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Metacoleoptera
Taxonomy
Metacoleoptera was named by Boudinot et al. (2022). It is extant.
It was assigned to Mesocoleoptera by Boudinot et al. (2022).
It was assigned to Mesocoleoptera by Boudinot et al. (2022).
Synonymy list
| Year | Name and author |
|---|---|
| 2022 | Metacoleoptera Boudinot et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Unr. Metacoleoptera Boudinot et al. 2022
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Fm. †Ademosynidae Ponomarenko 1968
+
G. †Ademosyne Handlirsch 1906
+
†Ademosyne adunca Dunstan 1923
†Ademosyne arcucciae Martins-Neto and Gallego 2006
†Ademosyne australiensis Tillyard 1916
†Ademosyne bacca Ponomarenko 1969
†Ademosyne brevis Dunstan 1923
†Ademosyne cameroni Tillyard 1916
†Ademosyne congener Tillyard 1916
†Ademosyne curvata Dunstan 1923
†Ademosyne elliptica Ponomarenko 1969
†Ademosyne elongatus Martins-Neto and Gallego 2006
†Ademosyne hexacostata Martins-Neto and Gallego 2006
†Ademosyne intermedia Dunstan 1923
†Ademosyne kirghizica Ponomarenko 1969
†Ademosyne lata Dunstan 1923
†Ademosyne llantenesensis Brauckmann et al. 2010
†Ademosyne major Handlirsch 1906
†Ademosyne olliffi Handlirsch 1906
†Ademosyne parva Dunstan 1923
†Ademosyne polyzetete Ponomarenko 2011
†Ademosyne prisca Riek 1974
†Ademosyne punctata Tillyard 1916
†Ademosyne punctuada Martins-Neto and Gallego 2006
†Ademosyne ramocostata Dunstan 1923
†Ademosyne reducta Riek 1976
†Ademosyne rosenfeldi Brauckmann et al. 2010
†Ademosyne rugulosa Dunstan 1923
†Ademosyne sibirica Rohdendorf 1961
†Ademosyne speciosa Riek 1974
†Ademosyne umutu Martins-Neto and Gallego 2009
†Ademosyne vittamargina Dunstan 1923
†Ademosyne wianamattensis Tillyard 1918
G. †Dolichosyne Ponomarenko 1969
+
†Dolichosyne confragosa Ponomarenko 1969
†Dolichosyne rostrata Ponomarenko 1969
†Dolichosyne sulcata Ponomarenko 1969
Subor. Adephaga Clairville 1806
+
G. †Amphoxyne Bode 1953
+
†Amphoxyne darbiensis Ponomarenko 1986
†Amphoxyne minuta Bode 1953
+
Invalid names: Coilotrachelus lineatus Bode 1953 [synonym], Cricotrachelites rotundatus Bode 1953 [synonym], Loxoncus procerus Bode 1953 [synonym]
Invalid names: Coilotrachelus Bode 1953 [synonym], Cricotrachelites Bode 1953 [synonym], Loxostelidotus Bode 1953 [synonym]
Fm. Carabidae Latreille 1802 [ground beetle]
+
G. †Antemnacrassa Gomez Pallerola 1979
+
†Antemnacrassa albomaculata Nikolajev 2004
†Antemnacrassa albosulcata Nikolajev 2004
†Antemnacrassa geminata Nikolajev 2004
†Antemnacrassa incerta Ponomarenko 1990
†Antemnacrassa kazakhstanica Nikolajev 2007
†Antemnacrassa latitibialis Nikolajev 1992
†Antemnacrassa maculata Nikolajev 1992
†Antemnacrassa magna Nikolajev 2004
†Antemnacrassa monreali Gomez Pallerola 1979
†Antemnacrassa nebulosa Nikolajev 2004
†Antemnacrassa nigrimontana Nikolajev 1992
†Antemnacrassa picturata Nikritin 1977
†Antemnacrassa punctata Ponomarenko 1986
†Antemnacrassa sulcata Nikritin 1977
†Antemnacrassa vitimensis Nikritin 1977
G. Apotomopterus Hope 1838
G. Bembidium Latreille 1802
+
Subg. Bembidium (Campa) Motschulsky 1844
+
†Bembidium (Campa) boryslavicum Lomnicki 1894
†Bembidium (Campa) subcontaminatum Lomnicki 1894
Subg. Bembidium (Testidium) Motschulsky 1864
†Bembidium absolutum Giebel 1856
†Bembidium cyaneomicans Piton 1940
†Bembidium exoletum Scudder 1876
†Bembidium expletum Scudder 1900
†Bembidium florissantensis Wickham 1913
†Bembidium fragmentum Scudder 1890
†Bembidium glaciatum Scudder 1890
†Bembidium haywardi Scudder 1900
†Bembidium infernum Heer 1856
†Bembidium laevigatum Förster 1891
†Bembidium obductum Scudder 1900
†Bembidium praeteritum Scudder 1900
†Bembidium saportanum Oustalet 1874
†Bembidium tumulorum Scudder 1900
†Bembidium vanum Scudder 1900
†Bembidium vestigium Scudder 1900
Subfm. Broscinae Hope 1838
+
Subfm. Carabinae Latreille 1802
+
G. †Atrirabus Hong and Wang 1990
+
†Atrirabus shandongensis Hong and Wang 1990
†Atrirabus tuanwangensis Hong and Wang 1990
Tr. Carabini Latreille 1802
+
G. Calosoma Weber 1801
G. Carabus Linnaeus 1758
Invalid names: Calosomini Bonelli 1810 [synonym]
Tr. Cychrini Perty 1830
+
G. Cychrus Fabricius 1794
G. Scaphinotus Dejean 1826
G. Sphaeroderus Dejean 1831 [small snail eating beetle]
G. †Carabites Ponomarenko 1977
+
†Carabites caraboides Ponomarenko 1989
†Carabites nigriventris Ponomarenko 1986
G. †Carabites Heer 1852
+
†Carabites anthracinus Heer 1852
†Carabites arapahoensis Cockerell 1918
†Carabites capellinii Cecconi 1892
†Carabites creta Ponomarenko 1977
†Carabites cuneatus Cockerell 1920
†Carabites dubius Geinitz 1894
†Carabites ellipticus Haupt 1956
†Carabites eocenicus Cockerell 1921
†Carabites exanimus Scudder 1892
†Carabites exilis Heer 1862
†Carabites feildenianus Heer 1878
†Carabites gardneri Cockerell 1920
†Carabites geinitzi Handlirsch 1906
†Carabites harringtoni Cockerell 1936
†Carabites islandicus Heer 1868
†Carabites kincaidi Cockerell 1908
†Carabites laesicollis Heer 1862
†Carabites parallelus Cockerell 1920
†Carabites peracutus Cockerell 1920
†Carabites rugosus Meunier 1898
†Carabites russelli Cockerell 1928
†Carabites schueli Cockerell 1926
†Carabites scoticus Cockerell 1921
†Carabites vitimensis Ponomarenko 1977
G. †Carabopteron Martynov 1926
+
†Carabopteron punctatolineatum Martynov 1926
†Carabopteron rugosum Ponomarenko 1985
Unr. Celeripedes Latreille 1802
+
Invalid names: Barbati Latreille 1802 [empty], Crepitantes Latreille 1802 [empty], Longipalpati Latreille 1802 [empty], Melanchlaeni Latreille 1802 [empty], Metallici Latreille 1802 [empty]
Subfm. †Conjunctiinae Ponomarenko 1977
+
G. †Conjunctia Ponomarenko 1977
+
†Conjunctia longa Zhang 1997
†Conjunctia prodroma Ponomarenko 1977
Subfm. Elaphrinae Latreille 1802
+
G. Blethisa Bonelli 1810
+
Blethisa catenaria Brown 1944
Blethisa inexpectata Goulet and Smetana 1983
Blethisa multipunctata Linnaeus 1758
†Blethisa pleistocenica Lomnicki 1894
Invalid names: Helobium Leach 1815 [objective synonym]
G. Diacheila Motschulsky 1844
+
Diacheila arctica Gyllenhal 1810
†Diacheila matthewsi Böcher 1995
Diacheila polita Faldermann 1835
G. Elaphrus Fabricius 1775
+
Subg. Elaphrus (Neoelaphrus) Hatch 1951
Elaphrus americanus Dejean 1831
Elaphrus angusticollis Sahlberg 1844
Elaphrus californicus Mannerheim 1843
Elaphrus cupreus Duftschmid 1812
Elaphrus finitimus Casey 1920
Elaphrus japonicus Ueno 1954
Elaphrus lapponicus Gyllenhal 1810
Elaphrus lecontei Crotch 1876
†Elaphrus pliocenicus Piton 1939
Elaphrus purpurans Hausen 1891
Elaphrus riparius Linnaeus 1758
Elaphrus ruscarius Say 1834
Elaphrus sibiricus Motschulsky 1846
Elaphrus tuberculatus Maklin 1877
Invalid names: Elaphrus irregularis Scudder 1890 [nomen dubium], Elaphrus ruscarius foveatus Pierce 1948 [synonym]
G. †Fonsecacarabus Martins-Neto and Mendes 2002
+
†Fonsecacarabus placidus Martins-Neto and Mendes 2002
Subfm. Gehringiinae Darlington 1933
+
G. †Hadrocephalus Handlirsch 1906
+
†Hadrocephalus anglicus Handlirsch 1906
†Hadrocephalus liasinus Handlirsch 1906
†Hadrocephalus minor Handlirsch 1906
Subfm. Harpalinae Bonelli 1810
+
Tr. Lebiini Bonelli 1810
+
G. †Agatoides Motschulsky 1856
Subtr. Calleidina Chaudoir 1873
Subtr. Cymindidina Laporte 1834
G. †Cymindoides Motschulsky 1856
Subtr. Dromiusina Bonelli 1810
Subtr. Lebiina Bonelli 1810
G. †Miolebidia Zhang 1989
Subtr. Pericalina Hope 1838
G. †Protoscalidion Schaufuss 1889
Invalid names: Brachinus repressus Scudder 1900 [nomen dubium]
Tr. Sphodrini Laporte 1834
+
Subtr. Calathina Laporte 1834
Subtr. Sphodrina Laporte 1834
Subtr. Synuchina Lindroth 1956
Invalid names: Dryptini Bonelli 1810 [empty], Licininae Bonelli 1810 [synonym]
Subfm. Migadopinae Chaudoir 1861
+
Subfm. Nebriinae Laporte 1834
+
Tr. Nebriini Laporte 1834
+
G. †Archaeonebria Kavanaugh and Schmidt 2019
G. Leistus Frölich 1799
G. Nebria Latreille 1802
G. †Nebrorabus Ponomarenko 1989
+
†Nebrorabus baculum Ponomarenko 1989
†Nebrorabus capitatus Ponomarenko 1989
†Nebrorabus nebrioides Ponomarenko 1989
Subfm. Omophroninae Bonelli 1810
+
Subfm. Paussinae Latreille 1806 [ant nest beetle]
+
Tr. Paussini Latreille 1806
+
Subtr. †Arthropteritina Luna de Carvalho 1961
Subtr. Carabidomemnina Wasmann 1928
Subtr. Cerapterina Billberg 1820
Subtr. †Eopaussina Luna de Carvalho 1951
Subtr. Paussina Latreille 1806
G. †Pleurarthropterus Wasmann 1929
+
Subg. †Pleurarthropterus (Acmarthropterus) Wasmann 1929
Subg. †Pleurarthropterus (Balticarthropterus) Nagel 1987
Subg. †Pleurarthropterus (Pleurarthropterus) Wasmann 1929
G. †Succinarthropterus Kolbe 1926
+
†Succinarthropterus helmi Schaufuss 1896
†Succinarthropterus kolbei Wasmann 1926
†Succinarthropterus kuhlii Stein 1877
Invalid names: Arthropterillus Wasmann 1926 [synonym]
Invalid names: Ozaenidae Hope 1838 [synonym], Ozaenini [empty]
Subfm. Platyninae Bonelli 1810
+
Tr. Platynini Bonelli 1810
+
G. Agonum Bonelli 1810
G. Anchomenus Bonelli 1810
G. Oxypselaphus Chaudoir 1843
G. Paranchus Lindroth 1974
Subtr. Platynina Bonelli 1810
G. Platynus Bonelli 1810
G. †Praeanchodemus Schmidt et al. 2017
G. †Procarabites Handlirsch 1906
Subfm. Protopaussinae Gestro 1892
+
Subfm. †Protorabinae Ponomarenko 1977
+
G. †Cordorabus Ponomarenko 1977
+
†Cordorabus antennatus Ponomarenko 1977
†Cordorabus minimus Ponomarenko 1977
†Cordorabus notatus Ponomarenko 1977
†Cordorabus striatus Ponomarenko 1986
†Cordorabus vittatus Ponomarenko 1980
†Cordorabus wuqiensis Hong 1995
G. †Cretorabus Ponomarenko 1977
+
†Cretarabus medius Hong and Wang 1990
†Cretorabus capitatus Ponomarenko 1977
†Cretorabus latus Ponomarenko 1977
†Cretorabus orientalis Ponomarenko 1989
†Cretorabus ovalis Ponomarenko 1989
†Cretorabus rasnitsyni Wang and Zhang 2011
†Cretorabus sulcatus Ponomarenko et al. 2005
G. †Ovrabites Ponomarenko 1977
+
†Ovrabites incertus Ponomarenko 1993
†Ovrabites jurassicus Ponomarenko 1977
†Ovrabites ovalis Ponomarenko 1977
G. †Protorabus Ponomarenko 1977
+
†Protorabus crassus Ponomarenko 1989
†Protorabus elongatus Ponomarenko 1986
†Protorabus kobdoensis Ponomarenko 1986
†Protorabus magnus Ponomarenko 1977
†Protorabus minisculus Zhang 1997
†Protorabus nigrimonticola Ponomarenko 1977
†Protorabus planus Ponomarenko 1977
†Protorabus polyphlebius Ren 1995
†Protorabus tsagensis Ponomarenko 1989
G. Stereocerus Kirby 1837
+
Stereocerus haematopus Dejean 1831
Stereocerus rubripes Motschulsky 1860
Subfm. Trechinae Bonelli 1810
+
Unr. Trechitae Bonelli 1810
+
G. †Balticeler Schmidt and Maddison 2021
Tr. Bembidiini Stephens 1827
Tr. Pogonini Laporte 1834
Tr. Tachyini Motschulsky 1862
Tr. Trechini Bonelli 1810
Invalid names: Bembidion succini Giebel 1856 [nomen dubium], Patrobinae Kirby 1837 [synonym]
Invalid names: Cratacanthus florissantensis Wickham 1917 [nomen dubium], Fossores Latreille 1802 [empty], Loricera exita Scudder 1900 [nomen dubium], Rhysodinae Laporte 1840 [empty], Siagoninae Bonelli 1813 [empty], Trechus fractus Wickham 1912 [nomen dubium]
Infraor. Carabiformia Lameere 1900
+
Superfm. Caraboidea Latreille 1802
+
G. †Carabilarva Ponomarenko 1985
+
†Carabilarva jurassica Ponomarenko 1985
†Carabilarva robusta Makarov 1995
†Carabilarva triassica Makarov and Prokin 2013
G. †Eurycoleus Hong 1982
+
†Eurycoleus arcuatus Hong 1982
†Eurycoleus clypeolatus Hong 1982
†Eurycoleus dimorphocellatus Hong 1982
†Eurycoleus foveolatus Hong 1984
†Eurycoleus parvus Hong 1982
G. †Jiangxicoleus Hong 1988
+
†Jiangxicoleus guixiense Hong 1988
†Jiangxicoleus yiyangense Hong 1988
G. †Procalosoma Meunier 1895
+
†Procalosoma giardi Meunier 1895
†Procalosoma incertum Ponomarenko 2014
†Procalosoma major Handlirsch 1906
†Procalosoma mimor Handlirsch 1906
†Procalosoma minor Handlirsch 1906
G. †Yumenocoleus Hong 1982
+
†Yumenocoleus intermedius Hong 1982
†Yumenocoleus lineatus Hong 1982
†Yumenocoleus longus Hong 1982
†Yumenocoleus nantianmenensis Hong 1984
†Yumenocoleus tenuis Hong 1988
Invalid names: Unda chonggezhuangensis Wang 1998 [nomen dubium], Unda pandurata Ren 1995 [nomen dubium]
Fm. Cicindelidae Latreille 1802 [tiger beetle]
+
Tr. Cicindelini Latreille 1802
+
G. †Hujia Song et al. 2022
Invalid names: Oxycheilini Thomson 1857 [empty]
Fm. †Colymbotethidae Ponomarenko 1993
+
Fm. †Coptoclavidae Ponomarenko 1961
+
G. †Amblycephalonius Bode 1953
+
†Amblycephalonius tenuistriatus Bode 1953
+
Invalid names: Barycephalus nudatus Bode 1953 [synonym], Tricyrtus tenuistriatus Bode 1953 [synonym]
Invalid names: Barocephalus Carpenter 1986 [synonym], Barycephalus Bode 1953 [synonym], Tricyrtus Bode 1953 [synonym]
Subfm. †Charonoscaphinae Ponomarenko 1977
+
G. †Charonoscapha Ponomarenko 1977
+
†Charonoscapha grossa Ponomarenko 1977
†Charonoscapha ovata Ponomarenko 1977
Subfm. †Coptoclavinae Ponomarenko 1961
+
G. †Bolbonectes Ponomarenko 1987
+
†Bolbonectes intermedius Ponomarenko 1987
†Bolbonectes occidentalis Ponomarenko 1993
†Bolbonectus lithographicus Ponomarenko and Martínez-Delclòs 2000
G. †Megacoptoclava Ponomarenko and Martínez-Delclòs 2000
+
†Megacoptoclava longiurogomphia Ponomarenko and Martínez-Delclòs 2000
Subfm. †Coptoclaviscinae Soriano et al. 2007
+
G. †Coptoclavella Ponomarenko 1980
+
†Coptoclavella elegans Ponomarenko 1980
†Coptoclavella inexpecta Soriano et al. 2007
†Coptoclavella jurassica Ponomarenko 2014
†Coptoclavella minor Ponomarenko 1980
†Coptoclavella purbeckensis Ponomarenko et al. 2005
†Coptoclavella striata Ponomarenko 1986
†Coptoclavella vittata Ponomarenko 1986
G. †Coptoclavisca Ponomarenko 1987
+
†Coptoclavisca grandioculus Zhang 1992
†Coptoclavisca nigricollinus Ponomarenko 1987
G. †Holcoptera Handlirsch 1906
+
†Holcoptera alisonae Kelly et al. 2017
†Holcoptera giebeli Handlirsch 1906
†Holcoptera pigmentatus Kelly et al. 2017
†Holcoptera rasnitsyni Ponomarenko and Fedorenko 2022
†Holcoptera solitensis Kelly et al. 2017
Invalid names: Holcoelytrum Handlirsch 1906 [synonym], Stargelytron Ponomarenko 2015 [synonym]
Subfm. †Hispanoclavinae Soriano et al. 2007
+
G. †Hispanoclavina Soriano et al. 2007
+
†Hispanoclavina diazromerali Soriano et al. 2007
†Hispanoclavina gratshevi Soriano et al. 2007
Subfm. †Timarchopsinae Wang et al. 2010
+
G. †Timarchopsis Brauer et al. 1889
+
†Timarchopsis aquaticus Ponomarenko 1977
†Timarchopsis cyrenaicus Ponomarenko 1977
†Timarchopsis czekanowskii Brauer et al. 1889
†Timarchopsis gigas Ponomarenko 1977
†Timarchopsis gobiensis Ponomarenko 1987
†Timarchopsis latus Ponomarenko 1977
†Timarchopsis longus Ponomarenko 2014
†Timarchopsis mongolicus Ponomarenko 1985
†Timarchopsis sainshandensis Ponomarenko 1987
Invalid names: Necronectes Ponomarenko 1977 [synonym]
Invalid names: Necronectinae Ponomarenko 1977 [replaced]
Fm. Dytiscidae Leach 1815 [predaceous diving beetle]
+
Subfm. Agabinae Thomson 1867
+
G. Agabus Leach 1817
+
Subg. Agabus (Acatodes) Thomson 1859
Subg. Agabus (Agabus) Leach 1817
Subg. Agabus (Gaurodytes) Thomson 1859
†Agabus charon Wickham 1912
†Agabus congeneroides Lomnicki 1894
†Agabus corticeus Lomnicki 1894
†Agabus florissantensis Wickham 1913
†Agabus hochi Statz 1939
†Agabus latissimus Ponomarenko 2019
†Agabus lithax Říha 1960
†Agabus niedzwiedzkii Lomnicki 1894
†Agabus parvulus Lomnicki 1894
†Agabus perditus Scudder 1900
†Agabus praelugens Wickham 1917
†Agabus rathbuni Scudder 1900
†Agabus reductus von Heyden and von Heyden 1866
†Agabus reitteri Lomnicki 1894
†Agabus savagei Wickham 1917
†Agabus ungeri Heer 1847
Invalid names: Necticus Hope 1838 [synonym]
G. Ilybius Erichson 1832
+
Ilybius apicalis Sharp 1873
†Ilybius boryslavicus Lomnicki 1894
Ilybius fenestratus Fabricius 1781
†Ilybius gratshevi Ponomarenko 2019
Ilybius quadrimaculatus Aubé 1838
Ilybius subaeneus Erichson 1837
Ilybius vittiger Gyllenhal 1827
Subfm. Copelatinae Van den Branden 1885
+
G. Copelatus Erichson 1832
+
†Copelatus aphroditae Balke 2003
†Copelatus convexus Förster 1891
†Copelatus fossilis Říha 1974
†Copelatus ponomarenkoi Říha 1974
†Copelatus predaveterus Miller 2003
†Copelatus stavropolitanus Říha 1974
Invalid names: Copelatini Van den Branden 1885 [empty]
Subfm. Coptotominae Van den Branden 1885
+
G. †Cretodytes Ponomarenko 1977
+
†Cretodytes incertus Prokin et al. 2013
†Cretodytes latipes Ponomarenko 1977
Subfm. Cybisterinae Sharp 1880
+
G. Cybister Curtis 1827
+
Subg. Cybister (Cybister) Curtis 1827
†Cybister agassizi Heer 1862
†Cybister atavus Heer 1862
†Cybister fractus Říha 1974
†Cybister imperfectus Říha 1974
†Cybister mancus Říha 1974
†Cybister rotundatus Říha 1974
G. Megadytes Sharp 1882
+
Subg. Megadytes (Paramegadytes) Trémouilles and Bachmann 1980
†Megadytes nicoleti Heer 1862
Subfm. Dytiscinae Leach 1815
+
Subfm. Hydroporinae Aubé 1836
+
Tr. Bidessini Sharp 1880
+
G. Bidessus Sharp 1882
Invalid names: Bidessus laminarum Wickham 1914 [nomen dubium], Hydroporus sedimentorum Wickham 1914 [nomen dubium]
Tr. Hydroporini Aubé 1836
+
Subtr. Deronectina Galewski 1994
G. Hydrocolus Roughley and Larson 2000
G. Hydroporus Clairville 1806
G. †Pteroporus Guignot 1933
Subfm. Laccophilinae Gistel 1848
+
Invalid names: Palaeogyrininae von Schlechtendal 1894 [synonym]
Subfm. †Liadytiscinae Prokin and Ren 2010
+
Tr. †Liadytiscini Prokin and Ren 2010
+
G. †Liadytiscus Prokin and Ren 2010
G. †Liadyxianus Prokin et al. 2013
Tr. †Mesoderini Prokin et al. 2013
+
G. †Mesoderus Prokin and Ren 2010
G. †Mesodytes Prokin et al. 2013
G. †Palaeodytes Ponomarenko 1987
+
†Palaeodytes baissiensis Prokin et al. 2013
†Palaeodytes gutta Ponomarenko 1987
†Palaeodytes incompleta Ponomarenko et al. 2005
†Palaeodytes sibiricus Ponomarenko 1987
Invalid names: Aubehydrinae Guignot 1942 [empty], Necticus palustris Aymard 1856 [nomen nudum]
Fm. Gyrinidae Latreille 1810 [whirligig beetle]
+
Subfm. Gyrininae Latreille 1810
+
Tr. Enhydrusini Branden 1882
+
Invalid names: Dineutina Desmarest 1851 [empty], Dineutini Desmarest 1851 [replaced]
G. †Protogyrininus Hatch 1927
+
†Protogyrininus sculpturatus Mjöberg 1905
+
Invalid names: Protogyrininus confinis Hatch 1927 [synonym]
Invalid names: Spanglerogyrinae Folkerts 1979 [empty]
Subfm. Haliplidae Aubé 1836 [crawling water beetle]
+
G. †Cretihaliplus Ren et al. 1995
+
†Cretihaliplus chifengensis Ren et al. 1995
†Cretihaliplus sidaojingensis Ren et al. 1995
G. Haliplus Latreille 1802
+
†Haliplus cretaceus Prokin and Ponomarenko 2013
Haliplus obliquus Fabricius 1787
Haliplus ruficollis De Geer 1774
G. Peltodytes Regimbart 1878
Fm. Hygrobiidae Regimbart 1878
+
G. Hygrobia Latreille 1804
+
†Hygrobia cretzschmari von Heyden and von Heyden 1866
Invalid names: Paelobius Schoenherr 1808 [synonym]
Invalid names: Paelobiidae Erichson 1837 [synonym]
Fm. †Liadytidae Ponomarenko 1977
+
G. †Liadytes Ponomarenko 1963
+
†Liadytes aspidytoides Prokin et al. 2013
†Liadytes avus Ponomarenko 1963
†Liadytes crassus Ponomarenko 1977
†Liadytes dajensis Ponomarenko 1987
†Liadytes longus Ponomarenko 1977
†Liadytes major Ponomarenko 1985
G. †Memptus Handlirsch 1906
+
†Memptus ganglbaueri Ponomarenko 1985
†Memptus guttula Ponomarenko 1985
†Memptus handlirschi Ponomarenko 1985
Fm. Noteridae Thomson 1860 [burrowing water beetle]
+
Subfm. Noterinae Thomson 1860
+
Tr. Noterini Thomson 1860
+
G. Canthydrus Sharp 1882
G. Hydrocanthus Say 1823
G. Noterus Clairville 1806
Invalid names: Hydrocanthini Sharp 1880 [synonym]
Fm. †Parahygrobiidae Ponomarenko 1977
+
Fm. Trachypachidae Thomson 1857 [false ground beetle]
+
Subfm. †Eodromeinae Ponomarenko 1977
+
G. †Ademosynoides Dunstan 1923
+
†Ademosynoides abnormis Dunstan 1923
†Ademosynoides alternata Dunstan 1923
†Ademosynoides angusta Tillyard 1916
†Ademosynoides antarcticus Zeuner 1959
†Ademosynoides japonicus Fujiyama 1973
†Ademosynoides magnifica Dunstan 1923
†Ademosynoides minor Handlirsch 1906
†Ademosynoides obtusa Tillyard 1916
†Ademosynoides striatella Dunstan 1923
G. †Eodromeus Ponomarenko 1977
+
†Eodromeus antiquus Ponomarenko 1977
†Eodromeus daohugouensis Wang et al. 2012
†Eodromeus dissectus Ponomarenko 1977
†Eodromeus major Ponomarenko 1977
†Eodromeus mongolicus Ponomarenko 1989
†Eodromeus robustus Wang et al. 2012
†Eodromeus sternalis Ponomarenko 1977
†Eodromeus sulcatus Ponomarenko 1989
G. †Evertus Ponomarenko 1986
+
†Evertus cornutus Ponomarenko 1986
†Evertus notatus Ponomarenko 1986
†Evertus pervalimand Jia and Ren 2011
Invalid names: Fortiseode Jia and Ren 2011 [synonym]
G. †Karadromeus Ponomarenko 1977
+
†Karadromeus capitatus Ponomarenko 1986
†Karadromeus elongatus Ponomarenko 1977
†Karadromeus erensis Ponomarenko 1986
†Karadromeus gobiensis Ponomarenko 1980
†Karadromeus latus Ponomarenko 1977
†Karadromeus mongolicus Ponomarenko 1977
†Karadromeus rostratus Ponomarenko 1977
†Karadromeus verrucosus Ponomarenko 1989
G. †Novunda Kirejtshuk and Ansorge 2022
+
†Novunda angulata Ponomarenko 1977
†Novunda chifengensis Wang et al. 2012
†Novunda cursoria Ponomarenko 1977
†Novunda microplata Ponomarenko 1977
†Novunda pachycephala Ponomarenko 1989
Invalid names: Unda Ponomarenko 1977 [replaced]
G. †Platycoxa Ponomarenko 1977
+
†Platycoxa armata Ponomarenko 1977
†Platycoxa jurassica Ponomarenko 1977
G. †Prosynactus Bode 1953
+
†Prosynactus gracilis Bode 1953
+
Invalid names: Cardioides incisus Bode 1953 [synonym], Cyclotrachelus exsecatus Bode 1953 [synonym], Cyphospheron virgatus Bode 1953 [synonym], Liassocarabites pulcher Bode 1953 [synonym], Mesalocistus constrictus Bode 1953 [synonym], Rhabdotus cingulatus Bode 1953 [synonym]
†Prosynactus scissus Bode 1953
+
Invalid names: Cardioides longealatus Bode 1953 [synonym], Euenarthrus mandibulatus Bode 1953 [synonym], Liassocarabites praefrictus Bode 1953 [synonym], Megelytrites mutilatus Bode 1953 [synonym], Trapezotrachelus longus Bode 1953 [synonym], Trigonocephalites sulcatus Bode 1953 [synonym]
Invalid names: Critotrachelus Carpenter 1986 [synonym], Cyclotrachelus Bode 1953 [synonym], Dysmorphus Bode 1953 [synonym], Euenarthrus Bode 1953 [synonym], Leptynticus Bode 1953 [synonym], Liassocarabites Bode 1953 [synonym], Megelytrites Bode 1953 [synonym], Mesalocistus Bode 1953 [synonym], Rhabdotus Bode 1953 [synonym], Tetragonotrachelus Bode 1953 [synonym], Trapezotrachelus Bode 1953 [synonym], Trigonocephalites Bode 1953 [synonym]
G. †Psacodromeus Ponomarenko 1977
+
†Psacodromeus crassus Ponomarenko 1977
†Psacodromeus gutta Ponomarenko 1977
†Psacodromeus minor Ponomarenko 1986
†Psacodromeus ovalis Ponomarenko 1977
†Psacodromeus rugosus Ponomarenko 1977
G. †Eodromeites Ponomarenko 1985
+
†Eodromeites minimus Ponomarenko 1985
†Eodromeites udensis Ponomarenko 1985
Subfm. Trachypachinae Thomson 1857
+
Invalid names: Karadromeus xiangfanggouensis Hong 1995 [nomen dubium]
Fm. †Triaplidae Ponomarenko 1977
+
G. †Catinius Ponomarenko 1968
+
†Catinius artus Zhang 1997
†Catinius ovatus Zhang 1997
†Catinius pelta Ponomarenko 1968
G. †Tomiaplus Ponomarenko 2021
+
†Tomiaplus minimus Ponomarenko 2021
†Tomiaplus sibiricus Volkov 2013
G. †Triaplus Ponomarenko 1977
+
†Triaplus laticoxa Ponomarenko 1977
†Triaplus macroplatus Ponomarenko 1977
G. †Triassocatinius Ponomarenko 1969
+
†Triassocatinius brachynotus Ponomarenko 1969
†Triassocatinius glabratus Ponomarenko 1969
G. †Tunguskagyrus Yan et al. 2018
+
†Tunguskagyrus planus Yan et al. 2018
†Tunguskagyrus yani Ponomarenko 2021
Invalid names: Catiniidae Ponomarenko 1968 [replaced]
Invalid names: Amphizoidae Leconte 1853 [empty], Dytiscoidea Leach 1815 [invalid subgroup], Pseudomorphinae Hope 1838 [empty], Tritarsusidae Hong 2002 [invalid subgroup]
Subor. Archostemata Kolbe 1908
+
G. †Argentinocupes Martins-Neto and Gallego 2006
+
†Argentinocupes abdalai Martins-Neto and Gallego 2006
†Argentinocupes pulcher Martins-Neto and Gallego 2006
†Argentinocupes sara Martins-Neto and Gallego 2009
Superfm. †Asiocoleoidea Rohdendorf 1961
+
Fm. †Tricoleidae Ponomarenko 1969
+
G. †Sogdelytron Ponomarenko 1969
+
†Sogdelytron latum Ponomarenko 1969
†Sogdelytron tristichus Hong 1985
Invalid names: Longxianocupes Hong 1985 [synonym]
G. †Tricoleodes Ponomarenko 1969
+
†Tricoleodes acutus Ponomarenko 1969
†Tricoleodes longus Ponomarenko 1969
G. †Tricoleus Ponomarenko 1969
+
†Tricoleus incertus Ponomarenko 1969
†Tricoleus punctatus Ponomarenko 1969
G. †Brachilatus Strelnikova and Yan 2023
+
†Brachilatus caducus Ponomarenko 1969
†Brachilatus dundulaensis Ponomarenko 1994
†Brachilatus foersteri Ponomarenko 1971
+
Invalid names: Notocupes reticulatus Ponomarenko 1968 [replaced]
†Brachilatus longicoxa Soriano and Delclos 2006
†Brachilatus nigrimonticola Ponomarenko 1968
†Brachilatus oculatus Soriano and Delclos 2006
†Brachilatus premeris Lee et al. 2022
†Brachilatus spinosus Li and Cai 2023
†Brachilatus viridis Soriano and Delclos 2006
G. †Conexicoxa Lin 1986
+
†Conexicoxa brachycephala Ponomarenko 1994
†Conexicoxa crassa Ponomarenko 1969
†Conexicoxa denticollis Jiang et al. 2020
†Conexicoxa epicharis Tan et al. 2005
†Conexicoxa eumeura Tan et al. 2006
†Conexicoxa exigua Ponomarenko 1994
†Conexicoxa homora Lin 1986
†Conexicoxa jurassica Hong 1983
†Conexicoxa khetanensis Ponomarenko 1993
†Conexicoxa kirghizica Ponomarenko 1969
†Conexicoxa longicollis Ponomarenko 1994
†Conexicoxa martinclosas Soriano and Delclos 2006
†Conexicoxa porrecta Tan et al. 2006
†Conexicoxa psilata Tan et al. 2005
†Conexicoxa siniestri Soriano and Delclos 2006
†Conexicoxa stabilis Tan et al. 2005
†Conexicoxa tripartita Oppenheim 1888
Superfm. Cupedoidea Laporte 1836
+
Fm. Cupedidae Laporte 1836 [reticulated beetle]
+
Subfm. Cupedinae Lacordaire 1857
+
G. †Anaglyphites Ponomarenko 1964
G. †Apriacma Kirejtshuk et al. 2016
G. †Asimma Ponomarenko 1966
G. †Barbaticupes Jarzembowski et al. 2017
G. †Cainomerga Kirejtshuk et al. 2016
G. †Cretomerga Kirejtshuk et al. 2016
G. Cupes Fabricius 1801
+
Invalid names: Cupoides Motschulsky 1856 [synonym], Distocupes Neboiss 1984 [synonym], Tenomerga Neboiss 1984 [synonym]
G. †Cupidium Ponomarenko 1968
G. †Cupopsis Kirejtshuk et al. 2016
G. †Ensicupes Hong 1976
G. †Furcicupes Tan and Ren 2006
G. †Gracilicupes Tan et al. 2006
G. †Mallecupes Jarzembowski et al. 2017
G. †Menatops Kirejtshuk et al. 2016
G. †Mesocupes Martynov 1926
G. †Miocupes Ponomarenko 1973
G. †Priacmopsis Ponomarenko 1966
G. †Pterocupes Ponomarenko 1966
G. †Taxopsis Kirejtshuk et al. 2016
Invalid names: Anaglyphites pluricavus Soriano and Delclos 2006 [nomen dubium], Cupedini Crowson 1962 [empty], Platycupes sogdianus Ponomarenko 1966 [nomen dubium], Priacma sanzii Soriano and Delclos 2006 [nomen dubium]
Invalid names: Mesocupedinae Ponomarenko 1969 [empty]
Fm. Micromalthidae Barber 1913 [telephone pole beetle]
+
G. Micromalthus LeConte 1878
+
†Micromalthus eocenicus Kirejtshuk et al. 2010
†Micromalthus priabonicus Perkovsky 2016
Fm. Ommatidae Sharp and Muir 1912
+
Subfm. Ommatinae Sharp and Muir 1912
+
G. †Allophalerus Kirejtshuk 2020
G. †Brochocoleus Hong 1982
G. †Burmocoleus Kirejtshuk 2020
G. †Cionocoleus Ren 1995
Tr. †Clessidrommatini Jarzembowski et al. 2017
G. †Diluticupes Ren 1995
G. †Echinocups Kirejtshuk and Jarzembowski 2020
G. †Eurydictyon Ponomarenko 1969
G. †Jarzembowskiops Kirejtshuk 2020
G. †Liassocupes Zeuner 1962
G. †Lithocupes Ponomarenko 1966
G. †Monticupes Ren 1995
G. †Notocupoides Ponomarenko 1966
G. †Odontomma Ren et al. 2006
Tr. Ommatini Sharp and Muir 1912
G. †Paraodontomma Yamamoto 2017
G. †Polyakius Kirejtshuk 2020
G. †Rhopalomma Ashman et al. 2015
G. †Stegocoleus Jarzembowski and Wang 2016
G. †Tetraphalerites Crowson 1962
G. Tetraphalerus Waterhouse 1901
Invalid names: Lithocupedini Ponomarenko 1969 [invalid subgroup], Notocupedinae Ponomarenko 1966 [synonym], Pronotocupedini Tan et al. 2012 [invalid subgroup], Tetraphalerini Crowson 1962 [invalid subgroup], Tetraphalerus brevicapitis Ponomarenko and Martínez-Delclòs 2000 [nomen dubium]
Invalid names: Brochocoleinae Hong 1982 [empty]
Invalid names: Crowsoniellidae Iablokoff-Khnzorian 1983 [empty], Magnocoleidae Hong 1998 [invalid subgroup]
G. †Notocupes Ponomarenko 1964
+
†Lupicupes trachylaenus Ren 1995
†Notocupes alienus Tan and Ren 2006
†Notocupes baojiatunensis Hong 1992
†Notocupes caudatus Ponomarenko 1966
†Notocupes cyclodontus Tan et al. 2006
†Notocupes daohugouensis Li and Cai 2023
†Notocupes diazromerali Soriano and Delclos 2006
†Notocupes elegans Ponomarenko 1994
†Notocupes excellens Ponomarenko 1966
†Notocupes kezuoensis Hong 1987
†Notocupes khasurtyiensis Strelnikova 2019
†Notocupes laetus Lin 1976
†Notocupes laiyangensis Hong and Wang 1990
†Notocupes lapidarius Ponomarenko 1968
†Notocupes latus Ponomarenko 1969
†Notocupes lini Ponomarenko et al. 2012
†Notocupes ludongensis Wang and Liu 1996
†Notocupes mongolicus Ponomarenko 1994
†Notocupes neli Tihelka et al. 2019
†Notocupes ohmkuhnlei Jarzembowski et al. 2019
†Notocupes picturatus Ponomarenko 1964
†Notocupes pingi Ponomarenko and Ren 2010
†Notocupes pulcher Ponomarenko 1968
†Notocupes reticulatus Oppenheim 1888
†Notocupes robustus Li and Cai 2023
†Notocupes rudis Tan et al. 2005
†Notocupes shiluoensis Hong 1984
†Notocupes tuanwangensis Hong and Wang 1990
†Notocupes validus Lin 1976
Invalid names: Amblomma Tan et al. 2005 [synonym], Chengdecupes Hong 1983 [synonym], Forticupes Hong and Wang 1990 [synonym], Lupicupes Ren 1995 [synonym], Ovatocupes Tan and Ren 2006 [synonym], Picticupes Hong and Wang 1990 [synonym], Sinocupes Lin 1976 [synonym]
G. †Pseudosilphites Zeuner 1930
+
†Pseudosilphites natalensis Zeuner 1960
†Pseudosilphites triassicus Zeuner 1930
G. †Rhabdocupes Ponomarenko 1966
+
†Rhabdocupes baculatus Ponomarenko 1969
†Rhabdocupes cellulosus Ponomarenko 1969
†Rhabdocupes dischides Zhang 1986
†Rhabdocupes issykkulensis Ponomarenko 1969
†Rhabdocupes laticella Ponomarenko 1969
†Rhabdocupes lentus Ren 1995
†Rhabdocupes longus Ponomarenko 1966
†Rhabdocupes minisculus Tan et al. 2006
†Rhabdocupes minor Ponomarenko 1966
†Rhabdocupes oxypygus Ponomarenko 1969
†Rhabdocupes protensus Tan et al. 2006
†Rhabdocupes rostratus Ponomarenko 1969
†Rhabdocupes sogutensis Ponomarenko 1969
†Rhabdocupes tenuis Ponomarenko 1969
†Rhabdocupes vitimensis Ponomarenko 1966
Superfm. †Rhombocoleoidea Rohdendorf 1961
+
Fm. †Rhombocoleidae Rohdendorf 1961
+
G. †Rhombocoleites Ponomarenko 1969
+
†Rhombocoleites adumbratus Ponomarenko 1969
†Rhombocoleites danutae Ponomarenko and Mostovski 2005
†Rhombocoleites uralensis Ponomarenko 2013
G. †Rhombocoleus Rohdendorf 1961
+
†Rhombocoleus andreae Rohdendorf 1961
†Rhombocoleus gomankovi Ponomarenko 2013
G. †Rossocoleus Rohdendorf 1961
+
†Rossocoleus altus Ponomarenko 2004
†Rossocoleus angustus Ponomarenko 2013
†Rossocoleus aristovi Ponomarenko 2013
†Rossocoleus avus Rohdendorf 1961
†Rossocoleus ellipticus Martynov 1937
†Rossocoleus grandis Rohdendorf 1961
†Rossocoleus latior Rohdendorf 1961
†Rossocoleus latus Rohdendorf 1961
†Rossocoleus novojilovi Ponomarenko 2013
†Rossocoleus parvus Rohdendorf 1961
†Rossocoleus sakmara Ponomarenko 2013
†Rossocoleus surijokovensis Rohdendorf 1961
†Rossocoleus verrucosus Rohdendorf 1961
Invalid names: Curculiopsis Martynov 1937 [replaced]
Superfm. †Schizocoleoidea Rohdendorf 1961
+
Fm. †Phoroschizidae Bouchard and Bousquet 2020
+
G. †Phoroschizus Bouchard and Bousquet 2020
+
†Phoroschizus crassus Ponomarenko 1968
Invalid names: Schizophorus Ponomarenko 1968 [replaced]
Invalid names: Schizophoridae Ponomarenko 1968 [replaced]
Fm. †Schizocoleidae Rohdendorf 1961
+
G. †Aenigmocoleus Rohdendorf 1961
+
†Aenigmocoleus borissiaki Rohdendorf 1961
†Aenigmocoleus uralensis Ponomarenko 2013
G. †Argentinosyne Martins-Neto and Gallego 2006
+
†Argentinosyne bonapartei Martins-Neto and Gallego 2006
†Argentinosyne duraznoensis Martins-Neto and Gallego 2008
†Argentinosyne frenguellii Martins-Neto and Gallego 2006
†Argentinosyne gonaldiae Martins-Neto and Gallego 2006
†Argentinosyne gualoensis Martins-Neto and Gallego 2006
†Argentinosyne ischichucaensis Martins-Neto and Gallego 2009
†Argentinosyne losrastrosensis Martins-Neto and Gallego 2006
†Argentinosyne rugosa Martins-Neto and Gallego 2006
G. †Erunakicupes Rohdendorf 1961
+
†Erunakicupes angustus Ponomarenko 2011
†Erunakicupes brevis Ponomarenko 2004
†Erunakicupes fomitshevi Rohdendorf 1961
†Erunakicupes grossus Ponomarenko 2013
†Erunakicupes kryshtofovichi Rohdendorf 1961
†Erunakicupes obrutshevi Rohdendorf 1961
†Erunakicupes schmalhauseni Rohdendorf 1961
Invalid names: Schizocupes Rohdendorf 1961 [synonym]
G. †Karakanocoleus Rohdendorf 1961
+
†Karakanocoleus europeus Ponomarenko 2004
†Karakanocoleus janischewskyi Rohdendorf 1961
†Karakanocoleus magnus Rohdendorf 1961
†Karakanocoleus major Rohdendorf 1961
+
Invalid names: Karakanocoleodes latissimus Rohdendorf 1961 [synonym]
†Karakanocoleus mutovinensis Ponomarenko 2013
†Karakanocoleus pincombeae Tillyard 1924
†Karakanocoleus striatus Rohdendorf 1961
†Karakanocoleus venjukovi Rohdendorf 1961
+
Invalid names: Karakanocoleus lebedevi Rohdendorf 1961 [synonym]
Invalid names: Karakanocoleodes Rohdendorf 1961 [synonym]
G. †Mesolobites Carpenter 1986
+
†Mesolobites granulatus Dunstan 1923
†Mesolobites margacrispus Lin and Mou 1989
†Mesolobites punctatoides Lin 1986
†Mesolobites quadripartita Dunstan 1923
†Mesolobites tenuiclathrata Dunstan 1923
†Mesolobites trivittatus Dunstan 1923
†Mesolobites tuberculatus Dunstan 1923
Invalid names: Lobites Dunstan 1923 [replaced], Shijingocoleus Lin and Mou 1989 [synonym]
G. †Metrorhynchites Tillyard 1916
+
†Metrorhynchites angustior Rohdendorf 1961
†Metrorhynchites angustum Tillyard 1918
†Metrorhynchites angustus Rohdendorf 1961
†Metrorhynchites baculum Ponomarenko 2011
†Metrorhynchites dilutes Lin 1986
†Metrorhynchites dunstani Tillyard 1916
†Metrorhynchites elongatus Ponomarenko 2011
†Metrorhynchites grandis Tillyard 1918
†Metrorhynchites longus Rohdendorf 1961
†Metrorhynchites putativus Lin 1986
†Metrorhynchites sydneiensis Tillyard 1916
Invalid names: Mancicupes Lin 1986 [synonym]
G. †Mimemala Strand 1936
+
†Mimemala giganteum Whalley 1985
†Mimemala punctatum Handlirsch 1906
Invalid names: Mimema Handlirsch 1906 [replaced]
G. †Pseudochrysomelites Handlirsch 1906
+
†Pseudochrysomelites anser Ponomarenko 2008
†Pseudochrysomelites bashkuevi Ponomarenko 2013
†Pseudochrysomelites circumflexus Ponomarenko 2015
†Pseudochrysomelites convexus Ponomarenko 2015
†Pseudochrysomelites dilatatus Ponomarenko 2008
†Pseudochrysomelites dvinensis Ponomarenko 2018
†Pseudochrysomelites elongatus Ponomarenko 2004
†Pseudochrysomelites humeralis Ponomarenko 2016
†Pseudochrysomelites latissimus Ponomarenko 2015
†Pseudochrysomelites latum Ponomarenko 2004
†Pseudochrysomelites longus Ponomarenko 2013
†Pseudochrysomelites major Ponomarenko 2011
†Pseudochrysomelites medialis Ponomarenko 2013
†Pseudochrysomelites ovum Ponomarenko 2004
†Pseudochrysomelites planus Ponomarenko 2008
†Pseudochrysomelites rothenbachi Heer 1877
†Pseudochrysomelites sphenoidalis Ponomarenko 2013
†Pseudochrysomelites tumidus Ponomarenko 2016
G. †Reeveana Dunstan 1923
+
†Reeveana intermedia Dunstan 1923
†Reeveana major Dunstan 1923
†Reeveana minor Dunstan 1923
G. †Schizocoleus Rohdendorf 1961
+
†Schizocoleus depressus Ponomarenko 2013
†Schizocoleus glabrus Ponomarenko 2004
†Schizocoleus kuznetskiensis Rohdendorf 1961
†Schizocoleus longus Ponomarenko 2004
†Schizocoleus major Ponomarenko 2013
†Schizocoleus minimus Ponomarenko 2013
†Schizocoleus robustus Ponomarenko 2013
†Schizocoleus septentrionalis Ponomarenko 2008
†Schizocoleus triassicus Ponomarenko 2016
G. †Shepherdia Dunstan 1923
+
†Shepherdia quadrivittata Dunstan 1923
†Shepherdia trivittata Ponomarenko 1985
G. †Stegosyne Rohdendorf 1961
+
†Stegosyne borealis Ponomarenko 2008
†Stegosyne rugosa Rohdendorf 1961
G. †Uskatocoleus Rohdendorf 1961
+
†Uskatocoleus artus Ponomarenko 2015
†Uskatocoleus avitus Rohdendorf 1961
†Uskatocoleus convexus Ponomarenko 2013
†Uskatocoleus convexus Rohdendorf 1961
†Uskatocoleus eurypygus Ponomarenko 2011
†Uskatocoleus foveolatus Rohdendorf 1961
†Uskatocoleus hirsutus Ponomarenko 2015
†Uskatocoleus iljinskiensis Rohdendorf 1961
†Uskatocoleus kaltanicus Rohdendorf 1961
†Uskatocoleus kushejakovicus Rohdendorf 1961
†Uskatocoleus lutugini Rohdendorf 1961
†Uskatocoleus micron Ponomarenko 2013
†Uskatocoleus minimus Rohdendorf 1961
†Uskatocoleus minor Ponomarenko 2015
†Uskatocoleus rhynchophorus Ponomarenko 2011
†Uskatocoleus sukhonensis Ponomarenko 2013
†Uskatocoleus surijokovicus Rohdendorf 1961
†Uskatocoleus uralensis Ponomarenko 2013
†Uskatocoleus zalesskyi Rohdendorf 1961
Invalid names: Schizophoroidea Ponomarenko 1968 [synonym]
G. †Tentyridium Westwood 1854
+
†Tentyridium dilatatum Handlirsch 1906
†Tentyridium peleus Westwood 1854
G. †Tillyardiopsis Dunstan 1923
+
†Tillyardiopsis granulata Dunstan 1923
†Tillyardiopsis tuberculata Dunstan 1923
†Tillyardiopsis variotuberculata Dunstan 1923
G. †Zygadenia Handlirsch 1906
+
†Zygadenia alexrasnitsyni Strelnikova and Yan 2021
†Zygadenia angliae Giebel 1856
†Zygadenia cornigera Lee et al. 2023
†Zygadenia floodpagei Jarzembowski et al. 2015
†Zygadenia giebeli Ponomarenko 2014
†Zygadenia handlirschi Ponomarenko 2014
†Zygadenia liui Jarzembowski et al. 2015
†Zygadenia martinae Jarzembowski et al. 2015
†Zygadenia semen Ponomarenko 2000
†Zygadenia sibirica Ponomarenko 2000
†Zygadenia simpsoni Jarzembowski et al. 2015
†Zygadenia sinitzae Ponomarenko 2000
†Zygadenia tuberculata Giebel 1856
†Zygadenia westraliensis Riek 1968
Invalid names: Camtodontus Dejean 1826 [synonym], Kakoselia Handlirsch 1906 [synonym]
Invalid names: Cupediformia [invalid subgroup], Moravocoleus perditus Kukalová 1968 [nomen dubium], Notocupedini Ponomarenko 1966 [invalid subgroup], Priacma corrupta Ponomarenko 1986 [nomen dubium], Priacma longicapitis Ponomarenko 1997 [nomen dubium], Priacma oculata Ponomarenko 1997 [nomen dubium], Priacma striata Ponomarenko 2000 [nomen dubium]
Fm. †Asiocoleidae Rohdendorf 1961
+
G. †Bicoleus Ponomarenko 2013
+
†Bicoleus kuplensis Ponomarenko 2013
†Bicoleus laticella Ponomarenko 2013
G. †Loculitricoleus Tan and Ren 2009
+
†Loculitricoleus flatus Tan and Ren 2009
†Loculitricoleus mongolicus Ponomarenko 2014
†Loculitricoleus tenuatus Tan and Ren 2009
G. †Schizotaldycupes Rohdendorf 1961
+
†Schizotaldycupes ananjevi Rohdendorf 1961
†Schizotaldycupes beckermigdisovae Rohdendorf 1961
†Schizotaldycupes pubescens Ponomarenko 2013
Invalid names: Carinicupes Rohdendorf 1961 [synonym]
G. †Tetracoleus Ponomarenko 2008
+
†Tetracoleus aristovi Ponomarenko 2018
†Tetracoleus golubevi Ponomarenko 2015
†Tetracoleus meyeni Ponomarenko 2013
†Tetracoleus minimus Ponomarenko 2008
†Tetracoleus orenburgensis Ponomarenko 2013
†Tetracoleus sennikovi Ponomarenko 2011
†Tetracoleus tricoleoides Ponomarenko 2013
†Tetracoleus tshalyshevi Ponomarenko 2008
†Tetracoleus vasilenkoi Ponomarenko 2018
Subor. Myxophaga Crowson 1955
+
Superfm. Lepiceroidea Hinton 1936
+
Fm. Lepiceridae Hinton 1936
+
G. †Haplochelus Kirejtshuk and Poinar 2006
+
†Haplochelus ankylosaurus Jałoszyński et al. 2020
†Lepicerus georissoides Kirejtshuk and Poinar 2006
Invalid names: Lepiceratus Jałoszyński et al. 2020 [synonym]
G. Lepicerus Motschulsky 1855
G. †Lepichelus Kirejtshuk and Poinar 2017
+
†Lepichelus mumia Jałoszyński and Yamamoto 2017
†Lepichelus pretiosus Kirejtshuk and Poinar 2013
Invalid names: Lepiceroides Kirejtshuk and Poinar 2013 [replaced]
Invalid names: Cyathoceridae Scharp 1882 [synonym], Haplochelidae Kirejtshuk and Poinar 2006 [synonym], Lepicerinae Hinton 1936 [empty]
Superfm. Sphaeriusoidea
+
Fm. Hydroscaphidae LeConte 1874 [skiff beetle]
+
Subfm. †Triamyxinae Qvarnström et al. 2021
+
G. †Palaeoscapha Kirejtshuk et al. 2023
G. †Triamyxa Qvarnström et al. 2021
Invalid names: Sphaeriidae Erichson 1845 [empty], Torridincolidae Steffan 1964 [empty]
Invalid names: Sphaeriiformia [empty]
Fm. †Peltosynidae Yan et al. 2017
+
G. †Peltosyne Ponomarenko 1977
+
†Peltosyne triassica Ponomarenko 1977
†Peltosyne varyvrosa Yan et al. 2017
Subor. Polyphaga Emery 1886
+
G. †Angarolarva Ponomarenko 1985
+
†Angarolarva aquatica Ponomarenko 1985
†Angarolarva incerta Ponomarenko 1990
Infraor. Bostrichiformia Forbes 1926
+
Superfm. Bostrichoidea Latreille 1802
+
Fm. Bostrichidae Latreille 1802 [false powderpost beetle]
+
Subfm. †Alitrepaninae Peng et al. 2022
Subfm. Bostrichinae Latreille 1802
Subfm. Dinoderinae Thomson 1863
Subfm. Lyctinae Billberg 1820
Subfm. Polycaoninae Lesne 1896
G. †Protapate Wickham 1912
Invalid names: Dysidinae Lesne 1921 [empty], Euderiinae Lesne 1934 [empty], Lyctidae [replaced], Psoinae Blanchard 1851 [empty]
Fm. Dermestidae Latreille 1804 [skin beetle]
+
Subfm. Attageninae Laporte 1840
Subfm. Dermestinae Latreille 1804
Subfm. Megatominae Leach 1815
Subfm. Orphilinae Le Conte 1861
Subfm. Trinodinae Casey 1900
G. †Tryoniopsis Dunstan 1923
Fm. Ptinidae Latreille 1802 [spider beetle]
+
Subfm. Anobiinae Fleming 1821
G. †Cretasernus Peris and Philips 2019
Subfm. Dorcatominae Thomson 1859
Subfm. Dryophilinae Gistel 1848
Subfm. Ernobiinae Pic 1912
Subfm. Eucradinae Leconte 1861
Subfm. Mesocoelopodinae Mulsant and Rey 1864
Subfm. Ptininae Latreille 1802
Subfm. Xyletininae Gistel 1848
G. †Xyletinites von Heyden and von Heyden 1866
Invalid names: Alvarenganiellinae Viana and Martinez 1971 [empty], Anobiidae Fleming 1821 [synonym], Ptilininae Shuckard 1840 [empty]
Invalid names: Endecatomidae LeConte 1861 [empty]
Unr. Clambiformia Cai and Tihelka 2022
+
Superfm. Clamboidea Fischer von Waldheim 1821
+
Fm. Clambidae Fischer von Waldheim 1821 [minute beetle]
+
Subfm. Acalyptomerinae Crowson 1979
Subfm. Clambinae Fischer von Waldheim 1821
Invalid names: Calyptomerinae Crowson 1955 [empty], Ptismidae Kirejtshuk and Azar 2016 [synonym]
Fm. Derodontidae LeConte 1861 [tooth-necked fungus beetle]
+
Subfm. Laricobiinae Mulsant and Rey 1863
G. †Mesocryptogastra Lin 1977
Subfm. Peltasticinae LeConte 1861
Invalid names: Derodontinae LeConte 1861 [empty]
Fm. Eucinetidae Lacordaire 1857 [plate-thigh beetle]
+
G. †Cretohlezkus Jałoszyński 2019
G. Eucinetus Germar 1818
G. †Huaxiacinectus Hong 1995
Fm. †Mesocinetidae Kirejtshuk and Ponomarenko 2010
+
G. †Manoelodes Kirejtshuk and Ponomarenko 2010
G. †Manopsis Kirejtshuk and Ponomarenko 2010
G. †Mesocinetus Ponomarenko 1986
G. †Parashartegus Kirejtshuk and Ponomarenko 2010
G. †Shartegus Kirejtshuk and Ponomarenko 2010
Invalid names: Derodontoidea LeConte 1861 [synonym], Eucinetoidea Lacordaire 1857 [synonym]
G. †Cryptophagites Ponomarenko 1990
+
†Cryptophagites capitatus Ponomarenko 1990
†Cryptophagites clavatus Ponomarenko 1990
†Cryptophagites elongatus Ponomarenko 1990
Infraor. Cucujiformia Lameere 1938
+
Fm. †Apophisandridae Molino-Olmedo 2017
+
Superfm. Cleroidea Latreille 1802
+
Fm. Biphyllidae Leconte 1861 [false skin beetle]
+
G. Diplocoelus Guerin-Meneville 1844
G. †Paleobiphyllus Makarov and Perkovsky 2019
Fm. Byturidae Gistel 1848
+
Invalid names: Byturinae Gistel 1848 [empty], Platydascillinae Pic 1914 [empty]
Fm. Cleridae Latreille 1802 [checkered beetle]
+
G. †Anebomorpha Poinar and Brown 2020
Subfm. Clerinae Latreille 1802
+
Invalid names: Hydnocerinae Spinola 1844 [synonym], Phyllobaeninae Lacordaire 1857 [synonym]
Subfm. Epiclininae Gunter et al. 2013
Subfm. Korynetinae Laporte 1836
+
Invalid names: Enopliinae Gistel 1856 [synonym], Epiphloeinae Kuwert 1893 [synonym], Tarsosteninae Jacquelin du Val 1860 [synonym]
G. †Plastophagus Zhang 1989
G. †Protoclerus Kolibáč and Huang 2016
G. †Pyrrhoclerus Zhang 1989
Subfm. Tillinae Fischer von Waldheim 1813
G. †Titanoclerus Zhang 1989
Invalid names: Clerus adonis Heer 1847 [nomen dubium], Tanocleria Hong 2002 [nomen nudum], Trichodes stebingeri Cockerell 1936 [nomen dubium]
Fm. Lophocateridae Crowson 1964
+
G. Ancyrona Reitter 1876
G. Antillipeltis Lawrence et al. 2014
G. †Gracilenticrus Yu et al. 2021
G. Grynocharis Thomson 1859
G. Lophocateres Olliff 1883
G. †Parayixianteres Yu et al. 2020
G. †Zaiwa Lyubarsky et al. 2021
Invalid names: Ancyronini Reitter 1876 [invalid subgroup], Lophocaterini Crowson 1964 [invalid subgroup]
Fm. Melyridae Leach 1815 [soft-winged flower beetle]
+
Subfm. Dasytinae Laporte 1840
Subfm. Malachiinae Fleming 1821
Subfm. Melyrinae Leach 1815
G. Troglocollops Wittmer 1965
Fm. Peltidae Kirby 1837 [shield beetle]
+
G. Peltis Müller 1764
Invalid names: Peltinae Kirby 1837 [invalid subgroup], Peltini Latreille 1806 [invalid subgroup], Phenolia incapax Scudder 1876 [nomen dubium]
Fm. Prionoceridae Lacordaire 1857
+
G. †Cretaidgia Zhao et al. 2021
G. †Idgiaites Liu et al. 2015
G. †Prionocerites Lawrence et al. 2008
Fm. Rhadalidae LeConte 1861
+
G. †Cretorhadalus Kolibáč and Prokop 2023
Subfm. Rhadalinae LeConte 1861
G. Xamerpus Fairmaire 1886
Invalid names: Gietellinae Constantin and Menier 1987 [empty]
Fm. Thanerocleridae Chapin 1924
+
Subfm. Zenodosinae Kolibáč 1992
Invalid names: Thaneroclerinae Chapin 1924 [empty]
Fm. Thymalidae Leveille 1888
+
Invalid names: Decamerinae Crowson 1964 [empty], Thymalinae Leveille 1888 [empty]
Fm. Trogossitidae Latreille 1802 [bark gnawing beetle]
+
Subfm. Calityinae Winkler 1924
Subfm. Lophocaterinae Crowson 1964
G. †Marginulatus Yu et al. 2014
G. †Paracretocateres Yu et al. 2015
G. †Protrogosita Piton and Théobald 1937
Subfm. Trogossitinae Latreille 1802
Invalid names: Calitinae Winkler 1924 [empty], Ergoliinae Lacordaire 1854 [empty], Larinotinae Slipinski 1992 [empty], Ostomatidae Reitter 1882 [synonym], Temnochilidae [replaced]
Invalid names: Chaetosomatidae Crowson 1952 [empty], Phloiophilidae Kiesenwetter 1863 [empty], Phycosecidae Crowson 1952 [empty], Protopeltidae Crowson 1966 [empty], Rentoniidae Crowson 1966 [empty]
Superfm. Coccinelloidea Latreille 1807
+
Fm. Anamorphidae Strohecker 1953
+
G. †Discolomopsis Shockley 2006
G. †Giltine Alekseev and Tomaszewska 2018
G. †Gramboale Alekseev and Tomaszewska 2018
G. †Kleinzaches Alekseev and Tomaszewska 2018
G. †Palaecoryphus Alekseev and Tomaszewska 2018
G. Symbiotes Redtenbacher 1849
Fm. Cerylonidae Billberg 1820
+
Subfm. Ceryloninae Billberg 1820
Subfm. Ostomopsinae Sen Gupta and Crowson 1973
Invalid names: Loeblioryloninae Slipinski 1990 [empty]
Fm. Coccinellidae Latreille 1807 [ladybird beetle]
+
Subfm. Coccinellinae Latreille 1807
+
Invalid names: Coccidulinae Mulsant 1846 [synonym], Scymninae Mulsant 1846 [synonym]
Subfm. Microweiseinae Leng 1920
Invalid names: Chilocorinae Mulsant 1846 [empty]
Fm. Corylophidae Le Conte 1852 [minute hooded beetle]
+
Subfm. Corylophinae LeConte 1852
Invalid names: Peltinodinae Paulian 1950 [empty], Sericoderinae Matthews 1888 [empty]
Fm. Discolomatidae Horn 1878
+
Subfm. Aphanocephalinae Grouvelle 1912
Invalid names: Cephalophaninae John 1954 [empty], Discolomatinae Horn 1878 [empty], Notiophyginae Jacobson 1915 [empty], Pondonatinae John 1954 [empty]
Fm. Endomychidae Leach 1815 [handsome fungus beetle]
+
Subfm. Endomychinae Leach 1815
Subfm. Leiestinae Thomson 1863
Subfm. Lycoperdininae Redtenbacher 1844
Subfm. Pleganophorinae Jacquelin du Val 1858
G. †Rhomeocalpsua Li et al. 2022
Subfm. Xenomycetinae Strohecker 1962
Invalid names: Epipocinae Gorham 1873 [empty]
Fm. Latridiidae Erichson 1842 [minute brown scavenger beetle]
+
Subfm. Corticariinae Curtis 1829
Subfm. Latridiinae Erichson 1842
Fm. Teredidae Seidlitz 1888
+
Subfm. Teredinae Seidlitz 1888
Invalid names: Anommatinae Ganglbauer 1899 [empty], Xylariophilinae Pal and Lawrence 1986 [empty]
Fm. †Tetrameropsidae Kirejtshuk and Azar 2008
+
G. †Atetrameropsis Kirejtshuk 2013
G. †Tetrameropsis Kirejtshuk and Azar 2008
Invalid names: Alexiidae Imhoff 1856 [empty], Eupsilobiidae Casey 1895 [empty], Euxestidae Grouvelle 1908 [empty], Murmidiidae Jacquelin du Val 1858 [empty], Mycetaeidae Jacquelin du Val 1857 [empty]
Superfm. Cucujoidea Latreille 1802
+
Fm. Cryptophagidae Kirby 1837 [silken fungus beetle]
+
Subfm. Atomariinae LeConte 1861
Subfm. Cryptophaginae Kirby 1837
G. †Lebanophytum Kirejtshuk and Azar 2008
Invalid names: Hypocoprinae Reitter 1879 [empty], Lebanophytidae Kirejtshuk 2013 [synonym]
Fm. Cyclaxyridae Gimmel et al. 2009
+
G. Cyclaxyra Broun 1893
G. †Electroxyra Gimmel et al. 2019
G. †Neolitochropus Lyubarsky and Perkovsky 2016
+
Invalid names: Olibrolitus Lyubarsky and Perkovsky 2020 [synonym]
G. †Pacyclaxyra Tihelka et al. 2021
Fm. Hobartiidae Sen Gupta and Crowson 1966
+
Invalid names: Hobartiinae Sen Gupta and Crowson 1969 [empty]
Fm. Laemophloeidae Ganglbauer 1899
+
G. Laemophloeus Dejean 1835
G. Leptophloeus Casey 1916
G. †Lithocoryne Scudder 1900
G. Parandrita LeConte and Horn 1883
G. Placonotus MacLeay 1871
Invalid names: Laemophloeinae [invalid subgroup], Propalticidae Crowson 1952 [synonym]
Subfm. †Meligethiellinae Kirejtshuk and Ponomarenko 1990
+
G. †Meligethiella Medvedev 1969
G. †Ostomalynus Kirejtshuk and Ponomarenko 1990
Fm. Passandridae Blanchard 1845 [parasitic flat bark beetle]
+
Subfm. †Mesopassandrinae Jin et al. 2019
Subfm. Passandrinae Blanchard 1845
Fm. Phalacridae Leach 1815
+
Subfm. Phalacrinae Leach 1815
Invalid names: Phaenocephalinae Matthews 1899 [empty]
Fm. Phloeostichidae Reitter 1911
+
G. †Pleuroceratos Poinar and Kirejtshuk 2008
Invalid names: Hymaeinae Sen Gupta and Crowson 1866 [empty], Phloeostichinae Reitter 1911 [empty]
Fm. Silvanidae Kirby 1837
+
Subfm. Brontinae Blanchard 1845
Subfm. Silvaninae Kirby 1837
Invalid names: Hyliotinae [empty]
Invalid names: Agapythidae Sen Gupta and Crowson 1969 [empty], Brachypteridae Erichson 1845 [empty], Cavognathidae Sen Gupta and Crowson 1966 [empty], Clavicornia [invalid subgroup], Heteromera [invalid subgroup], Lamingtoniidae Sen Gupta and Crowson 1969 [empty], Myraboliidae Lawrence and Britton 1991 [empty], Priasilphidae Crowson 1973 [empty], Sinisilvanidae Hong 2002 [invalid subgroup], Tasmosalpingidae Lawrence and Britton 1991 [empty]
Superfm. Erotyloidea Latreille 1802
+
Fm. Boganiidae Sen Gupta and Crowson 1966
+
Subfm. Boganiinae Sen Gupta and Crowson 1966
G. †Palaeoboganium Liu et al. 2017
Invalid names: Paracucujinae Endrody-Younga and Crowson 1986 [empty]
Fm. Erotylidae Latreille 1802 [pleasing fungus beetle]
+
Subfm. Cryptophilinae Casey 1900
Subfm. Erotylinae Latreille 1802
Subfm. Languriinae Crotch 1873
Subfm. Pharaxonothinae
Subfm. Xenoscelinae Ganglbauer 1899
Invalid names: Dacninae Gistel 1856 [empty], Encaustinae Crotch 1876 [empty], Loberinae [empty], Megalodacninae Sen Gupta 1969 [empty]
Superfm. Lymexyloidea Fleming 1821
+
Fm. Lymexylidae Fleming 1821 [ship-timber beetle]
+
Subfm. Atractocerinae Laporte 1840
Subfm. Hylecoetinae Germar 1818
Subfm. Lymexylinae Fleming 1821
Subfm. Melittommatinae Wheeler 1986
Superfm. Nitiduloidea Latreille 1802
+
Fm. Helotidae Reitter 1876
+
G. †Burmahelota Liu et al. 2019
G. †Lobatihelota Li et al. 2023
G. Metahelotella Kirejtshuk 2000
G. †Palaeohelota Liu et al. 2019
G. †Trihelota Tihelka et al. 2020
Fm. Kateretidae Kirby 1837
+
G. Amartus Le Conte 1861
Subfm. †Antirhelinae Kirejtshuk 2023
G. Brachypterolus Grouvelle 1913
G. Cercus Latreille 1796
G. †Cretaretes Peris and Jelínek 2019
G. †Electrumeretes Peris and Jelínek 2019
G. †Eoceniretes Kirejtshuk and Nel 2008
G. †Furcalabratum Poinar and Brown 2017
G. Heterhelus Jacquelin du Val 1858
G. Kateretes Herbst 1793
G. †Lebanoretes Kirejtshuk and Azar 2008
G. †Pelretes Tihelka et al. 2021
G. †Polliniretes Peris and Jelínek 2019
G. †Protokateretes Zhao et al. 2023
G. †Scaporetes Zhao et al. 2023
Fm. Monotomidae Laporte 1840 [root eating beetle]
+
G. †Jurorhizophagus Cai et al. 2015
Subfm. Monotominae Laporte 1840
Subfm. Rhizophaginae Redtenbacher 1845
Invalid names: Rhizophagidae Redtenbacher 1845 [synonym]
Fm. Nitidulidae Latreille 1802 [sap beetle]
+
Subfm. Carpophilinae Erichson 1842
Subfm. Cillaeinae Kirejtshuk and Audisio 1986
Subfm. Cryptarchinae Thomson 1859
Subfm. Cybocephalinae Jacquelin du Val 1858
Subfm. Epuraeinae Kirejtshuk 1986
Subfm. Meligethinae Thomson 1859
G. †Miophenolia Wickham 1916
G. †Nitidulina Martynov 1926
Subfm. Nitidulinae Latreille 1802
G. †Nitidulites Heer 1864
G. †Oligamphotis Theobald 1937
Subfm. Prometopiinae Böving and Craighead 1931
G. †Protonitidula Zhao et al. 2022
G. †Sinonitidulina Hong 1983
Invalid names: Calonecrinae Kirejtshuk 1982 [empty], Cateretinae Kirby 1837 [empty]
Fm. Smicripidae Horn 1879 [palmetto beetle]
+
G. †Mesosmicrips Kirejtshuk 2017
G. Smicrips LeConte 1878
Fm. Sphindidae Jacquelin du Val 1860 [cryptic slime mold beetle]
+
Subfm. †Libanopsinae Kirejtshuk 2015
Subfm. Protosphindinae Sen Gupta and Crowson 1979
Subfm. Sphindinae Jacquelin du Val 1860
G. †Trematosphindus Li and Cai 2021
Invalid names: Aspidiphoridae Kiesenwetter 1877 [synonym], Odontosphindinae Sen Gupta and Crowson 1979 [empty], Sphindiphorinae Sen Gupta and Crowson 1979 [empty]
Fm. †Parandrexidae Kirejtshuk 1994
+
Subfm. †Parandrexinae Kirejtshuk 1994
+
G. †Martynopsis Soriano et al. 2006
G. †Parandrexis Martynov 1926
Unr. Phytophaga
+
Superfm. Chrysomeloidea Latreille 1802
+
Fm. Bruchidae Latreille 1802
Fm. Cerambycidae Latreille 1802 [longhorned beetle]
+
Invalid names: Pseudonepidae Jordan 1967 [synonym], Spondylididae Audinet-Serville 1832 [synonym]
Fm. Chrysomelidae Latreille 1802 [leaf beetle]
Fm. Disteniidae Thomson 1860
Fm. Megalopodidae Latreille 1802
Fm. Orsodacnidae Thomson 1859
G. †Paloedemera Wickham 1914
Fm. Vesperidae Mulsant 1839
Invalid names: Cerambycoidea Latreille 1802 [synonym], Oxypeltidae Lacordaire 1868 [empty]
Superfm. Curculionoidea Latreille 1802 [weevil]
+
Fm. Anthribidae Billberg 1820 [fungus weevil]
Fm. Apionidae Schoenherr 1823
Fm. Attelabidae Billberg 1820 [leaf rolling weevil]
+
Invalid names: Rhynchitidae Gistel 1848 [synonym]
Fm. Belidae Schoenherr 1826
Fm. Caridae Thompson 1992
Fm. Cimberididae de Gozis 1882
Fm. Curculionidae Latreille 1802 [snout beetle]
+
Invalid names: Brachyceridae Billberg 1820 [synonym], Dryophthoridae Schoenherr 1825 [synonym]
Fm. †Mesophyletidae Poinar 2008
Fm. Nemonychidae Bedel 1882 [pine flower snout beetle]
Fm. †Obrieniidae Zherikhin and Gratshev 1993
Fm. †Ulyanidae Zherikhin 1993
Invalid names: Akulosamphus Aymard 1856 [nomen nudum], Brenthidae Billberg 1820 [empty], Erirhinidae Schoenherr 1825 [empty], Loricera glacialis Scudder 1877 [nomen dubium]
Superfm. Tenebrionoidea Latreille 1802
+
Fm. Aderidae Csiki 1909
+
Subfm. Aderinae Csiki 1909
G. Cnopus Champion 1893
G. Escalerosia Nardi 2007
G. †Palaeocnopus Alekseev and Grzymala 2015
G. Picemelinus Nakane 1987
G. Vanonus Casey 1895
Invalid names: Circaeidae Iablokoff-Khnzorian 1961 [synonym], Xylophilidae [replaced]
Fm. Anthicidae Latreille 1819 [ant like flower beetle]
+
Subfm. Anthicinae Latreille 1819
Subfm. Eurygeniinae LeConte 1862
Subfm. Lemodinae Lawrence and Britton 1991
Subfm. Macratriinae LeConte 1862
Subfm. Steropinae Jacquelin du Val 1863
Subfm. Tomoderinae Bonadona 1961
Invalid names: Copobaeninae Abdullah 1969 [empty]
Fm. Boridae Thomson 1859
+
Invalid names: Borinae Thomson 1859 [empty], Synercticinae Lawrence and Pollock 1994 [empty]
Fm. Ciidae Leach 1819 [minute tree-fungus beetle]
+
Subfm. Ciinae Leach 1819
Invalid names: Sphindociinae Lawrence 1974 [empty]
Subfm. Ischaliidae Blair 1920 [false fire-colored beetle]
+
G. Ischalia Pascoe 1860
G. †Prototrichalus Molina-Olmedo et al. 2020
G. †Kulindrobor Tihelka et al. 2021
+
†Kulindrobor enigmaticus Tihelka et al. 2021
†Kulindrobor magnus Tihelka et al. 2021
Fm. Melandryidae Leach 1815 [false darkling beetle]
+
G. †Cicindelopsis Cockerell 1921
Subfm. Melandryinae Leach 1815
Invalid names: Osphyinae Mulsant 1856 [empty]
Fm. Meloidae Gyllenhal 1810 [blister beetle]
+
Subfm. Meloinae Gyllenhal 1810
G. †Microbasis Martins-Neto 1998
Subfm. Nemognathinae Laporte 1840
Invalid names: Eleticinae Wellman 1910 [empty]
Fm. Mordellidae Latreille 1802 [tumbling flower beetle]
+
Subfm. †Apotomourinae Bao et al. 2018
G. Falsomordellistena Ermisch 1941
G. Glipostena Ermisch 1941
G. Mordellaria Ermisch 1950
G. †Mordellina Germar 1813
Subfm. Mordellinae Latreille 1802
G. †Petrimordella Batelka et al. 2023
Subfm. †Praemordellinae Shchegoleva-Barovskaya 1929
Invalid names: Ctenidiinae Franciscolo 1951 [empty], Praemordellidae Shchegoleva-Barovskaya 1929 [synonym]
Fm. Mycetophagidae Leach 1815 [hairy fungus beetle]
+
Subfm. Mycetophaginae Leach 1815
Invalid names: Bergininae Leng 1920 [empty], Esarcinae Reitter 1882 [empty]
Fm. Mycteridae Oken 1843 [palm and flower beetle]
+
Subfm. Mycterinae Oken 1843
Invalid names: Hemipeplinae Lacordaire 1854 [empty]
Fm. Oedemeridae Latreille 1810 [false blister beetle]
+
Subfm. Calopodinae Costa 1852
Subfm. Oedemerinae Latreille 1810
Fm. Prostomidae Thomson 1859 [jugular-horned beetle]
+
G. Prostomis Latreille 1825
G. †Vetuprostomis Engel and Grimaldi 2008
Fm. Pyrochroidae Latreille 1807
+
G. †Palaeopyrochroa Abdullah 1965
Subfm. Pedilinae Lacordaire 1859
Subfm. Pyrochroinae Latreille 1807
Subfm. †Waidelotinae Bukejs et al. 2019
Invalid names: Cretaceimelittomoides Vulcano and Pereira 1987 [nomen nudum], Cretaceimelittomoides cearensis Vulcano and Pereira 1987 [nomen nudum], Pilipalpinae Abdullah 1964 [empty], Tydessinae Nikitsky 1986 [empty]
Fm. Rhipiphoridae Gemminger 1870 [wedge shaped beetle]
+
G. Rhipiphorus Bosc 1792
Invalid names: Rhipidiinae Gerstaecker 1855 [empty], Rhipiphorinae Gemminger 1870 [empty]
Fm. Ripiphoridae Gemminger 1870 [wedge shaped beetle]
+
Subfm. Ripidiinae Gerstaecker 1855
Subfm. Ripiphorinae Gemminger 1870
G. †Salignacicola Batelka and Prokop 2021
Invalid names: Hemirhipidiinae Heller 1921 [empty], Ptilophorinae Gerstaecker 1855 [empty]
Fm. Salpingidae Leach 1815 [narrow-waisted bark beetle]
+
G. †Arra Peris et al. 2014
Subfm. Inopeplinae Grouvelle 1908
Subfm. Prostominiinae Grouvelle 1914
Subfm. Salpinginae Leach 1815
Invalid names: Aegialitinae LeConte 1862 [empty], Dacoderinae LeConte 1862 [empty], Othniinae LeConte 1861 [empty]
Fm. Scraptiidae Gistel 1848 [false flower beetle]
+
Subfm. Anaspidinae Mulsant 1856
Subfm. Scraptiinae Gistel 1848
G. †Scraptiomima Medvedev 1969
Fm. Stenotrachelidae Thomson 1859
+
Invalid names: Cephaloinae LeConte 1862 [empty], Nematoplinae LeConte 1862 [empty], Stenotrachelinae Thomson 1859 [empty], Stoliinae Nikitsky 1985 [empty]
Fm. Tenebrionidae Latreille 1802 [darkling beetle]
+
Subfm. Alleculinae Laporte 1840 [comb-clawed beetle]
Subfm. Blaptinae Leach 1815
Subfm. Diaperinae Latreille 1802
G. †Elytrophagus Zhang 1989
G. †Eoallognosis Haupt 1950
G. †Eocallidium Haupt 1950
G. †Eohelaeus Haupt 1950
Subfm. Lagriinae Latreille 1825
G. †Megalocera Hope 1842
Subfm. Nilioninae Oken 1843 [false ladybird beetle]
G. †Paropiophorus Haupt 1950
Subfm. Phrenapatinae Solier 1834
Subfm. Pimeliinae Latreille 1802
G. †Protoplatycera Wickham 1914
G. †Rhinohelaeites Haupt 1950
Subfm. Stenochiinae Kirby 1837
G. †Tagenopsis Heer 1864
Subfm. Tenebrioninae Latreille 1802
G. †Tenebrionites Cockerell 1920
Subfm. Zolodininae Watt 1874
Invalid names: Ancylochira oligostriata Piton and Théobald 1937 [nomen dubium], Tenebrio calculensis Scudder 1895 [nomen dubium]
Fm. Tetratomidae Billberg 1820 [polypore fungus beetle]
+
Subfm. Eustrophinae Gistel 1848
Subfm. Hallomeninae Gistel 1848
Subfm. Tetratominae Billberg 1820
Invalid names: Penthinae Lacordaire 1859 [empty], Piseninae Miyatake 1960 [empty]
Fm. Ulodidae Pascoe 1869
+
G. †Archaeophylax Leschen and Rhode 2002
G. †Waitomophylax Leschen and Rhode 2005
Fm. Zopheridae Solier 1834 [ironclad beetle]
+
Subfm. Colydiinae Billberg 1820
G. †Cretomysteria Deng et al. 2017
G. Syntarsus Fairmaire 1869
Subfm. Zopherinae Solier 1834
Invalid names: Colydiidae Billberg 1820 [replaced], Usechinae Horn 1867 [empty]
Invalid names: Afreminae Levey 1985 [empty], Archeocrypticidae Kaszab 1964 [empty], Asiamordella Hong 2002 [nomen nudum], Chalcodryidae Watt 1974 [empty], Cossyphodinae Wasmann 1899 [empty], Eurygenius wickhami Cockerell 1917 [nomen dubium], Fushunicucujus Hong 2002 [nomen nudum], Lagrioididae Abdullah and Abdullah 1968 [empty], Perimylopidae St. George 1939 [empty], Pterogeniidae Crowson 1953 [empty], Trachelostenidae Lacordaire 1859 [empty], Trictenotomidae Blanchard 1845 [empty]
Infraor. Elateriformia Crowson 1960
+
Superfm. Buprestoidea Leach 1815
+
Fm. Buprestidae Leach 1815 [jewel beetle]
+
Subfm. Agrilinae Laporte 1835
G. †Agrilium Westwood 1854
G. †Andakhudukia Alexeev 2008
G. †Archelater Carpenter 1992
Subfm. Buprestinae Leach 1815
Subfm. Chrysochroinae Laporte 1835
G. Coroebus Gory and Laporte 1839
G. †Ctenicerium Westwood 1854
G. †Dictyorachys Li et al. 2023
G. †Eurynucha Handlirsch 1906
G. †Eurythyreites Handlirsch 1906
Subfm. Julodinae Lacordaire 1857
G. †Macronotus Hong and Wang 1990
G. †Mesostigmodera Etheridge and Olliff 1890
Subfm. †Parathyreinae Alekseev 1994
G. †Planocoleus Hong 1982
Subfm. Polycestinae Lacordaire 1857
G. †Proctobuprestis Handlirsch 1906
Invalid names: Ancylochira pinguis Piton 1940 [nomen dubium], Ancylochira striatopunctata Piton 1940 [nomen dubium], Ancylochira violaceocyanescens Piton 1940 [nomen dubium], Chrysobothrites Heer 1864 [nomen nudum], Chrysochroa punctata Piton 1940 [nomen dubium], Iridotaenia hovassei Piton 1940 [nomen dubium], Stigmodera unicolor Piton 1940 [nomen dubium]
Fm. Schizopodidae LeConte 1859
+
G. †Mesoschizopus Cai et al. 2015
Subfm. Schizopodinae LeConte 1861
Invalid names: Mesohelophorus mongolicus Ponomarenko 1986 [nomen dubium]
Superfm. Byrrhoidea Latreille 1804
+
Fm. Byrrhidae Latreille 1804 [pill beetle]
+
G. †Amphicyrtella Ponomarenko 1990
Subfm. Amphicyrtinae LeConte 1861
Subfm. Byrrhinae Latreille 1804
G. †Byrrhydium Heer 1864
G. †Fangshanella Huang and Zhang 1997
G. Lamprobyrrhulus Ganglbauer 1902
Subfm. †Lidryopinae Kirejtshuk and Nel 2016
G. Lioligus Casey 1912
G. †Mesobyrrhus Huang and Zhang 1997
G. †Mesosimplocaria Ponomarenko 1990
G. †Nosotetocus Scudder 1892
G. Pelochares Mulsant and Rey 1869
Subfm. Syncalyptinae Mulsant and Rey 1869
Invalid names: Nosodendron tritavum Scudder 1890 [nomen dubium]
Invalid names: Chelonarium montanum Wickham 1914 [nomen dubium]
Superfm. Dascilloidea Gue'rin-Me'neville 1843
+
Fm. Dascillidae Guerin-Meneville 1843 [soft bodied plant beetle]
+
Subfm. Dascillinae Guerin-Meneville 1843
Subfm. Karumiinae Escalera 1913
G. †Lyprodascillus Zhang 1989
Invalid names: Karumiidae Escalera 1913 [empty]
Superfm. Dryopoidea
+
Fm. Dryopidae Billberg 1820 [long toed water beetle]
+
G. Helichus Erichson 1847
G. Helmis Latreille 1802
G. Pomatinus Sturm 1853
Invalid names: Parnidae Leach 1817 [replaced]
Fm. Elmidae Curtis 1830 [riffle beetle]
+
G. Cleptelmis Sanderson 1953
G. †Cretohypsilara Cai et al. 2018
Subfm. Elminae Curtis 1830
G. Heterelmis Sharp 1882
Invalid names: Larainae LeConte 1861 [empty]
Fm. Eulichadidae Crowson 1973 [forest stream beetle]
+
Subfm. Eulichadinae Crowson 1973
Invalid names: Lasiosyne punctata Yan 2012 [nomen dubium], Lasiosyne shartegiensis Yan 2012 [nomen dubium]
Fm. Heteroceridae MacLeay 1825 [Variegated mud-loving beetle]
+
G. †Excavotarsus Li et al. 2020
Subfm. Heterocerinae MacLeay 1825
G. †Heterocerites Ponomarenko 1986
G. Lanternarius Pacheco 1964
G. Lapsus Pacheco 1964
Invalid names: Elythomerinae Pacheco 1964 [empty]
Fm. Limnichidae Erichson 1846 [minute marsh loving beetle]
+
Subfm. Cephalobyrrhinae Champion 1925
Subfm. Limnichinae Erichson 1846
G. †Palaeoersachus Pütz et al. 2004
G. Platypelochares Champion 1923
Invalid names: Hyphalinae Britton 1971 [empty], Thaumastodinae Champion 1924 [empty]
Fm. †Mastigocoleidae Tihelka et al. 2022
+
G. †Cretaceocoleus Tihelka and Cai 2022
G. †Mastigocoleus Tihelka and Cai 2022
Fm. Psephenidae Lacordaire 1854 [water penny beetle]
+
Subfm. Eubrianacinae Jacobson 1913
Subfm. Eubriinae Lacordaire 1857
G. †Protacnaeus Wickham 1914
G. †Psephenella Martins-Neto 1998
Subfm. Psepheninae Lacordaire 1854
Subfm. Psephenoidinae Hinton 1939
G. †Pseudomataeopsephus Li and Cai 2021
Fm. Ptilodactylidae Laporte 1836
+
Subfm. Anchytarsinae Champion 1897
G. †Aphebodactyla Chatzimanolis et al. 2012
Subfm. Cladotominae Pic 1914
Subfm. Ptilodactylinae Laporte 1836
G. †Ptilodactyloides Motschulsky 1856
Invalid names: Aploglossinae Champion 1897 [empty], Araeopidiinae Lawrence 1991 [empty]
Invalid names: Actinocerites Hong 2002 [nomen nudum], Brachypodites Hong 2002 [nomen nudum], Callirhipidae Emden 1924 [empty], Cneoglossidae Champion 1897 [empty], Expansifemoria Hong 2002 [nomen nudum]
Superfm. Elateroidea Leach 1815
+
Fm. Artematopodidae Lacordaire 1857 [soft bodied plant beetle]
+
G. †Bipogonia Li et al. 2022
G. †Carinibipogonia Li et al. 2022
Subfm. Electribiinae Crowson 1975
G. †Forticatinius Tan and Ren 2007
G. †Protartematopus Crowson 1973
G. †Sinobrevipogon Cai et al. 2015
G. †Tarsomegamerus Zhang 2005
G. †Valdopogon Tihelka and Cai 2023
Invalid names: Artematopodinae Lacordaire 1857 [empty], Eurypogonidae [replaced], Notocupes undatabdominus Lin 1980 [nomen dubium]
Fm. †Berendtimiridae Winkler 1987
+
Subfm. †Berendtimirinae Winkler 1987
Subfm. †Retromalisinae Fanti et al. 2023
Fm. Brachypsectridae LeConte and Horn 1883 [Texas beetle]
+
G. Brachypsectra Le Conte 1874
G. †Cretopsectra Zhao et al. 2020
G. †Hongipsectra Tihelka et al. 2019
G. †Vetubrachypsectra Qu and Cai 2019
Fm. Cantharidae Imhoff 1856 [soldier beetle]
+
Subfm. Cantharinae Imhoff 1856
Subfm. Chauliognathinae LeConte 1861
G. †Cretocantharis Hsiao et al. 2021
G. †Katyacantharis Kazantsev and Perkovsky 2019
Subfm. Malthininae Kiesenwetter 1852
G. †Myamalycocerus Fanti and Ellenberger 2016
G. †Ornatomalthinus Poinar and Fanti 2016
Subfm. Silinae Mulsant 1862
Invalid names: Cantharidiae Latreille 1802 [replaced], Dysmorphocerinae Brancucci 1980 [empty], Telephoridae [replaced]
Fm. Cerophytidae Latreille 1834
+
G. †Amberophytum Yu et al. 2019
G. †Aphytocerus Zherikhin 1977
G. †Baissophytum Chang et al. 2011
G. †Baissopsis Kirejtshuk 2013
G. Brachycerophytum Costa et al. 2003
G. Cerophytum Latreille 1809
G. †Elaterocoleus Dolin 1973
G. †Jurassophytum Yu et al. 2019
G. †Necromera Martynov 1926
+
Invalid names: Idiomerus Dolin 1980 [synonym], Leptocnemus Hong and Wang 1990 [synonym]
G. †Necromeropsis Yu et al. 2019
G. †Praelaterium Dolin 1973
G. †Wongaroo Oberprieler et al. 2016
Invalid names: Aphytocerinae Kirejtshuk 2013 [invalid subgroup], Praelateriidae Dolin 1973 [synonym]
Fm. Elateridae Leach 1815 [click beetle]
+
G. †Adocetus Scudder 1900
Subfm. Agrypninae Candèze 1857
G. †Artinama Lin 1986
G. †Bilineariselater Chang and Ren 2008
Subfm. Cardiophorinae Candèze 1859
G. †Cretocardiophorus Qiu and Ruan 2023
G. †Cretoelaterium Alexeev 2008
G. †Cryptagriotes Wickham 1916
G. †Cryptocoelus Dolin and Nel 2002
Subfm. Dendrometrinae Gistel 1848
Subfm. Denticollinae Stein and Weise 1877
Subfm. Elaterinae Leach 1815
G. †Elaterites Heer 1847
G. †Elaterium Westwood 1854
G. †Gripecolous Lin 1986
Subfm. Hypnoidinae Schwarz 1906
Subfm. Lissominae Laporte 1835
G. †Ludiophanes Wickham 1916
G. †Mercata Lin 1986
G. †Mionelater Becker 1963
Subfm. Negastriinae Nakane and Kishii 1956
G. †Ovivagina Zhang 1997
G. †Paralithomerus Chang et al. 2008
Subfm. Pityobiinae Hyslop 1917
Subfm. †Protagrypninae Dolin 1973
G. †Protocardiophorus Dolin 1976
G. †Pseudocardiophorites Dolin 1976
G. †Silicernius Von Heyden 1859
G. †Sinoelaterium Ping 1928
G. Spheniscosomus Schwarz 1892
G. †Tetraraphes Yablokov-Khnzorian 1961
G. †Turonelater Alexeev 2011
Invalid names: Acmaeodera burmitina Cockerell 1917 [nomen dubium], Cebrionidae Latreille 1802 [replaced], Cebrioninae Latreille 1802 [empty], Elater maculatus Gistl 1831 [nomen dubium], Hemiopinae Fleutiaux 1941 [empty], Omalisinae Lacordaire 1857 [empty], Physodactylinae Lacordaire 1857 [empty], Subprotelaterinae Fleutiaux 1920 [empty], Tetralobinae Laporte 1840 [empty]
Fm. Eucnemidae Eschscholtz 1829 [false click beetle]
+
Subfm. Anischiinae Fleutiaux 1936
G. †Beattieellus Oberprieler et al. 2016
G. †Cretopoena Alexeev 2008
Subfm. Eucneminae Eschscholtz 1829
G. †Fiegelia Muona 2020
Subfm. Macraulacinae Fleutiaux 1923
Subfm. Melasinae Fleming 1821
Subfm. Palaeoxeninae Muona 1993
Subfm. Schizophilinae Muona 1993
Invalid names: Perothopinae Lacordaire 1857 [empty], Phlegoninae Muona 1993 [empty], Phyllocerinae Reitter 1905 [empty], Pseudomeninae Muona 1993 [empty]
Fm. Lampyridae Rafinesque 1815
+
Subfm. Lamprohizinae Kazantsev 2010
Subfm. Lampyrinae Rafinesque 1815
Subfm. Luciolinae Lacordaire 1857
Invalid names: Amydetinae Olivier 1907 [empty], Cyphonocerinae Crowson 1972 [empty], Photurinae Lacordaire 1857 [empty], Pterotinae LeConte 1861 [empty]
Fm. Lycidae Laporte 1836 [net-winged beetle]
+
Subfm. Dexorinae Bocak and Bocakova 1989
Subfm. Lycinae Laporte 1836
G. †Miocaenia Wickham 1914
Invalid names: Ateliinae Kleine 1928 [empty], Calochrominae Lacordaire 1857 [empty], Leptolycinae Leng and Mutchler 1922 [empty], Metriorrhynchinae Kleine 1926 [empty]
Fm. †Mysteriomorphidae Alekseev and Ellenberger 2019
+
G. †Mysteriomorphus Alekseev and Ellenberger 2019
+
Invalid names: Trapezioceps Qu et al. 2023 [synonym]
Fm. Omethidae Leconte 1861
+
G. †Electromethes Kazantsev 2012
Invalid names: Driloniinae Crowson 1972 [empty], Matheteinae LeConte 1881 [empty], Omethinae LeConte 1861 [empty], Telegeusinae Leng 1920 [empty]
Fm. Throscidae Laporte 1840
+
G. †Archaeolus Lin 1986
G. †Captopus Li and Cai 2021
G. †Electrothroscus Li and Cai 2021
G. †Jaira Muona 1993
G. Pactopus Leconte 1868
G. †Potergosoma Kovalev and Kirejtshuk 2013
G. Potergus de Bonvouloir 1871
G. †Pseudopactopus Li and Cai 2021
G. †Pseudothroscus Muona 2019
G. †Rhomboaspis Kovalev and Kirejtshuk 2013
G. †Trixagosoma Li et al. 2020
G. Trixagus Kugelann 1794
+
Invalid names: Palaeothroscus Yablokov-Khnzorian 1962 [synonym], Throscus Latreille 1796 [synonym]
G. †Tyrannothroscus Muona 2019
Invalid names: Megacephalites Hong 2002 [nomen nudum], Palaeothroscini Yablokov-Khnzorian 1962 [invalid subgroup], Throscinae Laporte 1840 [empty], Trixagidae Gistel 1848 [synonym]
Invalid names: Artematopoidea Lacordaire 1857 [synonym], Eudicronychinae Girard 1971 [empty], Rhagophthalmidae Olivier 1907 [empty]
Invalid names: Mesosperchus angulatus Ponomarenko 1985 [nomen dubium], Podabrocephalidae Pic 1930 [empty]
G. †Jingxidiscus Hong 1990
+
†Jingxidiscus lushangfenensis Hong 1981
Invalid names: Discus Hong 1981 [replaced]
Fm. Jurodidae Ponomarenko 1985
+
G. †Jurodes Ponomarenko 1985
+
†Jurodes daohugouensis Yan et al. 2014
†Jurodes ignoramus Ponomarenko 1985
†Jurodes minor Ponomarenko 1990
†Jurodes pygmaeus Yan et al. 2014
†Jurodes shartegiensis Yan 2014
†Jurodes shef Yan et al. 2021
Invalid names: Sikhotealiniidae Lafer 1996 [synonym]
G. †Larvula Ponomarenko 1985
+
†Larvula cassa Ponomarenko 1985
†Larvula triassica Prokin and Ponomarenko 2013
Fm. †Lasiosynidae Kirejtshuk et al. 2010
+
G. †Anacapitis Yan 2009
+
†Anacapitis incertus Yan 2009
†Anacapitis karataviensis Yan 2009
†Anacapitis oblongus Yan 2009
†Anacapitis plata Tan and Ren 2009
Invalid names: Brachysyne Tan and Ren 2009 [synonym]
G. †Glaphyropterodes Handlirsch 1906
+
†Glaphyropterodes gehreti Heer 1852
†Glaphyropterodes sibiricus Ponomarenko 1985
G. †Lasiosyne Tan et al. 2007
+
†Lasiosyne cataphracta Yan 2012
†Lasiosyne daohugouensis Kirejtshuk et al. 2010
†Lasiosyne decora Yan 2012
†Lasiosyne euglyphea Tan et al. 2007
+
Invalid names: Pappisyne eucallus Tan and Ren 2009 [synonym], Pappisyne lasiospatha Tan and Ren 2009 [synonym]
†Lasiosyne fedorenkoi Kirejtshuk et al. 2010
†Lasiosyne gratiosa Kirejtshuk et al. 2010
†Lasiosyne insculpta Yan 2012
†Lasiosyne integera Guo et al. 2022
†Lasiosyne longitarsus Yan 2012
†Lasiosyne multituberata Lin 1980
†Lasiosyne parva Guo et al. 2022
†Lasiosyne quadricollis Kirejtshuk et al. 2010
Invalid names: Pappisyne Tan and Ren 2009 [synonym]
Invalid names: Lasiosyne laxa Yan et al. 2014 [nomen dubium]
Unr. Nosodendriformia Cai and Tihelka 2022
+
Superfm. Nosodendroidea Erichson 1846
+
Fm. Nosodendridae Erichson 1846 [wounded tree beetle]
+
G. †Archaenosodendron Li and Cai 2021
G. †Basinosa Tihelka et al. 2020
G. †Mesonosa Tihelka et al. 2021
G. Nosodendron Latreille 1804
G. †Plastelater Handlirsch 1906
+
†Plastelater glabratus Ponomarenko 1985
†Plastelater neptuni Giebel 1856
G. †Polysitum Dunstan 1923
+
†Polysitum elongatum Ponomarenko 1985
†Polysitum hosbayari Ponomarenko 1986
†Polysitum kuznetskiense Rohdendorf 1961
†Polysitum latum Ponomarenko 1985
†Polysitum majus Ponomarenko 1990
†Polysitum minutus Dunstan 1923
†Polysitum ovale Ponomarenko 1990
†Polysitum punctatus Dunstan 1923
†Polysitum rugosum Ponomarenko 1986
†Polysitum sharategense Ponomarenko 2014
†Polysitum wudenghaoensis Hong 1976
Unr. Rhinorhipiformia Cai et al. 2022
+
Unr. Scirtiformia Lawrence et al. 2011
+
Superfm. Scirtoidea Fleming 1821
+
Fm. Scirtidae Fleming 1821 [marsh beetle]
+
Subfm. †Mesernobiinae Engel 2010
G. †Miocyphon Wickham 1914
G. †Molinernobius Molino-Olmedo 2017
Subfm. Scirtinae Fleming 1821
+
Invalid names: Cyphoninae Stephens 1829 [synonym], Helodinae Shuckard 1839 [synonym]
Invalid names: Elodiidae Shuckard 1839 [synonym], Sinodryopitidae Hong 2002 [invalid subgroup]
Invalid names: Decliniidae Nikitsky et al. 1994 [empty]
G. †Sphaerocantharis Bode 1953
+
†Sphaerocantharis defossa Bode 1953
†Sphaerocantharis sibirica Ponomarenko 1990
Invalid names: Mikrocarpides Bode 1953 [synonym], Theornithion Bode 1953 [synonym]
Infraor. Staphyliniformia Lameere 1900
+
Superfm. Histeroidea Gyllenhal 1808
+
Fm. Histeridae Gyllenhal 1808 [clown beetle]
+
Subfm. Abraeinae MacLeay 1819
G. †Amplectister Caterino and Maddison 2018
Subfm. †Antigracilinae Zhou et al. 2020
G. Chaetabraeus Portevin 1929
Subfm. Dendrophilinae Reitter 1909
Subfm. Haeteriinae Marseul 1857
Subfm. Histerinae Gyllenhal 1808
Subfm. Onthophilinae MacLeay 1819
G. Plegaderus Erichson 1834
Subfm. Saprininae Blanchard 1845
Subfm. Trypanaeinae Marseul 1857
Invalid names: Chlamydopsinae Bickhardt 1914 [empty], Hetaeriinae Marseul 1857 [empty], Niponiinae Fowler 1912 [empty], Tribalinae Bickhardt 1914 [empty], Trypeticinae Bickhardt 1914 [empty]
Invalid names: Sphaeritidae Shuckard 1839 [empty]
Superfm. Hydrophiloidea Latreille 1802 [water scavenger beetle]
+
Fm. Hydrophilidae Latreille 1802 [water scavenger beetle]
+
G. †Alegorius Fikáček et al. 2014
G. †Aposphinctus Bode 1953
+
Invalid names: Hexameristus Bode 1953 [synonym], Spalacoides Bode 1953 [synonym], Streblocardioides Bode 1953 [synonym]
Subfm. Chaetarthriinae Bedel 1881
G. †Creniphilites Wickham 1913
G. †Cretosperchus Ponomarenko 1997
Subfm. Enochrinae Short and Fikáček 2013
Subfm. Georissinae Laporte 1819
G. Hydrocharis Berthold 1827
Subfm. Hydrochinae Thomson 1859
Subfm. Hydrophilinae Latreille 1802
G. †Orphnospercheus Lin 1992
G. †Paraspercheus Ponomarenko 1977
G. †Penitus Zhang et al. 1994
G. Philhydrus Solier 1834
G. †Prospercheus Prokin 2009
G. †Protochares Fikáček et al. 2014
G. Regimbartia Zaitzev 1908
Subfm. Rygmodinae Orchymont 1916
+
Invalid names: Andotypini Hansen 1991 [synonym], Borborophorini Hansen 1991 [synonym], Tormissini Hansen 1991 [synonym]
Subfm. Sphaeridiinae Latreille 1802
G. Sternolophus Solier 1834
Invalid names: Escheria Heer 1847 [nomen dubium], Escheria bella Heer 1862 [nomen dubium], Escheria ovalis Heer 1847 [nomen dubium], Escheriidae Kolbe 1880 [synonym], Mesosperchus schultzi Ponomarenko 1985 [nomen dubium], Spercheinae Erichson 1837 [empty], Sphaeridium melanarium Gistl 1831 [nomen dubium]
G. †Laetopsia Fikáček et al. 2012
+
†Laetopsia baissensis Ponomarenko 1987
†Laetopsia bontsaganica Prokin 2009
†Laetopsia hydraenoides Prokin et al. 2010
†Laetopsia mongolica Ponomarenko 1987
†Laetopsia shatrovskiyi Prokin et al. 2010
Invalid names: Epimetopidae Zaitzev 1908 [empty], Georissidae Laporte 1819 [empty]
Superfm. Scarabaeoidea Latreille 1802 [scarab beetle]
+
Fm. Bolboceratidae Mulsant 1842 [dor beetle]
+
Subfm. Athyreinae Lynch Arribálzaga 1878
Subfm. Bolboceratinae Mulsant 1842
G. †Cretobolbus Nikolajev 1996
Fm. Geotrupidae Latreille 1802 [earth-boring dung beetle]
+
Subfm. Geotrupinae Latreille 1802
G. †Orrhodomala Zhang 1989
G. Typhaeus Leach 1815
Invalid names: Lethrinae Mulsant and Rey 1871 [empty], Taurocerastinae [empty]
G. †Geotrupoides Handlirsch 1906
+
†Geotrupoides jiaoheense Hong 1992
†Geotrupoides kezuoensis Hong 1992
†Geotrupoides lithographicus Deichmüller 1886
†Geotrupoides saxosus Zhang 1997
†Geotrupoides songyingziensis Hong 1984
Fm. Glaphyridae MacLeay 1819 [bumble bee scarab beetle]
+
Subfm. Amphicominae Blanchard 1845
Subfm. †Cretoglaphyrinae Nikolajev 2005
G. †Cretohypna Yan et al. 2012
Subfm. Glaphyrinae MacLeay 1819
G. Lichnanthe Burmeister 1844
G. †Lithohypna Nikolajev et al. 2011
G. †Mesohypna Nikolajev and Ren 2013
G. †Paraglaphyrus Nikolajev 2015
Fm. Glaresidae Prudhomme de Borre 1886 [enigmatic scarab beetle]
+
G. †Cretoglaresis Nikolajev 2007
G. Glaresis Erichson 1848
G. †Lithoglaresis Nikolajev 2007
Invalid names: Aphodiites Heer 1864 [nomen dubium]
G. †Holcorobeus Nikritin 1977
+
†Holcorobeus atrosulcatus Nikolajev 1992
†Holcorobeus evittatus Zhang 1992
†Holcorobeus nigrovittatus Nikolajev 1992
†Holcorobeus vittatus Nikritin 1977
Invalid names: Holcorobeus (Holcorobeus) Nikritin 1977 [invalid subgroup]
G. †Hongscarabaeus Krell 2000
+
†Hongscarabaeus brunneus Hong 1982
Invalid names: Proscarabaeus Hong 1982 [replaced]
Fm. Hybosoridae Erichson 1847 [scavenger scarab beetle]
+
Subfm. Anaidinae Nikolajev 1996
Subfm. Ceratocanthinae Martinez 1968
G. †Coprologus Heer 1847
Subfm. Hybosorinae Erichson 1847
Subfm. Liparochrinae Ocampo 2006
Subfm. †Mimaphodiinae Nikolajev 2007
G. †Procoilodes Ocampo 2002
G. †Pulcherhybosorus Yan et al. 2012
G. †Sinochaetodus Lu et al. 2018
G. †Sinohybosorus Nie et al. 2018
G. †Tyrannasorus Ratcliffe and Ocampo 2001
Fm. Lucanidae Latreille 1804 [stag beetle]
+
Subfm. Aesalinae MacLeay 1819
Subfm. †Ceruchitinae Nikolajev 2006
G. †Cretolucanus Nikolajev 2007
G. †Dorcasoides Motschulsky 1856
Subfm. Lampriminae MacLeay 1819
Subfm. †Litholampriminae Nikolajev 2015
G. †Paleognathus Waga 1883
Subfm. †Paralucaninae Nikolajev 2000
G. †Prosinodendron Bai et al. 2012
Subfm. †Protolucaninae Nikolajev 2007
Subfm. Syndesinae MacLeay 1819
G. †Mesoscarabaeus Hong 1982
+
†Mesoscarabaeus corneus Hong 1982
†Mesoscarabaeus morulosus Hong 1982
Fm. Ochodaeidae Streubel 1846 [sand-loving scarab beetle]
+
Subfm. Chaetocanthinae Scholtz 1988
Subfm. †Cretochodaeinae Nikolajev 1995
G. †Mesochodaeus Nikolajev and Ren 2010
Subfm. Ochodaeinae Streubel 1846
Fm. Passalidae Leach 1815 [bess beetle]
+
Subfm. Aulacocyclinae Kaup 1868
Subfm. Passalinae Leach 1815
Fm. †Passalopalpidae Boucher and Bai 2016
+
G. †Mesoceratocanthus Nikolajev et al. 2010
G. †Passalopalpus Boucher and Bai 2016
Fm. Pleocomidae Leconte 1861 [rain beetle]
+
Subfm. †Archescarabaeinae Nikolajev 2010
Subfm. †Cretocominae Nikolajev 2002
Subfm. Pleocominae Leconte 1861
Fm. Scarabaeidae Latreille 1802 [scarab beetle]
+
Subfm. Aclopinae Blanchard 1850
Subfm. Aegialiinae Laporte 1840
G. †Antiqusolidus Lu et al. 2023
Subfm. Aphodiinae Leach 1815
Subfm. Aulonocneminae Janssens 1946
G. Caccobius Thompson 1859
Subfm. Cetoniinae Leach 1815 [flower chafer]
Subfm. †Cretoscarabaeinae Nikolajev 1995
Subfm. Dynastinae MacLeay 1819 [rhinoceras beetle]
Subfm. †Electrorubesopsinae Bai and Wang 2018
Subfm. Eremazinae Stebnicka 1977
G. Liothorax Motschulsky 1859
Subfm. †Lithoscarabaeinae Nikolajev 1992
Subfm. Melolonthinae Leach 1819
G. †Mesydra Ponomarenko 1977
Subfm. †Prototroginae Nikolajev 2000
Subfm. Rutelinae MacLeay 1819
Subfm. Scarabaeinae Latreille 1802
G. Triodonta Mulsant 1842
Invalid names: Allidiostomatinae Arrow 1940 [empty], Coprophaginae [empty], Dynamopodinae Arrow 1911 [empty], Oniticellus amplicollis Heer 1862 [nomen dubium], Orphninae Erichson 1847 [empty], Pachypodinae Erichson 1840 [empty], Phaenomeridinae Erichson 1847 [empty]
Fm. Trogidae MacLeay 1819 [hide beetle]
+
Subfm. †Kresnikinae Tihelka et al. 2019
Subfm. Omorginae Nikolajev 2005
Subfm. Troginae MacLeay 1819
Invalid names: Trox oustaleti Scudder 1879 [nomen dubium]
Invalid names: Belohinidae Paulian 1959 [empty], Copris subterraneus Heer 1862 [nomen dubium], Gymnopleurus deperditus Heer 1862 [nomen dubium], Holcorobeini Nikolajev 1992 [invalid subgroup], Onthophagus urusheeri Krell 2000 [nomen dubium]
Superfm. Staphylinoidea Latreille 1802
+
Fm. Agyrtidae Thomson 1859 [primitive carrion beetle]
+
Subfm. Agyrtinae Thomson 1859
G. †Eosilphites Haupt 1950
Subfm. Necrophilinae Newton 1997
G. †Ponomarenkia Perkovsky 2001
G. †Prosilpha Hong 1982
Subfm. Pterolomatinae Thomson 1862
G. †Sinosilphia Hong and Wang 1990
Fm. Hydraenidae Mulsant 1844 [minute moss beetle]
+
Subfm. Hydraeninae Mulsant 1844
G. †Ochtebiites Ponomarenko 1977
Subfm. Ochthebiinae Thomson 1859
Invalid names: Prosthetopinae Perkins 1994 [empty]
Fm. Jacobsoniidae Heller 1926
+
Subfm. Derolathrinae Sen Gupta 1979
G. Sarothrias Grouvelle 1918
Invalid names: Sarothriidae [replaced]
Fm. Leiodidae Fleming 1821 [round fungus beetle]
+
G. Agaricophagus Schmidt 1841
Subfm. Camiarinae Jeannel 1911
Subfm. Catopocerinae Hatch 1927
Subfm. Cholevinae Kirby 1837
G. Cholevinus Reitter 1901
Subfm. Leiodinae Fleming 1821
Invalid names: Leptininae LeConte 1866 [empty], Platypsyllinae Ritsema 1869 [empty]
Fm. Ptiliidae Erichson 1845 [feather-winged beetle]
+
G. Microptilium Matthews 1872
Subfm. Nossidiinae Sörensson and Delgado 2019
Subfm. Ptiliinae Erichson 1845
Fm. Staphylinidae Latreille 1802 [rove beetle]
+
Subfm. Aleocharinae Fleming 1821
G. †Aleocharopsis Wickham 1913
Subfm. Apateticinae Fauvel 1895
G. †Apticax Schomann and Solodovnikov 2012
G. †Bembicidiodes Schaufuss 1889
G. Boletobius Leach 1819
G. †Cardiolita Strand 1933
G. †Caririderma Martins-Neto 1990
Subfm. Coomaniinae Żyła and Solodovnikov 2020
Subfm. Dasycerinae Reitter 1887
G. Ellipsotomus Motschulsky 1845
Subfm. Euaesthetinae Thomson 1859
Subfm. Glypholomatinae Jeannel 1962
G. Heterosoma Bernhauer 1903
G. †Laostaphylinus Zhang 1988
Subfm. Megalopsidiinae Leng 1920
G. †Mesapatetica Cai et al. 2014
Subfm. Micropeplinae Leach 1815
Subfm. Olisthaerinae Thomson 1858
Subfm. Omaliinae MacLeay 1825
Subfm. Osoriinae Erichson 1839
Subfm. Oxyporinae Fleming 1821
Subfm. Oxytelinae Fleming 1821
Subfm. Paederinae Fleming 1821
G. †Paroxyporus Cai 2021
Subfm. Phloeocharinae Erichson 1839
Subfm. Piestinae Erichson 1839
Subfm. †Protactinae Heer 1847
Subfm. Proteininae Erichson 1839
Subfm. Protopselaphinae Newton and Thayer 1995
Subfm. Pselaphinae Latreille 1802
Subfm. Pseudopsinae Ganglbauer 1895
Subfm. Scaphidiinae Latreille 1807
G. †Scaphylinus Kaddumi 2005
Subfm. Scydmaeninae Leach 1815 [antlike stone beetle]
G. †Seniaulus von Heyden and von Heyden 1866
Subfm. Silphinae Latreille 1807
G. †Sinostaphylina Hong and Wang 1990
G. †Sinostaphylius Hong 1992
Subfm. Solieriinae Newton and Thayer 1992
Subfm. Staphylininae Latreille 1802
+
Invalid names: Platyprosopinae Lynch Arribálzaga 1884 [synonym], Xantholininae Erichson 1839 [synonym]
G. †Staphylinites Scudder 1876
Subfm. Steninae Macleay 1825
G. †Sulcelytrinus Tikhomirova 1968
Subfm. Tachyporinae MacLeay 1825
Subfm. Trichophyinae Thomson 1858
Subfm. Trigonurinae Reiche 1821
G. †Tunicopterus Tikhomirova 1968
Invalid names: Empelinae Newton and Thayer 1992 [empty], Habrocerinae Mulsant and Rey 1877 [empty], Leptotyphlinae Fauvel 1874 [empty], Microsilphinae Crowson 1950 [empty], Neophoninae Fauvel 1905 [empty], Silphidae Latreille 1807 [synonym], Triga coeni Scudder 1900 [nomen dubium]
Invalid names: Dasyceridae Reitter 1887 [empty], Empelidae [empty]
Invalid names: Scarabaeiformia Crowson 1960 [synonym]
Invalid names: Agyrtes primoticus Scudder 1900 [nomen dubium], Derodontiformia Le Conte 1861 [invalid subgroup], Escheria crassipunctata Förster 1891 [nomen dubium], Escheria dimidiata Förster 1891 [nomen dubium], Escheria punctulata Förster 1891 [nomen dubium], Eucinetiformia [empty], Limnius perrini Theobald 1937 [nomen dubium], Malachius vertummni Heer 1847 [nomen dubium], Mesosperchus notatus Ponomarenko 1977 [nomen dubium], Tomocoleites Hong 2002 [nomen nudum], Troglops punctatissima Théobald 1935 [nomen dubium]
Fm. †Ponomarenkiumidae Lawrence and Bouchard 2022
+
G. †Ponomarenkium Yan et al. 2018
+
†Ponomarenkium belmonthense Yan et al. 2017
Invalid names: Ponomarenkia Yan et al. 2017 [replaced]
Invalid names: Ponomarenkiidae Yan et al. 2017 [invalid subgroup]
Fm. †Taldycupedidae Rohdendorf 1961
+
G. †Afrotaldycupes Kirejtshuk 2020
+
†Afrotaldycupes africanus Ponomarenko 2005
†Afrotaldycupes lidgettoniensis Ponomarenko 2005
G. †Mesothoris Tillyard 1916
+
†Mesothoris clathrata Tillyard 1916
†Mesothoris grandis Dunstan 1923
†Mesothoris punctomarginum Dunstan 1923
G. †Proterocupes Ponomarenko 2015
+
†Proterocupes major Ponomarenko 2015
†Proterocupes nedubrovensis Ponomarenko 2015
†Proterocupes permiana Ponomarenko 2013
Invalid names: Lobanovia Kirejtshuk 2020 [objective synonym]
G. †Simmondsia Dunstan 1923
+
†Simmondsia cylindrica Dunstan 1923
†Simmondsia subpiriformis Dunstan 1923
G. †Taldycupes Rohdendorf 1961
+
†Taldycupes cellulosus Ponomarenko 2013
†Taldycupes khalfini Rohdendorf 1961
†Taldycupes moltshanovi Rohdendorf 1961
†Taldycupes pingi Ponomarenko et al. 2014
†Taldycupes reticulatus Ponomarenko 1969
†Taldycupes rosanovi Ponomarenko 2006
†Taldycupes vasjuchitshevi Rohdendorf 1961
Invalid names: Cryptocupes Rohdendorf 1961 [synonym], Taldycupidium Rohdendorf 1961 [synonym]
G. †Tecticupes Rohdendorf 1961
+
†Tecticupes heckeri Rohdendorf 1961
+
Invalid names: Stegocupes efremovi Rohdendorf 1961 [synonym], Stegocupes fedotovi Rohdendorf 1961 [synonym]
†Tecticupes indistinctus Rohdendorf 1961
+
Invalid names: Tychticupes tchihatcheffi Rohdendorf 1961 [synonym]
†Tecticupes martynovi Ponomarenko 2013
Invalid names: Stegocupes Rohdendorf 1961 [synonym]
G. †Tychticupes Rohdendorf 1961
+
†Tychticupes beljanini Rohdendorf 1961
†Tychticupes radtshenkoi Rohdendorf 1961
†Tychticupes ragozini Rohdendorf 1961
Invalid names: Tychticupes ussovi Rohdendorf 1961 [nomen dubium], Uskaticupes Rohdendorf 1961 [nomen dubium], Uskaticupes javorskyi Rohdendorf 1961 [nomen dubium]
Fm. †Triadocupedidae Ponomarenko 1966
+
G. †Cupesia Ponomarenko 1966
+
†Cupesia monilicornia Ponomarenko 1966
†Cupesia sepulta Ponomarenko 1969
†Cupesia serricornia Ponomarenko 1969
G. †Platycupes Ponomarenko 1966
+
†Platycupes dolichocerus Ponomarenko 1966
†Platycupes major Ponomarenko 1966
†Platycupes pusillus Ponomarenko 1966
†Platycupes reticulatus Ponomarenko 1966
G. †Triadocupes Ponomarenko 1966
+
†Triadocupes ellipticus Ponomarenko 1966
†Triadocupes ferghanensis Ponomarenko 1966
†Triadocupes latus Ponomarenko 1966
Diagnosis
No diagnoses are available
Measurements
No measurements are available
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| Source: c = class, p = phylum | |||||
| References: Kiessling 2004, Bush and Bambach 2015 | |||||
Age range
Maximum range based only on fossils: base of the Asselian to the top of the Holocene or 298.90000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 295.5 Ma
Minimum age of oldest fossil (stem group age): 295.5 Ma
Collections (1821 total)
Oldest occurrences | |||
| Time interval | Ma | Country or state | Original ID and collection number |
|---|---|---|---|
| Asselian | Czech Republic | Moravocoleus perditus (114186) | |
| Roadian | Russian Federation (Kemerovo) | Asiocoleus novojilovi, Carinicupes beckermigdisovae, Schizotaldycupes ananjevi, Stegocupes efremovi, Uskatocoleus kaltanicus (104704) Rossocoleus avus (122567) Schizocoleus kuznetskiensis (124604) | |
| Roadian | Russian Federation | Stegosyne rugosa (124611) | |
| Roadian | Russian Federation (Kirov) | Rossocoleus altus (105058) | |
| Wordian | Russian Federation (Udmurtia) | Rossocoleus novojilovi, Rossocoleus angustus, Schizophoridae indet. (103502) Schizotaldycupes pubescens, Rhombocoleites uralensis (104750) | |
| Wordian | Russian Federation (Kemerovo) | Rossocoleus verrucosus, Uskatocoleus convexus (124603) Schizocupes obrutshevi (124600) Tychticupes tchihatcheffi, Tecticupes indistinctus, Stegocupes fedotovi, Tecticupes heckeri, Rossocoleus latus, Rossocoleus latior, Rossocoleus parvus, Rossocoleus grandis, Rossocoleus surijokovensis, Karakanocoleus janischewskyi, Uskatocoleus iljinskiensis, Uskatocoleus surijokovicus, Uskatocoleus avitus (122566) | |
| Wordian - Capitanian | Russian Federation | Uskatocoleus kushejakovicus (124610) | |
| Capitanian | Russian Federation (Orenburg) | Bicoleus kuplensis, Tetracoleus orenburgensis, Rhombocoleus gomankovi, Erunakicupes brevis, Schizocoleus longus, Schizocoleus minimus, Schizocoleus glabrus, Schizocoleus robustus, Karakanocoleus europeus, Palademosyne ovum (104717) Curculiopsis ellipticus (104994) Rossocoleus sakmara, Aenigmocoleus uralensis, Schizocoleus major (152235) Tecticupes martynovi, Rhombocoleites adumbratus, Kargalarva permosialis (104057) | |
| Capitanian | China (Anhui) | Chaocoleus limnebius, Asiocoleopsis hongi, Taldycupes pingi, Dikerocoleus divisus (103239) | |
| Wuchiapingian | Russian Federation (Vologda) | Bicoleus laticella, Tetracoleus tricoleoides, Taldycupes cellulosus, Simmondsia permiana, Taldycupedidae indet., Rossocoleus aristovi, Erunakicupes grossus, Schizocoleus depressus, Karakanocoleus mutovinensis, Pseudochrysomelites bashkuevi, Pseudochrysomelites longus, Pseudochrysomelites medialis, Pseudochrysomelites sphenoidalis, Uskatocoleus convexus, Uskatocoleus sukhonensis, Uskatocoleus micron (104754) | |
| Wuchiapingian | Russian Federation | Ademosyne sibirica, Tychticupes vasjuchitshevi, Taldycupes moltshanovi, Cryptocupes rjabinini, Taldycupidium bergi, Tychticupoides grjazevi, Rhombocoleus andreae, Aenigmocoleus borissiaki, Erunakicupes schmalhauseni, Erunakicupes fomitshevi, Erunakicupes kryshtofovichi, Karakanocoleus striatus, Karakanocoleus magnus, Karakanocoleodes latissimus, Karakanocoleus major, Karakanocoleus lebedevi, Erunakicupes venjukovi, Uskatocoleus angustus, Uskatocoleus longus, Uskatocoleus angustior, Uskatocoleus zalesskyi, Uskatocoleus foveolatus, Uskatocoleus lutugini, Uskatocoleus minimus, Polysitum kuznetskiense (124608) Taldycupes khalfini, Uskaticupes javorskyi, Tychticupes ragozini, Tychticupes radtshenkoi, Taldycupes beljanini, Tychticupes ussovi, Uskatocoleus zalesskyi (122565) | |
| Wuchiapingian | Russian Federation (Orenburg) | Uskatocoleus uralensis (152236) | |
| Changhsingian | Russian Federation (Evenki Autonomous Okrug) | Tunguskagyrus planus, Tunguskagyrus yani (118027) Tunguskagyrus yani (105049) | |
| Changhsingian | South Africa (Kwazulu-Natal) | Taldycupes africanus (14356) Taldycupes lidgettoniensis, Rhombocoleites danutae (14357) | |
| Changhsingian | Australia (New South Wales) | Permosyne pincombeae (122414) Ponomarenkia belmonthensis (122433) Tetracoleus sp. (111680) | |
| Changhsingian | Russian Federation (Vladimir) | Erunakicupes angustus, Pseudochrysomelites major, Metrorhynchites elongatus, Metrorhynchites baculum, Uskatocoleus eurypygus, Uskatocoleus rhynchophorus, Schizocoleidae indet. (113351) Tetracoleus sennikovi (104756) | |
| Changhsingian | Russian Federation (Vologda) | Tetracoleus golubevi, Proterocupes nedubrovensis, Proterocupes major, Pseudochrysomelites latissimus, Pseudochrysomelites convexus, Pseudochrysomelites circumflexus, Palademosyne latum, Uskatocoleus minor, Uskatocoleus hirsutus, Uskatocoleus artus, Schizocoleidae indet. (105046) Tetracoleus vasilenkoi, Tetracoleus aristovi, Tetracoleus meyeni, Pseudochrysomelites dvinensis (105041) | |
| Changhsingian | Kazakhstan (Zaisan) | Taldycupes reticulatus (122972) | |
| Changhsingian | Russian Federation (Krasnoyarsk) | Taldycupes rosanovi (108358) | |
| Changhsingian - Induan | Russian Federation | Palaeoscapha tunguskaensis (229880) | |