PBDB Taxon

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Hesperhys vagrans

Mammalia - Tayassuidae

Taxonomy

Hesperhys vagrans was named by Douglass (1903) [genotype].

Synonyms

Desmathyus validus was named by Matthew (1932). Its type specimen is AMNH 20506, a maxilla (upper jaw, young, with dp3-m1 and p3-4 preformed.), and it is a 3D body fossil. Its type locality is Thomson Quarry, which is in a Hemingfordian terrestrial horizon in the Sheep Creek Formation of Nebraska.

It was synonymized subjectively with Hesperhys vagrans by Wright (1998), Prothero (2015) and Prothero (2021).

Synonymy list

Year	Name and author
1903	Hesperhys vagrans Douglass
1932	Desmathyus validus Matthew p. 3 fig. 4
1998	Hesperhys vagrans Wright p. 395
2015	Hesperhys vagrans Prothero p. 241 figs. 4 - 6
2021	Hesperhys vagrans Prothero p. 10

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Rank	Name	Author
kingdom	Animalia	()
—	Bilateria
—	Eubilateria	Ax 1987
—	Deuterostomia
phylum	Chordata	Haeckel 1874
subphylum	Vertebrata
superclass	Gnathostomata
—	Osteichthyes	()
subclass	Sarcopterygii	()
subclass	Dipnotetrapodomorpha	(Nelson 2006)
subclass	Tetrapodomorpha	()
—	Tetrapoda
—	Reptiliomorpha
—	Anthracosauria
subclass	Amphibiosauria	Kuhn 1967
—	Cotylosauria	()
—	Amniota
subclass	Synapsida
—	Therapsida	()
infraorder	Cynodontia	()
—	Mammaliamorpha
—	Mammaliaformes

Rank	Name	Author
class	Mammalia
—	Theriamorpha	(Rowe 1993)
—	Theriiformes	()
—	Trechnotheria
—	Cladotheria
—	Zatheria
subclass	Tribosphenida	()
subclass	Theria
—	Eutheria	()
—	Placentalia
—	Boreoeutheria
—	Laurasiatheria
—	Scrotifera
—	Euungulata
—	Artiodactylamorpha
—	Artiodactyla	()
superfamily	Suoidea
—	Suina	(Gray 1868)
family	Tayassuidae
subfamily	Hesperhyinae
genus	Hesperhys
species	vagrans

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

†Hesperhys vagrans Douglass 1903

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Invalid names: Desmathyus validus Matthew 1932 [synonym]

Diagnosis

Reference	Diagnosis
W. D. Matthew 1932 (Desmathyus validus)	-Slightly larger than D. pinensis; p3 sub-oval in form, nearly as large as p4 and similar in cusp construction. In D. pinensis the inner crescent of p3 is smaller and imperfect, giving the tooth a sub-trigonal outline.
D. R. Prothero 2015	[Same as for genus] Very large hesperhyine with very massive, robust skull and aring orbits and zygomatic arches, and broad rostrum; cheek teeth more robust and bulbous than in any other hesperhyine. The plesiochoanal fossa is reduced to a shallow pocket. P3 has a large protocone. The i1 and i2 are deep dorsoventrally at the base.