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Taxonomy
Borophagus secundus was named by VanderHoof (1931) [named as subspecies of Aelurodon saevus by Matthew and Cook 1909; raised to species level].
It was recombined as Osteoborus secundus by Stirton and VanderHoof (1933), VanderHoof and Gregory (1940), Skinner et al. (1977), Richey (1979) and Munthe (1998).
It was recombined as Osteoborus secundus by Stirton and VanderHoof (1933), VanderHoof and Gregory (1940), Skinner et al. (1977), Richey (1979) and Munthe (1998).
Synonyms
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Synonymy list
Year | Name and author |
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1924 | Hyaenognathus direptor Matthew |
1928 | Hyaenognathus cyonoides Martin p. 235 |
1930 | Borophagus cyonoides Matthew and Stirton |
1931 | Borophagus cyonoides VanderHoof |
1931 | Borophagus secundus VanderHoof |
1932 | Borophagus cyonoides Reed and Longenecker |
1933 | Osteoborus cyonoides Stirton and VanderHoof p. 177 |
1933 | Osteoborus direptor Stirton and VanderHoof p. 177 |
1933 | Osteoborus secundus Stirton and VanderHoof p. 178 |
1935 | Borophagus cyonoides Bode |
1936 | Osteoborus cyonoides Hesse |
1937 | Osteoborus cyonoides Johnston |
1939 | Osteoborus cyonoides Johnston |
1940 | Osteoborus secundus VanderHoof and Gregory |
1941 | Osteoborus cyonoides Stirton |
1944 | Osteoborus cyonoides McGrew |
1948 | Osteoborus cyonoides Macdonald |
1969 | Osteoborus cyonoides Dalquest |
1977 | Osteoborus secundus Skinner et al. |
1979 | Osteoborus cyonoides Bennett |
1979 | Osteoborus cyonoides Richey |
1979 | Osteoborus direptor Richey |
1979 | Osteoborus secundus Richey |
1980 | Osteoborus cyonoides Dalquest and Mooser p. 3 |
1980 | Borophagus direptor Kurten and Anderson p. 165 |
1983 | Osteoborus cyonoides Harrison |
1984 | Osteoborus cyonoides Webb and Perrigo |
1990 | Osteoborus cyonoides Schultz |
1998 | Osteoborus cyonoides Miller and Carranza-Castaneda |
1998 | Osteoborus cyonoides Munthe |
1998 | Osteoborus secundus Munthe |
1999 | Borophagus secundus Wang et al. |
2008 | Borophagus secundus Wang et al. |
2016 | Borophagus secundus Tseng and Geisler p. 2 figs. Table 1 |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
†Borophagus secundus VanderHoof 1931
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Invalid names: Hyaenognathus cyonoides Martin 1928 [synonym], Hyaenognathus direptor Matthew 1924 [synonym]
Diagnosis
Reference | Diagnosis | |
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R. A. Stirton and V. L. VanderHoof 1933 | Very closely related to O. cyonoides; p1 apparently more frequently present in secundus than in O. cyonoides. A series from Snake Creek may show that p1 is predominantly present and this possibility is sufficiently great to warrant the maintenance of O. secundus as a species. p2 has two roots; p3 is not so narrow nor so long-crowned as O. cyonoides, neither is it wider posteriorly than anteriorly. | |
R. A. Stirton and V. L. VanderHoof 1933 (Osteoborus cyonoides) | Rostrum short, frontals abruptly elevated from level of nasals; frontals extremely convex both antero-posteriorly and laterally; zygomatic arches wide; palate short and wide; P1-3 with very low median cusps; p2 with double or single roots and vestigial anterior and posterior cusps in some individuals; P3 with widely separated roots; crown of P3 elongate and narrow, seldom high; talon of M1 not as wide as in O. littoralis; horizontal ramus deflected outwardly below p4 giving tooth row a curved alignment; p1 present in only one specimen from Coffee Ranch, alveolus shallow, root single; p3 with separate roots, or one grooved root, or one round root, low median cusps on p2; p3 with two roots; p3 usually wider posteriorly than anteriorly, with a low to moderately high median cusp and a vestigial posterior cusp. | |
R. A. Stirton and V. L. VanderHoof 1933 (Osteoborus direptor) | Horizontal ramus deflected outwardly below p4 giving tooth row a curved alignment; p1 absent; p2 with one root; p3 with a distinct external goove in its single root. |