupes Hong 1983 [synonym], Forticupes Hong and Wang 1990 [synonym], Lupicupes Ren 1995 [synonym], Ovatocupes Tan and Ren 2006 [synonym], Picticupes Hong and Wang 1990 [synonym], Sinocupes Lin 1976 [synonym]
G. †Pseudosilphites Zeuner 1930
+
†Pseudosilphites natalensis Zeuner 1960
†Pseudosilphites triassicus Zeuner 1930
G. †Rhabdocupes Ponomarenko 1966
+
†Rhabdocupes baculatus Ponomarenko 1969
†Rhabdocupes cellulosus Ponomarenko 1969
†Rhabdocupes dischides Zhang 1986
†Rhabdocupes issykkulensis Ponomarenko 1969
†Rhabdocupes laticella Ponomarenko 1969
†Rhabdocupes lentus Ren 1995
†Rhabdocupes longus Ponomarenko 1966
†Rhabdocupes minisculus Tan et al. 2006
†Rhabdocupes minor Ponomarenko 1966
†Rhabdocupes oxypygus Ponomarenko 1969
†Rhabdocupes protensus Tan et al. 2006
†Rhabdocupes rostratus Ponomarenko 1969
†Rhabdocupes sogutensis Ponomarenko 1969
†Rhabdocupes tenuis Ponomarenko 1969
†Rhabdocupes vitimensis Ponomarenko 1966
Superfm. †Rhombocoleoidea Rohdendorf 1961
+
Fm. †Rhombocoleidae Rohdendorf 1961
+
G. †Rhombocoleites Ponomarenko 1969
G. †Rhombocoleus Rohdendorf 1961
G. †Sinorhombocoleus Tan and Ren 2009
Superfm. †Schizocoleoidea Rohdendorf 1961
+
Fm. †Phoroschizidae Bouchard and Bousquet 2020
+
G. †Phoroschizus Bouchard and Bousquet 2020
+
Invalid names: Schizophorus Ponomarenko 1968 [replaced]
Invalid names: Schizophoridae Ponomarenko 1968 [replaced]
Fm. †Schizocoleidae Rohdendorf 1961
+
G. †Aenigmocoleus Rohdendorf 1961
G. †Argentinosyne Martins-Neto and Gallego 2006
G. †Mesolobites Carpenter 1986
+
Invalid names: Lobites Dunstan 1923 [replaced], Shijingocoleus Lin and Mou 1989 [synonym]
G. †Mesosagrites Martynov 1935
G. †Pseudochrysomelites Handlirsch 1906
G. †Reeveana Dunstan 1923
G. †Schizocoleus Rohdendorf 1961
G. †Shepherdia Dunstan 1923
G. †Stegosyne Rohdendorf 1961
G. †Uskatocoleus Rohdendorf 1961
Invalid names: Schizophoroidea Ponomarenko 1968 [synonym]
G. †Tentyridium Westwood 1854
+
†Tentyridium dilatatum Handlirsch 1906
†Tentyridium peleus Westwood 1854
G. †Tillyardiopsis Dunstan 1923
+
†Tillyardiopsis granulata Dunstan 1923
†Tillyardiopsis tuberculata Dunstan 1923
†Tillyardiopsis variotuberculata Dunstan 1923
Invalid names: Cupediformia [invalid subgroup], Moravocoleus perditus Kukalová 1968 [nomen dubium], Notocupedini Ponomarenko 1966 [invalid subgroup], Priacma corrupta Ponomarenko 1986 [nomen dubium], Priacma longicapitis Ponomarenko 1997 [nomen dubium], Priacma oculata Ponomarenko 1997 [nomen dubium], Priacma striata Ponomarenko 2000 [nomen dubium]
Fm. †Asiocoleidae Rohdendorf 1961
+
G. †Bicoleus Ponomarenko 2013
+
†Bicoleus kuplensis Ponomarenko 2013
†Bicoleus laticella Ponomarenko 2013
G. †Loculitricoleus Tan and Ren 2009
+
†Loculitricoleus flatus Tan and Ren 2009
†Loculitricoleus mongolicus Ponomarenko 2014
†Loculitricoleus tenuatus Tan and Ren 2009
G. †Schizotaldycupes Rohdendorf 1961
+
†Schizotaldycupes ananjevi Rohdendorf 1961
†Schizotaldycupes beckermigdisovae Rohdendorf 1961
†Schizotaldycupes pubescens Ponomarenko 2013
Invalid names: Carinicupes Rohdendorf 1961 [synonym]
G. †Tetracoleus Ponomarenko 2008
+
†Tetracoleus aristovi Ponomarenko 2018
†Tetracoleus golubevi Ponomarenko 2015
†Tetracoleus meyeni Ponomarenko 2013
†Tetracoleus minimus Ponomarenko 2008
†Tetracoleus orenburgensis Ponomarenko 2013
†Tetracoleus sennikovi Ponomarenko 2011
†Tetracoleus tricoleoides Ponomarenko 2013
†Tetracoleus tshalyshevi Ponomarenko 2008
†Tetracoleus vasilenkoi Ponomarenko 2018
Subor. Myxophaga Crowson 1955
+
Superfm. Lepiceroidea Hinton 1936
+
Fm. Lepiceridae Hinton 1936
+
G. †Haplochelus Kirejtshuk and Poinar 2006
+
Invalid names: Lepiceratus Jałoszyński et al. 2020 [synonym]
G. Lepicerus Motschulsky 1855
G. †Lepichelus Kirejtshuk and Poinar 2017
+
Invalid names: Lepiceroides Kirejtshuk and Poinar 2013 [replaced]
Invalid names: Cyathoceridae Scharp 1882 [synonym], Haplochelidae Kirejtshuk and Poinar 2006 [synonym], Lepicerinae Hinton 1936 [empty]
Superfm. Sphaeriusoidea
+
Fm. Hydroscaphidae LeConte 1874 [skiff beetle]
+
Subfm. Hydroscaphinae LeConte 1874
Subfm. †Leehermaniinae Kirejtshuk et al. 2023
Subfm. †Triamyxinae Qvarnström et al. 2021
Fm. Sphaeriusidae Erichson 1845
+
G. Bezesporum Fikáček et al. 2022
G. †Burmasporum Kirejtshuk 2009
G. †Crowsonaerius Li and Cai 2023
G. Sphaerius Waltl 1838
Invalid names: Microsporidae Crotch 1873 [synonym]
Invalid names: Sphaeriidae Erichson 1845 [empty], Torridincolidae Steffan 1964 [empty]
Invalid names: Sphaeriiformia [empty]
Fm. †Peltosynidae Yan et al. 2017
+
G. †Peltosyne Ponomarenko 1977
+
†Peltosyne triassica Ponomarenko 1977
†Peltosyne varyvrosa Yan et al. 2017
Subor. Polyphaga Emery 1886
+
G. †Angarolarva Ponomarenko 1985
+
†Angarolarva aquatica Ponomarenko 1985
†Angarolarva incerta Ponomarenko 1990
Infraor. Bostrichiformia Forbes 1926
+
Superfm. Bostrichoidea Latreille 1802
+
Fm. Dermestidae Latreille 1804 [skin beetle]
Invalid names: Endecatomidae LeConte 1861 [empty]
Unr. Clambiformia Cai and Tihelka 2022
+
Superfm. Clamboidea Fischer von Waldheim 1821
+
Fm. Clambidae Fischer von Waldheim 1821 [minute beetle]
+
Invalid names: Ptismidae Kirejtshuk and Azar 2016 [synonym]
Fm. Derodontidae LeConte 1861 [tooth-necked fungus beetle]
Fm. Eucinetidae Lacordaire 1857 [plate-thigh beetle]
Fm. †Mesocinetidae Kirejtshuk and Ponomarenko 2010
Invalid names: Derodontoidea LeConte 1861 [synonym], Eucinetoidea Lacordaire 1857 [synonym]
Infraor. Cucujiformia Lameere 1938
+
Superfm. Cleroidea Latreille 1802
+
Fm. Acanthocnemidae Crowson 1952
Fm. Biphyllidae Leconte 1861 [false skin beetle]
Fm. Byturidae Gistel 1848
Fm. Cleridae Latreille 1802 [checkered beetle]
G. †Foveapeltis Li et al. 2024
G. †Juraniscus Kolibáč and Huang 2019
Fm. Lophocateridae Crowson 1964
G. †Mathesius Kolibáč and Huang 2011
Fm. Melyridae Leach 1815 [soft-winged flower beetle]
Fm. Peltidae Kirby 1837 [shield beetle]
Fm. Prionoceridae Lacordaire 1857
Fm. Rhadalidae LeConte 1861
G. †Shartegopsis Kirejtshuk 2012
Fm. Thanerocleridae Chapin 1924
Fm. Thymalidae Leveille 1888
Fm. Trogossitidae Latreille 1802 [bark gnawing beetle]
+
Invalid names: Ostomatidae Reitter 1882 [synonym], Temnochilidae [replaced]
Invalid names: Chaetosomatidae Crowson 1952 [empty], Phloiophilidae Kiesenwetter 1863 [empty], Phycosecidae Crowson 1952 [empty], Protopeltidae Crowson 1966 [empty], Rentoniidae Crowson 1966 [empty]
Superfm. Coccinelloidea Latreille 1807
+
Fm. Anamorphidae Strohecker 1953
Fm. Bothrideridae Erichson 1845
Fm. Cerylonidae Billberg 1820
Fm. Coccinellidae Latreille 1807 [ladybird beetle]
Fm. Corylophidae Le Conte 1852 [minute hooded beetle]
Fm. Discolomatidae Horn 1878
Fm. Endomychidae Leach 1815 [handsome fungus beetle]
Fm. Latridiidae Erichson 1842 [minute brown scavenger beetle]
Fm. Teredidae Seidlitz 1888
Fm. †Tetrameropsidae Kirejtshuk and Azar 2008
Invalid names: Alexiidae Imhoff 1856 [empty], Eupsilobiidae Casey 1895 [empty], Euxestidae Grouvelle 1908 [empty], Murmidiidae Jacquelin du Val 1858 [empty], Mycetaeidae Jacquelin du Val 1857 [empty]
Superfm. Cucujoidea Latreille 1802
+
G. †Alloterocucus Li et al. 2022
Fm. Cryptophagidae Kirby 1837 [silken fungus beetle]
+
Invalid names: Lebanophytidae Kirejtshuk 2013 [synonym]
Fm. Cucujidae Latreille 1802
Fm. Cyclaxyridae Gimmel et al. 2009
Fm. Hobartiidae Sen Gupta and Crowson 1966
Subfm. †Meligethiellinae Kirejtshuk and Ponomarenko 1990
Fm. Passandridae Blanchard 1845 [parasitic flat bark beetle]
Fm. Phalacridae Leach 1815
Fm. Phloeostichidae Reitter 1911
Fm. Silvanidae Kirby 1837
Invalid names: Agapythidae Sen Gupta and Crowson 1969 [empty], Brachypteridae Erichson 1845 [empty], Cavognathidae Sen Gupta and Crowson 1966 [empty], Clavicornia [invalid subgroup], Heteromera [invalid subgroup], Lamingtoniidae Sen Gupta and Crowson 1969 [empty], Myraboliidae Lawrence and Britton 1991 [empty], Priasilphidae Crowson 1973 [empty], Sinisilvanidae Hong 2002 [invalid subgroup], Tasmosalpingidae Lawrence and Britton 1991 [empty]
Superfm. Erotyloidea Latreille 1802
+
Fm. Boganiidae Sen Gupta and Crowson 1966
Fm. Erotylidae Latreille 1802 [pleasing fungus beetle]
G. †Isocryptophilus Li and Cai 2024
Superfm. Nitiduloidea Latreille 1802
+
Fm. Helotidae Reitter 1876
Fm. Kateretidae Kirby 1837
Fm. Monotomidae Laporte 1840 [root eating beetle]
+
Invalid names: Rhizophagidae Redtenbacher 1845 [synonym]
Fm. Nitidulidae Latreille 1802 [sap beetle]
Fm. Protocucujidae Crowson 1954
Fm. Smicripidae Horn 1879 [palmetto beetle]
Fm. Sphindidae Jacquelin du Val 1860 [cryptic slime mold beetle]
+
Invalid names: Aspidiphoridae Kiesenwetter 1877 [synonym]
Fm. †Parandrexidae Kirejtshuk 1994
+
Subfm. †Cretoparacucujinae Kirejtshuk 2023
Subfm. †Parandrexinae Kirejtshuk 1994
Superfm. Tenebrionoidea Latreille 1802
+
Fm. Aderidae Csiki 1909
+
Invalid names: Circaeidae Iablokoff-Khnzorian 1961 [synonym], Xylophilidae [replaced]
Fm. Anthicidae Latreille 1819 [ant like flower beetle]
G. †Archaeoripiphorus Hsiao et al. 2017
Fm. Boridae Thomson 1859
Fm. Ciidae Leach 1819 [minute tree-fungus beetle]
Fm. †Elodophthalmidae Kirejtshuk and Azar 2008
G. †Glypta Hong 1984
Subfm. Ischaliidae Blair 1920 [false fire-colored beetle]
G. †Kulindrobor Tihelka et al. 2021
Fm. †Liaoximordellidae Wang 1993
Fm. Melandryidae Leach 1815 [false darkling beetle]
Fm. Meloidae Gyllenhal 1810 [blister beetle]
Fm. Mordellidae Latreille 1802 [tumbling flower beetle]
+
Invalid names: Praemordellidae Shchegoleva-Barovskaya 1929 [synonym]
Fm. Mycetophagidae Leach 1815 [hairy fungus beetle]
Fm. Mycteridae Oken 1843 [palm and flower beetle]
G. †Neopolypria Abdullah 1964
Fm. Oedemeridae Latreille 1810 [false blister beetle]
Fm. Prostomidae Thomson 1859 [jugular-horned beetle]
Fm. Pyrochroidae Latreille 1807
Fm. Pythidae Solier 1834
Fm. Rhipiphoridae Gemminger 1870 [wedge shaped beetle]
Fm. Ripiphoridae Gemminger 1870 [wedge shaped beetle]
Fm. Salpingidae Leach 1815 [narrow-waisted bark beetle]
G. †Samlandotoma Alekseev 2019
Fm. Scraptiidae Gistel 1848 [false flower beetle]
Fm. Stenotrachelidae Thomson 1859
Fm. Synchroidae Lacordaire 1859
Fm. Tenebrionidae Latreille 1802 [darkling beetle]
Fm. Tetratomidae Billberg 1820 [polypore fungus beetle]
Fm. Ulodidae Pascoe 1869
Invalid names: Afreminae Levey 1985 [empty], Archeocrypticidae Kaszab 1964 [empty], Asiamordella Hong 2002 [nomen nudum], Chalcodryidae Watt 1974 [empty], Cossyphodinae Wasmann 1899 [empty], Eurygenius wickhami Cockerell 1917 [nomen dubium], Fushunicucujus Hong 2002 [nomen nudum], Lagrioididae Abdullah and Abdullah 1968 [empty], Perimylopidae St. George 1939 [empty], Pterogeniidae Crowson 1953 [empty], Trachelostenidae Lacordaire 1859 [empty], Trictenotomidae Blanchard 1845 [empty]
Infraor. Elateriformia Crowson 1960
+
Superfm. Buprestoidea Leach 1815
+
Fm. Buprestidae Leach 1815 [jewel beetle]
Fm. Schizopodidae LeConte 1859
Invalid names: Mesohelophorus mongolicus Ponomarenko 1986 [nomen dubium]
Superfm. Byrrhoidea Latreille 1804
+
Fm. Byrrhidae Latreille 1804 [pill beetle]
G. †Serecoleus Yan et al. 2015
Invalid names: Chelonarium montanum Wickham 1914 [nomen dubium]
Superfm. Dascilloidea Gue'rin-Me'neville 1843
+
Fm. Dascillidae Guerin-Meneville 1843 [soft bodied plant beetle]
Fm. Rhipiceridae Latreille 1834 [cicada parasite beetle]
Invalid names: Karumiidae Escalera 1913 [empty]
Superfm. Dryopoidea
+
Fm. Chelonariidae Blanchard 1845
Fm. Dryopidae Billberg 1820 [long toed water beetle]
+
Invalid names: Parnidae Leach 1817 [replaced]
Fm. Elmidae Curtis 1830 [riffle beetle]
Fm. Eulichadidae Crowson 1973 [forest stream beetle]
Fm. Heteroceridae MacLeay 1825 [Variegated mud-loving beetle]
G. †Jiuquanocoleus Hong 1982
Fm. Limnichidae Erichson 1846 [minute marsh loving beetle]
Fm. Lutrochidae Kasap and Crowson 1975 [robust marsh loving beetle]
Fm. †Mastigocoleidae Tihelka et al. 2022
Fm. Psephenidae Lacordaire 1854 [water penny beetle]
Fm. Ptilodactylidae Laporte 1836
Invalid names: Actinocerites Hong 2002 [nomen nudum], Brachypodites Hong 2002 [nomen nudum], Callirhipidae Emden 1924 [empty], Cneoglossidae Champion 1897 [empty], Expansifemoria Hong 2002 [nomen nudum]
Superfm. Elateroidea Leach 1815
+
G. †Anoeuma Li et al. 2021
Fm. Artematopodidae Lacordaire 1857 [soft bodied plant beetle]
+
Invalid names: Eurypogonidae [replaced]
Fm. †Berendtimiridae Winkler 1987
Fm. Brachypsectridae LeConte and Horn 1883 [Texas beetle]
Fm. Cantharidae Imhoff 1856 [soldier beetle]
+
Invalid names: Cantharidiae Latreille 1802 [replaced], Telephoridae [replaced]
Fm. †Cretophengodidae Li et al. 2021
Fm. Eucnemidae Eschscholtz 1829 [false click beetle]
G. †Grammocolous Lin 1986
Fm. Lampyridae Rafinesque 1815
Fm. Lycidae Laporte 1836 [net-winged beetle]
G. †Microcoleus Hong 1982
Fm. †Mysteriomorphidae Alekseev and Ellenberger 2019
Fm. Omethidae Leconte 1861
Fm. Phengodidae LeConte 1861
Invalid names: Artematopoidea Lacordaire 1857 [synonym], Eudicronychinae Girard 1971 [empty], Rhagophthalmidae Olivier 1907 [empty]
Invalid names: Mesosperchus angulatus Ponomarenko 1985 [nomen dubium], Podabrocephalidae Pic 1930 [empty]
G. †Jingxidiscus Hong 1990
+
†Jingxidiscus lushangfenensis Hong 1981
Invalid names: Discus Hong 1981 [replaced]
Fm. Jurodidae Ponomarenko 1985
+
G. †Jurodes Ponomarenko 1985
+
†Jurodes daohugouensis Yan et al. 2014
†Jurodes ignoramus Ponomarenko 1985
†Jurodes minor Ponomarenko 1990
†Jurodes pygmaeus Yan et al. 2014
†Jurodes shartegiensis Yan 2014
†Jurodes shef Yan et al. 2021
Invalid names: Sikhotealiniidae Lafer 1996 [synonym]
G. †Larvula Ponomarenko 1985
+
†Larvula cassa Ponomarenko 1985
†Larvula triassica Prokin and Ponomarenko 2013
Fm. †Lasiosynidae Kirejtshuk et al. 2010
+
G. †Anacapitis Yan 2009
+
†Anacapitis incertus Yan 2009
†Anacapitis karataviensis Yan 2009
†Anacapitis oblongus Yan 2009
†Anacapitis plata Tan and Ren 2009
Invalid names: Brachysyne Tan and Ren 2009 [synonym]
G. †Glaphyropterodes Handlirsch 1906
+
†Glaphyropterodes gehreti Heer 1852
†Glaphyropterodes sibiricus Ponomarenko 1985
G. †Lasiosyne Tan et al. 2007
+
†Lasiosyne cataphracta Yan 2012
†Lasiosyne daohugouensis Kirejtshuk et al. 2010
†Lasiosyne decora Yan 2012
†Lasiosyne euglyphea Tan et al. 2007
+
Invalid names: Pappisyne eucallus Tan and Ren 2009 [synonym], Pappisyne lasiospatha Tan and Ren 2009 [synonym]
†Lasiosyne fedorenkoi Kirejtshuk et al. 2010
†Lasiosyne gratiosa Kirejtshuk et al. 2010
†Lasiosyne insculpta Yan 2012
†Lasiosyne integera Guo et al. 2022
†Lasiosyne longitarsus Yan 2012
†Lasiosyne multituberata Lin 1980
†Lasiosyne parva Guo et al. 2022
†Lasiosyne quadricollis Kirejtshuk et al. 2010
Invalid names: Pappisyne Tan and Ren 2009 [synonym]
Invalid names: Lasiosyne laxa Yan et al. 2014 [nomen dubium]
Unr. Nosodendriformia Cai and Tihelka 2022
+
G. †Plastelater Handlirsch 1906
+
†Plastelater glabratus Ponomarenko 1985
†Plastelater neptuni Giebel 1856
G. †Polysitum Dunstan 1923
+
†Polysitum elongatum Ponomarenko 1985
†Polysitum hosbayari Ponomarenko 1986
†Polysitum kuznetskiense Rohdendorf 1961
†Polysitum latum Ponomarenko 1985
†Polysitum majus Ponomarenko 1990
†Polysitum minutus Dunstan 1923
†Polysitum ovale Ponomarenko 1990
†Polysitum punctatus Dunstan 1923
†Polysitum rugosum Ponomarenko 1986
†Polysitum sharategense Ponomarenko 2014
†Polysitum wudenghaoensis Hong 1976
Unr. Rhinorhipiformia Cai et al. 2022
+
Unr. Scirtiformia Lawrence et al. 2011
+
Superfm. Scirtoidea Fleming 1821
+
Fm. †Boleopsidae Kirejtshuk and Nel 2013
Invalid names: Decliniidae Nikitsky et al. 1994 [empty]
G. †Sphaerocantharis Bode 1953
+
†Sphaerocantharis defossa Bode 1953
†Sphaerocantharis sibirica Ponomarenko 1990
Invalid names: Mikrocarpides Bode 1953 [synonym], Theornithion Bode 1953 [synonym]
Infraor. Staphyliniformia Lameere 1900
+
Superfm. Histeroidea Gyllenhal 1808
+
Fm. †Cretohisteridae Zhou et al. 2018
Fm. Histeridae Gyllenhal 1808 [clown beetle]
Fm. Synteliidae Lewis 1882
Invalid names: Sphaeritidae Shuckard 1839 [empty]
Superfm. Hydrophiloidea Latreille 1802 [water scavenger beetle]
+
G. †Cretotaenia Ponomarenko 1977
Fm. Helophoridae Leach 1815
Fm. Hydrochidae Thomson 1859
Fm. Hydrophilidae Latreille 1802 [water scavenger beetle]
+
Invalid names: Escheriidae Kolbe 1880 [synonym]
G. †Hydrophilopsia Ponomarenko 1987
G. †Laetopsia Fikáček et al. 2012
G. †Sinosperchopsis Prokin et al. 2010
Fm. Spercheidae Erichson 1837
Invalid names: Epimetopidae Zaitzev 1908 [empty], Georissidae Laporte 1819 [empty]
Superfm. Scarabaeoidea Latreille 1802 [scarab beetle]
+
G. †Aliscarabaeus Hong 1984
Fm. †Alloioscarabaeidae Bai et al. 2012
G. †Ateuchites Meunier 1898
Fm. Bolboceratidae Mulsant 1842 [dor beetle]
G. †Ceafornotensis Woolley 2016
G. †Crassiungula Li et al. 2025
G. †Cretaclopus Nikolajev 2004
G. †Cretonitis Nikolajev 2007
Fm. Diphyllostomatidae Holloway 1972
Fm. Geotrupidae Latreille 1802 [earth-boring dung beetle]
G. †Geotrupoides Handlirsch 1906
Fm. Glaphyridae MacLeay 1819 [bumble bee scarab beetle]
Fm. Glaresidae Prudhomme de Borre 1886 [enigmatic scarab beetle]
G. †Holcorobeus Nikritin 1977
Fm. Hybosoridae Erichson 1847 [scavenger scarab beetle]
G. †Hybosorites Nikolajev 1996
G. †Juraclopus Nikolajev 2004
Fm. Lucanidae Latreille 1804 [stag beetle]
G. †Mahengea Strümpher et al. 2023
G. †Mesaclopus Nikolajev 1992
G. †Mesoscarabaeus Hong 1982
G. †Mongolrobeus Nikolajev 2004
Fm. Ochodaeidae Streubel 1846 [sand-loving scarab beetle]
G. †Opiselleipon Bode 1953
Fm. Passalidae Leach 1815 [bess beetle]
Fm. †Passalopalpidae Boucher and Bai 2016
Fm. Pleocomidae Leconte 1861 [rain beetle]
G. †Prionocephale Lin 1980
G. †Prophaenognatha Bai et al. 2011
Fm. Scarabaeidae Latreille 1802 [scarab beetle]
G. †Scelocopris Zhang 1989
Fm. †Septiventeridae Bai et al. 2013
Fm. Trogidae MacLeay 1819 [hide beetle]
Invalid names: Belohinidae Paulian 1959 [empty], Copris subterraneus Heer 1862 [nomen dubium], Gymnopleurus deperditus Heer 1862 [nomen dubium], Holcorobeini Nikolajev 1992 [invalid subgroup], Onthophagus urusheeri Krell 2000 [nomen dubium]
Superfm. Staphylinoidea Latreille 1802
+
Fm. Agyrtidae Thomson 1859 [primitive carrion beetle]
Fm. Colonidae Horn 1880
Fm. Hydraenidae Mulsant 1844 [minute moss beetle]
Fm. Leiodidae Fleming 1821 [round fungus beetle]
Fm. Ptiliidae Erichson 1845 [feather-winged beetle]
Invalid names: Dasyceridae Reitter 1887 [empty], Empelidae [empty]
Invalid names: Scarabaeiformia Crowson 1960 [synonym]
Invalid names: Agyrtes primoticus Scudder 1900 [nomen dubium], Derodontiformia Le Conte 1861 [invalid subgroup], Escheria crassipunctata Förster 1891 [nomen dubium], Escheria dimidiata Förster 1891 [nomen dubium], Escheria punctulata Förster 1891 [nomen dubium], Eucinetiformia [empty], Limnius perrini Theobald 1937 [nomen dubium], Malachius vertummni Heer 1847 [nomen dubium], Mesosperchus notatus Ponomarenko 1977 [nomen dubium], Tomocoleites Hong 2002 [nomen nudum], Troglops punctatissima Théobald 1935 [nomen dubium]
Fm. †Ponomarenkiumidae Lawrence and Bouchard 2022
+
G. †Ponomarenkium Yan et al. 2018
+
†Ponomarenkium belmonthense Yan et al. 2017
Invalid names: Ponomarenkia Yan et al. 2017 [replaced]
Invalid names: Ponomarenkiidae Yan et al. 2017 [invalid subgroup]
Fm. †Taldycupedidae Rohdendorf 1961
+
G. †Afrotaldycupes Kirejtshuk 2020
+
†Afrotaldycupes africanus Ponomarenko 2005
†Afrotaldycupes lidgettoniensis Ponomarenko 2005
G. †Mesothoris Tillyard 1916
+
†Mesothoris clathrata Tillyard 1916
†Mesothoris grandis Dunstan 1923
†Mesothoris punctomarginum Dunstan 1923
G. †Proterocupes Ponomarenko 2015
+
†Proterocupes major Ponomarenko 2015
†Proterocupes nedubrovensis Ponomarenko 2015
†Proterocupes permiana Ponomarenko 2013
Invalid names: Lobanovia Kirejtshuk 2020 [objective synonym]
G. †Simmondsia Dunstan 1923
+
†Simmondsia cylindrica Dunstan 1923
†Simmondsia subpiriformis Dunstan 1923
G. †Taldycupes Rohdendorf 1961
+
†Taldycupes cellulosus Ponomarenko 2013
†Taldycupes khalfini Rohdendorf 1961
†Taldycupes moltshanovi Rohdendorf 1961
†Taldycupes pingi Ponomarenko et al. 2014
†Taldycupes reticulatus Ponomarenko 1969
†Taldycupes rosanovi Ponomarenko 2006
†Taldycupes vasjuchitshevi Rohdendorf 1961
Invalid names: Cryptocupes Rohdendorf 1961 [synonym], Taldycupidium Rohdendorf 1961 [synonym]
G. †Tecticupes Rohdendorf 1961
+
†Tecticupes heckeri Rohdendorf 1961
+
Invalid names: Stegocupes efremovi Rohdendorf 1961 [synonym], Stegocupes fedotovi Rohdendorf 1961 [synonym]
†Tecticupes indistinctus Rohdendorf 1961
+
Invalid names: Tychticupes tchihatcheffi Rohdendorf 1961 [synonym]
†Tecticupes martynovi Ponomarenko 2013
Invalid names: Stegocupes Rohdendorf 1961 [synonym]
G. †Tychticupes Rohdendorf 1961
+
†Tychticupes beljanini Rohdendorf 1961
†Tychticupes radtshenkoi Rohdendorf 1961
†Tychticupes ragozini Rohdendorf 1961
Invalid names: Tychticupes ussovi Rohdendorf 1961 [nomen dubium], Uskaticupes Rohdendorf 1961 [nomen dubium], Uskaticupes javorskyi Rohdendorf 1961 [nomen dubium]
Fm. †Triadocupedidae Ponomarenko 1966
+
G. †Cupesia Ponomarenko 1966
+
†Cupesia monilicornia Ponomarenko 1966
†Cupesia sepulta Ponomarenko 1969
†Cupesia serricornia Ponomarenko 1969
G. †Platycupes Ponomarenko 1966
+
†Platycupes dolichocerus Ponomarenko 1966
†Platycupes major Ponomarenko 1966
†Platycupes pusillus Ponomarenko 1966
†Platycupes reticulatus Ponomarenko 1966
G. †Triadocupes Ponomarenko 1966
+
†Triadocupes ellipticus Ponomarenko 1966
†Triadocupes ferghanensis Ponomarenko 1966
†Triadocupes latus Ponomarenko 1966
Fm. †Permocupedidae Martynov 1932
hide
Subfm. †Permocupedinae Martynov 1932
+
G. †Afrocupes Geertsema and van den Heever 1996
+
†Afrocupes firmae Geertsema and van den Heever 1996
G. †Archicupes Rohdendorf 1961
+
†Archicupes jacobsoni Rohdendorf 1961
+
Invalid names: Palaeocupes kaltanicus Rohdendorf 1961 [synonym]
Invalid names: Palaeocupes Rohdendorf 1961 [synonym]
G. †Eocupes Rohdendorf 1961
+
†Eocupes cellulosus Ponomarenko 1969
†Eocupes lukjanovitshi Rohdendorf 1961
G. †Frankencupes Ponomarenko and Bashkuev 2018
+
†Frankencupes ultimus Ponomarenko and Bashkuev 2018
G. †Ichthyocupes Rohdendorf 1961
+
†Ichthyocupes kuznetskiensis Ponomarenko 1969
†Ichthyocupes skoki Rohdendorf 1961
†Ichthyocupes tyzhnovi Rohdendorf 1961
G. †Kaltanicupes Rohdendorf 1961
+
†Kaltanicupes acutus Ponomarenko 1963
†Kaltanicupes kargalensis Ponomarenko 1963
†Kaltanicupes kitjakensis Ponomarenko 1963
†Kaltanicupes richteri Rohdendorf 1961
G. †Permocupes Martynov 1932
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†Permocupes distinctus Martynov 1932
†Permocupes grandis Ponomarenko 1963
†Permocupes latus Ponomarenko 2013
†Permocupes semenovi Martynov 1932
†Permocupes sojanensis Ponomarenko 1963
Invalid names: Permocupoides Martynov 1932 [synonym]
G. †Protocupoides Rohdendorf 1961
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†Protocupoides acer Rohdendorf 1961
†Protocupoides carinatus Rohdendorf 1961
†Protocupoides elongatus Ponomarenko 2013
†Protocupoides esini Ponomarenko 2013
†Protocupoides plavilstshikovi Rohdendorf 1961
†Protocupoides sharovi Rohdendorf 1961
Invalid names: Protocupoides rohdendorfi Pinto 1987 [nomen dubium], Tomiocupes Rohdendorf 1961 [synonym], Tricupes Rohdendorf 1961 [synonym]
Invalid names: Protocupes Rohdendorf 1961 [nomen dubium], Taldycupes chachlovi Rohdendorf 1961 [nomen dubium]
Invalid names: Archicupes reichardti Rohdendorf 1961 [nomen dubium], Kaltanocoleidae Rohdendorf 1961 [synonym], Protocupes martynovi Rohdendorf 1961 [nomen dubium]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Bush and Bambach 2015, Kiessling 2004 | |||||
Age range
Maximum range based only on fossils: base of the Asselian to the top of the Holocene or 298.90000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 293.52 Ma
Minimum age of oldest fossil (stem group age): 293.52 Ma
Collections (1849 total)
Oldest occurrences | |||
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Asselian | Czechia | Moravocoleus perditus (114186) | |
| Artinskian | South Africa (Western Cape) | Afrocupes firmae (156986) | |
| Kungurian | Russian Federation | Maricoleus valentinae (115214) | |
| Kungurian | Brazil (Rio Grande do Sul) | Protocupoides rohdendorfi, Kaltanicupes ponomarenkoi (116121) | |
| Roadian | Russian Federation (Kazan) | Permocupoides distinctus, Permocupes semenovi (116877) | |
| Roadian | Russian Federation (Kemerovo) | Eocupes lukjanovitshi, Cytocupoides elongatus, Archicupes jacobsoni, Protocupes martynovi, Rossocoleus avus (122567) Ichthyocupes kuznetskiensis (122572) Ichthyocupes tyzhnovi, Cytocupes angustus, Protocupoides plavilstshikovi, Tomiocupes sharovi, Eocupes cellulosus, Kaltanocoleus pospelovi, Palaeocupes kaltanicus, Archicupes reichardti, Stegocupes efremovi, Schizotaldycupes ananjevi, Carinicupes beckermigdisovae, Asiocoleus novojilovi, Uskatocoleus kaltanicus (104704) Kaltanicupes richteri (122568) Kaltanocoleus pospelovi, Protocupes martynovi (122569) Schizocoleus kuznetskiensis (124604) | |
| Roadian | Russian Federation | Stegosyne rugosa (124611) | |
| Roadian | Russian Federation (Arkangelsk) | Permocupes semenovi (104716) Permocupes sojanensis (104992) Permocupes sojanensis, Permocupes grandis (116933) | |
| Roadian | Russian Federation (Kirov) | Kaltanicupes major, Kaltanicupes kitjakensis, Kaltanicupes acutus, Rossocoleus altus (105058) | |
| Wordian | Russian Federation (Udmurtia) | Protocupoides elongatus, Permocupes latus, Rossocoleus novojilovi, Rossocoleus angustus, Schizophoridae indet. (103502) Schizotaldycupes pubescens, Rhombocoleites uralensis (104750) | |
| Wordian | South Africa | Permocupedidae indet. (226396) | |
| Wordian | Russian Federation (Kemerovo) | Protocupes ogloblini, Taldycupes chachlovi, Tecticupes indistinctus, Tychticupes tchihatcheffi, Tecticupes heckeri, Stegocupes fedotovi, Rossocoleus latior, Rossocoleus surijokovensis, Rossocoleus latus, Rossocoleus grandis, Rossocoleus parvus, Karakanocoleus janischewskyi, Uskatocoleus surijokovicus, Uskatocoleus iljinskiensis, Uskatocoleus avitus (122566) Rossocoleus verrucosus, Uskatocoleus convexus (124603) Schizocupes obrutshevi (124600) | |
| Wordian - Capitanian | Russian Federation | Uskatocoleus kushejakovicus (124610) | |
| Capitanian | Russian Federation (Orenburg) | Curculiopsis ellipticus (104994) Kaltanicupes kargalensis, Tecticupes martynovi, Rhombocoleites adumbratus, Kargalarva permosialis (104057) Rossocoleus sakmara, Schizocoleus major, Aenigmocoleus uralensis (152235) Tatarocupes granulatus, Bicoleus kuplensis, Tetracoleus orenburgensis, Rhombocoleus gomankovi, Palademosyne ovum, Schizocoleus longus, Schizocoleus robustus, Schizocoleus glabrus, Schizocoleus minimus, Erunakicupes brevis, Karakanocoleus europeus (104717) | |
| Capitanian | China (Anhui) | Linicupes yinpinensis, Taldycupes pingi, Asiocoleopsis hongi, Chaocoleus limnebius, Dikerocoleus divisus (103239) | |
| Wuchiapingian | Russian Federation | Permocupoides skoki, Tricupes acer, Tomiocupes carinatus, Tychticupoides grjazevi, Taldycupidium bergi, Cryptocupes rjabinini, Taldycupes moltshanovi, Tychticupes vasjuchitshevi, Ademosyne sibirica, Polysitum kuznetskiense, Rhombocoleus andreae, Aenigmocoleus borissiaki, Uskatocoleus angustior, Uskatocoleus angustus, Uskatocoleus longus, Erunakicupes kryshtofovichi, Erunakicupes schmalhauseni, Erunakicupes fomitshevi, Karakanocoleus major, Karakanocoleodes latissimus, Erunakicupes venjukovi, Karakanocoleus lebedevi, Karakanocoleus magnus, Karakanocoleus striatus, Uskatocoleus lutugini, Uskatocoleus minimus, Uskatocoleus foveolatus, Uskatocoleus zalesskyi (124608) Permocupoides skoki, Tychticupes ussovi, Uskaticupes javorskyi, Taldycupes khalfini, Tychticupes ragozini, Tychticupes radtshenkoi, Taldycupes beljanini, Uskatocoleus zalesskyi (122565) | |
| Wuchiapingian | Russian Federation (Vologda) | Protocupoides esini, Taldycupedidae indet., Simmondsia permiana, Taldycupes cellulosus, Bicoleus laticella, Tetracoleus tricoleoides, Rossocoleus aristovi, Pseudochrysomelites bashkuevi, Pseudochrysomelites medialis, Pseudochrysomelites longus, Pseudochrysomelites sphenoidalis, Schizocoleus depressus, Erunakicupes grossus, Karakanocoleus mutovinensis, Uskatocoleus micron, Uskatocoleus convexus, Uskatocoleus sukhonensis (104754) | |
| Wuchiapingian | Russian Federation (Orenburg) | Uskatocoleus uralensis (152236) | |
| Changhsingian | Russian Federation (Vladimir) | Schizocoleidae indet., Pseudochrysomelites major, Metrorhynchites baculum, Metrorhynchites elongatus, Erunakicupes angustus, Uskatocoleus eurypygus, Uskatocoleus rhynchophorus (113351) Tetracoleus sennikovi (104756) | |
| Changhsingian | South Africa (Kwazulu-Natal) | Taldycupes africanus (14356) Taldycupes lidgettoniensis, Rhombocoleites danutae (14357) | |