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Holoclemensia texana

Mammalia - Holoclemensiidae

Taxonomy
Clemensia texana was named by Slaughter (1968) ["genotype"]. Its type specimen is SMP-SMU 61997, a tooth (RM2, missing the protoconal region), and it is a 3D body fossil. Its type locality is Butler Farm, which is in an Aptian/Albian terrestrial claystone in the Antlers Formation of Texas. It is the type species of Holoclemensia, Clemensia.

It was recombined as Holoclemensia texana by Slaughter (1968), Kielan-Jaworowska et al. (1979), Kielan-Jaworowska et al. (2004), Winkler et al. (2011) and Davis and Cifelli (2011).

Synonyms
Synonymy list
YearName and author
1968Clemensia texana Slaughter
1968Holoclemensia texana Slaughter
1979Holoclemensia texana Kielan-Jaworowska et al. p. 183
1989Comanchea hilli Jacobs et al.
2004Comanchea hilli Kielan-Jaworowska et al. pp. 411, 422
2004Holoclemensia texana Kielan-Jaworowska et al. pp. 411, 421
2011Holoclemensia texana Davis and Cifelli p. 450
2011Holoclemensia texana Winkler et al. p. 8

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
RankNameAuthor
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
Mammaliaformes
classMammalia
Theriamorpha(Rowe 1993)
Theriiformes()
Trechnotheria
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
familyHoloclemensiidaeAplin and Archer 1987
genusHoloclemensia
speciestexana()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Holoclemensia texana Slaughter 1968
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Invalid names: Comanchea hilli Jacobs et al. 1989 [synonym]
Diagnosis
ReferenceDiagnosis
B. M. Davis and R. L. Cifelli 2011Basal eutherian mammal differing from all other basal eutherians in presence of a very large central stylar cusp (= mesostyle); differs from stem tribosphenidans in very small size of stylocone, the presence of a very large mesostyle, relatively narrow metastylar lobe at all loci, a prominent, flange−like parastylar lobe on mesial molars, very tall, bulbous metaconid on lower molars, and highly mesiodistally compressed trigonid; differs from deltatheroidans in presence of three molars, relatively weak postmetacrista, small stylocone, and small, inclined paraconid; differs from Slaughteria in larger size, presence of metaconid on p5, metaconid taller than paraconid on molars, and absence of distal metacristid; differs from Pappotherium in presence of three molars, shallower ectoflexus, more separated paracone and metacone, narrower metastylar lobe, metaconid taller than paraconid, and absence of distal metacristid.
Measurements
No measurements are available
Composition: phosphaticsubp
Environment: terrestrialsubc
Locomotion: actively mobilec
Life habit: scansorialsubc
Diet: insectivoreg
Reproduction: viviparoussubc
Created: 2005-09-07 09:54:10
Modified: 2005-09-07 11:54:10
Source: g = genus, subc = subclass, c = class, subp = subphylum
References: Hendy et al. 2009, Ji et al. 2002, Carroll 1988, Lillegraven 1979, Kielan-Jaworowska et al. 1979

Age range: base of the Albian to the top of the Early/Lower Albian or 113.00000 to 109.00000 Ma

Collections (5 total)


Time interval Ma Country or state Original ID and collection number
Late/Upper Aptian - Early/Lower Albian122.46 - 109.0USA (Texas) Clemensia texana (type locality: 14375) Holoclemensia texana (14388)
Late/Upper Aptian - Early/Lower Albian122.46 - 109.0USA (Oklahoma) Holoclemensia texana (14392)
Albian113.0 - 100.5USA (Texas) Comanchea hilli (14411) Holoclemensia texana (14420)