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Hymenopterida

Insecta

Taxonomy
Hymenopterida is an unranked clade. It is extant.

It was assigned to Holometabola by Engel (2022).

Sister taxa
Apatohymenoptera, Hymenoptera (syn. Aculeata, Piezata)

Synonymy list
YearName and author
2022Hymenopterida Engel

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Protostomia
Ecdysozoa
Panarthropoda
phylumArthropodaLatreille 1829
superclassHexapodaLatreille 1825
classInsecta
RankNameAuthor
Dicondylia
Paranotalia
subclassPterygota()
NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
Holometabola
Hymenopterida
Hymenopterida

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

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Or. †Apatohymenoptera Engel 2022
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Fm. †Avioxyelidae Nel et al. 2013
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G. †Avioxyela Nel et al. 2013
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Avioxyela gallica Nel et al. 2013
Or. Hymenoptera Linnaeus 1758
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Unr. Aculeati Latreille 1802
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Unr. Nemoglossata Latreille 1802
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Fm. Apiariae Latreille 1802
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Invalid names: Apes Latreille 1802 [empty], Bombi Latreille 1802 [empty], Ceratinae Latreille 1802 [empty], Eucerae Latreille 1802 [empty], Megachiles Latreille 1802 [empty], Parasitae Latreille 1802 [empty], Xylocopae Latreille 1802 [empty]
Unr. Platiglossata Latreille 1802
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Invalid names: Andrenetae Latreille 1802 [empty], Crabronites Latreille 1802 [empty], Formicariae Latreille 1802 [empty], Philantores Latreille 1802 [empty], Scolietae Latreille 1802 [empty], Sphegimae Latreille 1802 [empty]
Superfm. Anaxyeloidea Martynov 1925
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Fm. Anaxyelidae Martynov 1925
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Subfm. †Anaxyelinae Martynov 1925
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G. †Anasyntexis Rasnitsyn 1968
G. †Anaxyela Martynov 1925
G. †Brachysyntexis Rasnitsyn 1969
G. †Dolichostigma Rasnitsyn 1968
G. †Kempendaja Rasnitsyn 1968
G. †Kulbastavia Rasnitsyn 1968
G. †Mangus Kopylov 2019
G. †Sphenosyntexis Rasnitsyn 1969
G. †Syntexyela Rasnitsyn 1968
G. †Urosyntexis Rasnitsyn 1969
Invalid names: Dolichostigmatinae Rasnitsyn 1968 [synonym], Kempendajinae Rasnitsyn 1980 [synonym]
Subfm. Syntexinae Benson 1935
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G. †Cretosyntexis Rasnitsyn and Martínez-Delclòs 2000
G. †Curiosyntexis Kopylov 2019
G. †Curvitexis Jouault et al. 2022
G. †Daosyntexis Kopylov et al. 2020
G. †Dolichosyntexis Kopylov 2019
G. †Eosyntexis Rasnitsyn 1990
G. †Orthosyntexis Gao et al. 2021
G. †Parasyntexis Kopylov 2019
G. †Sclerosyntexis Wang et al. 2020
Invalid names: Paraxiphydriinae Gao et al. 2022 [synonym]
Fm. †Bryopompilidae Engel and Grimaldi 2006
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Invalid names: Bryopompilidae Rodriguez et al. 2016 [objective synonym]
G. †Nygmatius Hong 1983
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Superfm. Pamphilioidea Cameron 1890 [sawfly]
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Fm. Megalodontesidae Konow 1897
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Subfm. †Archoxyelydinae Wang et al. 2013
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G. †Archoxyelyda Wang et al. 2013
G. †Xyelydontes Rasnitsyn 1983
Subfm. †Decorisiricinae Wang et al. 2015
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G. †Brevisiricius Wang et al. 2015
G. †Decorisiricius Wang et al. 2015
G. †Limbisiricius Wang et al. 2015
G. †Jibaissodes Ren 1995
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Jibaissodes bellus Gao et al. 2016
Jibaissodes peichenae Wang et al. 2019
Subfm. Megalodontesinae Konow 1897
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Invalid names: Rudisiriciinae Gao et al. 2010 [synonym]
Subfm. †Praesiricinae Rasnitsyn 1968
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G. †Hoplitolyda Gao et al. 2013
G. †Praesirex Rasnitsyn 1968
G. †Turgidontes Rasnitsyn 1990
Invalid names: Praesiricidae Rasnitsyn 1968 [synonym]
Fm. †Mirolydidae Wang et al. 2017
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G. †Mirolyda Wang et al. 2017
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Mirolyda hirta Wang et al. 2017
Fm. Pamphiliidae Cameron 1890
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Subfm. Cephalciinae Benson 1945
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G. Acantholyda Costa 1894
G. Cephalcia Jurine 1801
Invalid names: Cephalciini Benson 1945 [invalid subgroup]
G. Cephaleia Jurine 1801
Subfm. Juralydinae Rasnitsyn 1977
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G. †Atocus Scudder 1892
G. †Dolicholyda Zhuang et al. 2022
G. †Juralyda Rasnitsyn 1977
G. †Scabolyda Wang et al. 2014
G. †Tapholyda Rasnitsyn 1983
Subfm. Pamphiliinae Cameron 1890
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G. Neurotoma Konow 1897 [Leaf-rolling sawfly]
G. Pamphilius Latreille 1802
Invalid names: Neurotomini Benson 1945 [invalid subgroup]
G. †Ulteramus Archibald and Rasnitsyn 2015
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Ulteramus republicensis Archibald and Rasnitsyn 2015
Invalid names: Lydites Newman 1834 [replaced]
G. †Prolyda Rasnitsyn 1968
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Prolyda depressa Rasnitsyn 1969
Prolyda dimidia Wang et al. 2016
Prolyda elegantula Wang et al. 2016
Prolyda karatavica Rasnitsyn 1968
Prolyda xyelocera Rasnitsyn 1968
Fm. †Xyelydidae Rasnitsyn 1968
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G. †Brevilyda Wang and Rasnitsyn 2016
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Brevilyda provecta Wang and Rasnitsyn 2016
G. †Chifengilyda Zheng and Chen 2018
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Chifengilyda robusta Zheng and Chen 2018
G. †Ferganolyda Rasnitsyn 1983
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Ferganolyda charybdis Rasnitsyn et al. 2006
Ferganolyda chungkuei Rasnitsyn et al. 2006
Ferganolyda cubitalis Rasnitsyn 1983
Ferganolyda eucalla Wang et al. 2015
Ferganolyda insolita Wang et al. 2015
Ferganolyda radialis Rasnitsyn 1983
Ferganolyda scylla Rasnitsyn et al. 2006
Ferganolyda sogdiana Rasnitsyn 1983
G. †Fissilyda Wang et al. 2015
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Fissilyda alba Wang et al. 2015
Fissilyda compta Wang et al. 2015
Fissilyda parilis Wang et al. 2015
G. †Medilyda Wang and Rasnitsyn 2016
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Medilyda distorta Wang and Rasnitsyn 2016
Medilyda procera Wang and Rasnitsyn 2016
G. †Mesolyda Rasnitsyn 1963
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Mesolyda jurassica Rasnitsyn 1963
Mesolyda sibirica Rasnitsyn 1983
Invalid names: Alloxyelula Ren 1995 [synonym]
G. †Novalyda Gao et al. 2013
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Novalyda cretacica Gao et al. 2013
Novalyda decora Wang et al. 2015
G. †Rectilyda Wang et al. 2014
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Rectilyda sticta Wang et al. 2014
G. †Sagulyda Rasnitsyn 1983
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Sagulyda arcuata Rasnitsyn 1983
Sagulyda ferganica Rasnitsyn 1983
Sagulyda magna Rasnitsyn 1983
G. †Sinoprolyda Hong 1987
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G. †Sogutia Rasnitsyn 1977
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Sogutia liassica Rasnitsyn 1977
G. †Strenolyda Wang and Rasnitsyn 2016
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Strenolyda marginalis Wang and Rasnitsyn 2016
Strenolyda retrorsa Wang and Rasnitsyn 2016
G. †Strophandria Rasnitsyn 1968
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Strophandria grossa Rasnitsyn 1968
Strophandria moderata Rasnitsyn 1983
G. †Xyelyda Rasnitsyn 1968
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Xyelyda excellens Rasnitsyn 1968
Invalid names: Megalodontoidea Konow 1897 [synonym]
Superfm. Tenthredinoidea Latreille 1802 [wood wasp]
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Fm. Argidae Konow 1890 [argid sawfly]
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Subfm. Arginae Konow 1890
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G. Arge Schrank 1802
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Invalid names: Hylotoma Latreille 1802 [synonym]
G. †Mioarge Nel 2004
Subfm. Sterictiphorinae Rohwer 1911
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G. Didymia Lepeletier and Serville 1828
G. Sterictiphora Billberg 1820
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Invalid names: Schizocerus Berthold 1827 [synonym]
G. †Atefia Krogmann et al. 2013
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Atefia rasnitsyni Krogmann et al. 2013
Fm. Athaliidae Rohwer 1911
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G. Athalia Leach 1817
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Athalia vetuecclesiae Wappler et al. 2005
Athalia wheeleri Cockerell 1906
Fm. Blasticotomidae Thomson 1871
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G. Blasticotoma Klug 1834
G. †Paremphytus Brues 1908
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Paremphytus ostentus Brues 1908
G. Runaria Malaise 1931
G. †Zippelia Rasnitsyn and Müller 2023
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Zippelia engeli Rasnitsyn and Müller 2023
Invalid names: Xyelotomidae Rasnitsyn 1968 [synonym]
Fm. Cimbicidae Kirby 1837 [cimbicid sawfly]
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Subfm. Abiinae Thomson 1871
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G. Abia Leach 1817
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Invalid names: Procimbex Hong 1984 [synonym], Zaraea Leach 1817 [synonym]
G. †Trichiosomites Brues 1908
Subfm. †Cenocimbicinae Archibald and Rasnitsyn 2023
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G. †Allenbycimbex Archibald and Rasnitsyn 2023
G. †Cenocimbex Nel 2004
G. †Leptostigma Archibald and Rasnitsyn 2023
Subfm. Cimbicinae Kirby 1837
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G. Cimbex Olivier 1790
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Invalid names: Clavellaria Olivier 1789 [replaced]
G. †Floricimbex Archibald and Rasnitsyn 2023
G. †Phenacoperga Cockerell 1908
G. Pseudoclavellaria Schultz 1906
G. †Sinocimbex Zhang et al. 1994
Subfm. Corynidinae Benson 1938
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G. Corynis Thunberg 1789
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Invalid names: Amasis Leach 1817 [objective synonym]
Subfm. Pachylostictinae Benson 1938
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G. †Eopachylosticta Malaise 1945
Fm. Diprionidae Rohwer 1910 [conifer sawfly]
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Subfm. Diprioninae Rohwer 1910
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G. Diprion Schrank 1802
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Invalid names: Lophyrus Latreille 1802 [synonym]
G. †Eodiprion Schedl 2007
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Eodiprion groehni Schedl 2007
Subfm. Monocteninae MacGillivray 1906
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G. †Oligomonoctenus Nel et al. 2023
G. †Paleomonoctenus Nel 2004
Fm. †Electrotomidae Rasnitsyn 1977
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G. †Electrotoma Rasnitsyn 1977
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Electrotoma kopylovi Rasnitsyn and Manukyan 2023
Electrotoma succini Rasnitsyn 1977
G. †Manevalia Piton 1940
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Fm. Pergidae Ashmead 1898
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Subfm. Perginae Ashmead 1898
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G. Perga Leach 1817
Subfm. Perreyiinae Cameron 1883
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Invalid names: Euryinae Konow 1898 [synonym]
Invalid names: Syzygoniinae Cameron 1883 [empty]
G. †Pseudocimbex Rohwer 1908
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Pseudocimbex clavatus Rohwer 1908
Fm. Tenthredinidae Latreille 1802
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Subfm. Allantinae Newman 1834
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Tr. Allantini Newman 1834
Tr. Empriini Rohwer 1911
G. Eriocampa Hartig 1837
G. †Hylotomites Meunier 1914
G. †Palaeathalia Zhang 1985
G. †Palaeotaxonus Brues 1908
G. Pseudosiobla Ashmead 1898
G. Taxonus Hartig 1837
Subfm. Blennocampinae Konow 1890
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Tr. Tomostethini Benson 1938
G. †Emprionopsis Zhang et al. 1994
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Emprionopsis doliabdominalis Zhang et al. 1994
Subfm. Heterarthrinae Benson 1952
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G. Caliroa Costa 1859
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Invalid names: Eriocampoides Konow 1890 [synonym]
G. Fenusa Leach 1817
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Invalid names: Lithoryssus Brues 1906 [synonym]
G. †Leptocampus Zhang et al. 1994
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Leptocampus amoenus Zhang et al. 1994
Subfm. Nematinae
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G. Cladius Illiger 1807
Tr. Dineurini Ashmead 1898
G. †Driocampus Zhang and Zhang 1990
G. †Eohemichroa Zhelochovtzev and Rasnitsyn 1972
G. †Florissantinus Zhelochovtzev and Rasnitsyn 1972
G. Hoplocampa Hartig 1837
G. Mesoneura Hartig 1837
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Invalid names: Lisconeura Rohwer 1908 [synonym]
G. Nematus Panzer 1801
G. †Paleonematus Nel 2022
Invalid names: Hoplocampini Konow 1890 [invalid subgroup]
G. †Phthonocampus Zhang et al. 1994
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Phthonocampus eos Zhang et al. 1994
G. Rhogogaster Konow 1884
Subfm. Selandriinae Thomson 1871
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Tr. Dolerini Thomson 1871
Tr. Selandriini Thomson 1871
Subfm. Tenthredininae Latreille 1802
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G. Macrophya Dahlbom 1835
G. †Nortonella Rohwer 1908
Tr. Perineurini Rohwer 1911
G. †Sambia Vilhelmsen and Engel 2012
G. †Taeniurites Cockerell 1917
G. Tenthredo Linnaeus 1758
G. †Tenthredoides Zhang 1989
G. Tenthredopsis Costa 1859
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Invalid names: Thomsonia Konow 1884 [synonym]
Invalid names: Tenthredinini Latreille 1802 [empty]
Invalid names: Nematus ellipticus Fritsch 1901 [nomen dubium]
Invalid names: Pteronus prodigus Brues 1908 [nomen dubium]
Unr. Terebrantes Latreille 1802
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Unr. Ventropedonculati Latreille 1802
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Fm. Ichneumonides Latreille 1802 [ichneumon wasp]
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Invalid names: Ichneumonides Latreille 1802 [empty], Sphegii Latreille 1802 [empty]
Invalid names: Cleptiosa Latreille 1802 [empty], Evaniales Latreille 1802 [empty]
Unr. Ventrosessiles Latreille 1802
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Invalid names: Tenthredines Latreille 1802 [empty], Urocerata Latreille 1802 [empty]
Unr. Unicalcarida Schulmeister et al. 2002
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Subor. Apocrita Gerstaecker 1867 [wasp]
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Unr. Aculeata Latreille 1802
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Superfm. Apoidea Latreille 1802 [apoid wasp]
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Invalid names: Sphecoidea Latreille 1802 [synonym]
Superfm. †Bethylonymoidea Rasnitsyn 1975
Superfm. Chrysidoidea Latreille 1802
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Invalid names: Bethyloidea [synonym]
G. †Cretopone Dlussky 1975
G. †Dolichomyrma Dlussky 1975
Superfm. Formicoidea Latreille 1802
Fm. †Holopsenellidae Engel et al. 2016
G. †Khetania Dlussky 1999
Fm. †Melittosphecidae Poinar and Danforth 2006
G. †Mesosirex Hong 1984
G. †Petropone Dlussky 1975
Superfm. Pompiloidea Latreille 1804
Superfm. Scolioidea Latreille 1802
Fm. †Stigmaphronidae Kozlov 1975
Superfm. Thynnoidea Shuckard 1841
Superfm. Tiphioidea Leach 1815
Superfm. Vespoidea Latreille 1802
Invalid names: Bocchus cenomanianus Olmi et al. 2010 [nomen dubium], Bombus pusillus Menge 1856 [nomen dubium]
G. †Apocritites Rasnitsyn and Jarzembowski 1998
+
Apocritites distinctus Rasnitsyn and Jarzembowski 1998
Fm. †Armaniidae Dlussky 1983
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G. †Archaeopone Dlussky 1975
G. †Armania Dlussky 1983
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Invalid names: Armaniella Dlussky 1983 [synonym]
G. †Orapia Dlussky et al. 2004
G. †Poneropterus Dlussky 1983
G. †Pseudarmania Dlussky 1983
Infraor. Brachycleistogastromorpha Kovalev 1994
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Superfm. †Brachycleistogastroidea Kovalev 1994
Invalid names: Cleistogastroidea Rasnitsyn 1975 [empty]
Superfm. Ceraphronoidea Haliday 1833
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Fm. Ceraphronidae Haliday 1833
Fm. Megaspilidae Ashmead 1893
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Invalid names: Calliceratidae [synonym]
Superfm. Evanioidea Latreille 1802
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Fm. †Andreneliidae Rasnitsyn and Martínez-Delclòs 2000
Fm. †Anomopterellidae Rasnitsyn 1975
Fm. †Baissidae Rasnitsyn 1975
+
Invalid names: Manlayidae Rasnitsyn 1986 [synonym]
Fm. Evaniidae Latreille 1802 [ensign wasp]
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Invalid names: Cretevaniidae Rasnitsyn 1975 [synonym]
Unr. Neoevanioides Engel 2006
Fm. †Othniodellithidae Engel and Huang 2016
Fm. †Praeaulacidae Rasnitsyn 1972
Fm. †Vectevaniidae Li et al. 2018
Invalid names: Asiapterites Hong 2002 [nomen nudum], Euryopterites Hong 2002 [nomen nudum], Oblongigastrula Hong 2002 [nomen nudum]
Superfm. Ichneumonoidea Latreille 1802
+
Fm. Braconidae Nees ab Esenbeck 1811
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Invalid names: Eoichneumonidae Jell and Duncan 1986 [synonym]
Fm. Ichneumonidae Latreille 1802 [ichneumon wasp]
Fm. †Ichneumonomimidae Rasnitsyn 1975
Fm. †Praeichneumonidae Rasnitsyn 1983
G. †Rasnitsynia Pagliano and Scaramozzino 1990
+
Invalid names: Oligoneuroides Zhang 1985 [replaced]
Fm. †Khutelchalcididae Rasnitsyn et al. 2004
+
G. †Khutelchalcis Rasnitsyn et al. 2004
Fm. †Kuafuidae Rasnitsyn and Zhang 2010
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G. †Arthrogaster Rasnitsyn 1975
G. †Kuafua Rasnitsyn and Zhang 2010
G. †Leptogastrella Rasnitsyn 1975
Fm. †Limnetidae Hong 1983
+
G. †Limnetus Hong 1983
G. †Longhuaia Hong 1984
+
Longhuaia rara Hong 1984
Superfm. Megalyroidea Schletterer 1889
+
G. †Cretodinapsis Rasnitsyn 1977
Fm. Megalyridae Schletterer 1890
Superfm. †Panguoidea Li et al. 2019
+
Fm. †Panguidae Li et al. 2019
+
Superfm. Cynipoidea Latreille 1802
+
Invalid names: Archaeocynipoidea Rasnitsyn and Kovalev 1988 [synonym]
G. †Koonwarrus Rasnitsyn and Öhm-Kühnle 2019
Unr. Microprocta Rasnitsyn and Öhm-Kühnle 2020
G. †Parviformosus Barling et al. 2013
Superfm. Proctotrupoidea Latreille 1802
Invalid names: Cynipomorpha Kovalev 1994 [synonym]
Superfm. Stephanoidea Leach 1815
+
Fm. †Aptenoperissidae Rasnitsyn and Poinar 2016
Fm. †Ephialtitidae Handlirsch 1906
+
Invalid names: Karataidae Rasnitsyn 1977 [synonym]
Fm. †Myanmarinidae Zhang and Rasnitsyn 2017
Fm. †Ohlhoffiidae Jouault et al. 2020
Fm. Stephanidae Leach 1815 [crown wasp]
Invalid names: Eostephanitidae Hong 2002 [invalid subgroup], Ephialtitoidea Handlirsch 1906 [synonym]
Superfm. Trigonalyoidea Cresson 1887
+
Fm. †Maimetshidae Rasnitsyn 1975
Fm. Trigonalidae Cresson 1887
Invalid names: Apiaria lapidea Germar 1842 [nomen dubium], Asiacynipites Hong 2002 [nomen nudum], Dendrocerus dubitatus Brues 1937 [nomen dubium], Eucynipites Hong 2002 [nomen nudum], Evaniomorpha Rasnitsyn 1988 [invalid subgroup], Stephanomorpha Leach 1815 [invalid subgroup]
Superfm. Cephoidea Newman 1834 [stem sawfly]
+
Fm. Cephidae Newman 1834 [stem sawfly]
+
Subfm. Cephinae Newman 1834
Subfm. †Cuspilonginae Kopylov and Rasnitsyn 2016
Fm. †Sepulcidae Rasnitsyn 1968
+
Subfm. †Ghilarellinae Rasnitsyn 1988
Subfm. †Parapamphiliinae Rasnitsyn 1968
Subfm. †Sepulcinae Rasnitsyn 1968
Subfm. †Trematothoracinae Rasnitsyn 1988
Subfm. †Xyelulinae Rasnitsyn 1993
G. †Cratoenigma Krogmann and Nel 2012
+
Cratoenigma articulata Krogmann and Nel 2012
Superfm. Orussoidea Newman 1834 [wood wasp]
+
Fm. †Burmorussidae Zhang et al. 2020
+
G. †Burmorussus Zhang et al. 2020
G. †Cretorussus Jouault et al. 2021
Fm. Orussidae Newman 1834 [parasitic wood wasp]
+
G. †Baltorussus Schedl 2011
Subfm. †Mesorussinae Rasnitsyn 1977
G. †Minyorussus Basibuyuk et al. 2000
Subfm. Ophrynopinae Benson 1935
Fm. †Paroryssidae Martynov 1925
+
G. †Microryssus Rasnitsyn 1968
G. †Paroryssus Martynov 1925
G. †Praeoryssus Rasnitsyn 1968
G. †Renaliculus Lin 1980
Invalid names: Karatavitoidea Rasnitsyn 1963 [synonym]
Superfm. Siricoidea Billberg 1820
+
Fm. †Protosiricidae Rasnitsyn and Zhang 2004
+
G. †Protosirex Rasnitsyn 1969
Fm. †Pseudosiricidae Handlirsch 1906
+
G. †Myrmicium Westwood 1854
+
Invalid names: Apiaria Germar 1839 [replaced], Fabellovena Oppenheim 1885 [synonym], Hagenia Weyenbergh 1869 [synonym], Pseudosirex Weyenbergh 1873 [synonym], Rhipidorhabdus Oppenheim 1885 [synonym]
Invalid names: Myrmiciidae Maa 1949 [synonym]
G. †Sinanxyela Hong 1982
+
Sinanxyela dorunensis Hong 1982
Fm. †Sinosiricidae Hong 1975
+
G. †Sinosirex Hong 1975
Fm. Siricidae Billberg 1820 [horntail]
+
Subfm. †Auliscinae Rasnitsyn 1968
Subfm. †Cratosiricinae Jouault and Nel 2020
G. †Cretosirex Wang et al. 2018
G. †Emanrisca Rasnitsyn 1993
G. †Eoteredon Archibald et al. 2021
Subfm. †Gigasiricinae Rasnitsyn 1968
+
Invalid names: Pararchexyelinae Rasnitsyn 1968 [synonym]
Subfm. †Megapteritinae Maa 1949
Subfm. Siricinae Billberg 1820
Subfm. Tremecinae Ashmead 1898
Subor. Vespina Laicharting 1781
+
Invalid names: Euhymenoptera [synonym]
Superfm. Xiphydrioidea Leach 1819
+
Invalid names: Xiphydriidae Leach 1819 [empty]
Invalid names: Orussomorpha Rasnitsyn 1980 [invalid subgroup]
Superfm. Xyeloidea Newman 1834 [sawfly]
+
Fm. †Syspastoxyelidae Engel and Huang 2016
+
G. †Burmoxyela Zheng and Rasnitsyn 2021
+
Burmoxyela lii Zheng and Rasnitsyn 2021
G. †Deltoxyela Wang et al. 2021
+
Deltoxyela engeli Wang et al. 2021
G. †Grandixyela Zheng 2021
+
Grandixyela rasnitsyni Zheng 2021
G. †Pinguixyela Zheng and Rasnitsyn 2021
+
Pinguixyela pinguis Wang et al. 2021
G. †Striaexyela Zheng 2019
+
Striaexyela simpla Wang et al. 2021
G. †Syspastoxyela Engel and Huang 2016
+
Syspastoxyela rhaphidia Engel and Huang 2016
Fm. Xyelidae Newman 1834 [sawfly]
+
Subfm. †Archexyelinae Rasnitsyn 1964
+
G. †Archexyela Riek 1955
G. †Asioxyela Rasnitsyn 1964
G. †Dinoxyela Rasnitsyn 1969
G. †Euryxyela Rasnitsyn 1964
G. †Ferganoxyela Rasnitsyn 1969
G. †Leioxyela Rasnitsyn 1969
G. †Lithoxyela Rasnitsyn 1969
G. †Madygenius Rasnitsyn 1969
G. †Moltenia Schlüter 2000
G. †Oryctoxyela Rasnitsyn 1969
G. †Triassoxyela Rasnitsyn 1964
G. †Xaxexis Pagliano and Scaramozzino 1990
+
Invalid names: Euryxyela Hong 1984 [replaced]
G. †Xiphoxyela Rasnitsyn 1969
G. †Xyelinus Rasnitsyn 1964
Subfm. †Daohugoinae Rasnitsyn and Zhang 2004
+
G. †Daohugoa Rasnitsyn and Zhang 2004
Subfm. Macroxyelinae Ashmead 1898
+
Tr. †Angaridyelini Rasnitsyn 1966
G. †Bolboxyela Rasnitsyn 1990
Tr. †Ceroxyelini Rasnitsyn 1966
Tr. †Gigantoxyelini Rasnitsyn 1969
Tr. Macroxyelini Ashmead 1898
Tr. Xyeleciini Benson 1945
Subfm. †Madygellinae Rasnitsyn 1969
+
G. †Chubakka Kopylov 2014
G. †Madygella Rasnitsyn 1969
G. †Samarkandykia Kopylov 2014
G. †Sirecomima Rasnitsyn 1969
Subfm. Xyelinae Newman 1834
+
G. †Aequixyela Wang et al. 2014
Tr. †Liadoxyelini Rasnitsyn 1966
G. Pleuroneura Konow 1897
Tr. Xyelini Newman 1834
Invalid names: Aculeata Scopoli 1763 [synonym], Cleptes steenstrupi Beck 1836 [nomen nudum], Piezata Fabricius 1798 [synonym], Pseudisobrachium oligocenicum Theobald 1937 [nomen dubium], Shandongodes necrodes Zhang 1985 [nomen dubium], Siricomorpha Rasnitsyn 1980 [invalid subgroup], Symphyta Gerstaecker 1867 [invalid subgroup]
Diagnosis
No diagnoses are available
Measurements
No measurements are available
Composition: chitinc
Environment: terrestrialc
Locomotion: actively mobilep
Created: 2017-04-17 09:56:18
Modified: 2017-04-17 09:56:18
Source: c = class, p = phylum
References: Kiessling 2004, Bush and Bambach 2015

Age range

Maximum range based only on fossils: base of the Westphalian C to the top of the Holocene or 314.60000 to 0.00000 Ma
Minimum age of oldest fossil (stem group age): 306.95 Ma

Collections (1013 total)

Oldest occurrences

Time interval Ma Country or state Original ID and collection number
Westphalian C - Westphalian D314.6 - 306.95France Avioxyela gallica (122504)
Ladinian242.0 - 237.0Kyrgyzstan (Osh) Madygella analoga, Xyelinus majus, Leioxyela grandis, Leioxyela mitis, Leioxyela mollis, Xiphoxyela procrusta, Asioxyela paurura, Oryctoxyela triassica, Dinoxyela armata, Madygenius extraradius, Lithoxyela fenestralis (110763) Madygella bashkuevi, Madygella aristovi, Madygella levivenosa, Samarkandykia ryzhkovae, Samarkandykia shmakovi (163986) Madygella kurochkini, Chubakka madygensis (163987) Triassoxyela foveolata (113468) Triassoxyela orycta, Oryctoxyela anomala, Madygenius primitivus, Ferganoxyela destructa, Ferganoxyela sogdiana (106088) Xyelinus angustiradius, Triassoxyela kirgizica, Triassoxyela orycta, Euryxyela lata, Asioxyela antiqua, Triassoxyela striata, Asioxyela parvula, Triassoxyela grandipennis, Asioxyela smilodon, Euryxyela euryptera (106090)
Carnian237.0 - 228.0Argentina Potrerilloxyela menendezi (151962)
Carnian237.0 - 228.0Japan (Yamaguchi) Xyeloidea indet., Madygella humioi (106482)
Carnian237.0 - 228.0South Africa (Eastern Cape) Moltenia rieki (23142)
Norian228.0 - 208.5Australia (Queensland) Archexyela crosbyi (106484) Archexyela ipswichensis (156521)
Sevatian212.0 - 205.6Germany Sepulenia sp. (123417)
Hettangian - Sinemurian201.3 - 190.8Kyrgyzstan (Tonskiy) Kirghizoxyela mirabilis (105087) Sogutia liassica (105084)
Pliensbachian190.8 - 182.7Kyrgyzstan Liadoxyela praecox (114701)
Early/Lower Toarcian183.0 - 182.0Germany (Niedersachsen) Liadobracona raduhna (139222) Sippelipterus liasinus (114995) Thilopterus lampei (154300) Xyelula benderi (154299)
Early/Lower Toarcian183.0 - 182.0Germany Xyelula benderi, Ephialtitidae indet. (114991)
Early/Lower Toarcian183.0 - 182.0Germany (Mecklenburg-Vorpommern) Grimmaratavites mirabilis (114993) Xyelula benderi (154298)
Early/Lower Toarcian183.0 - 182.0Germany (Bavaria) Xyelula benderi (117794)
Falciferum183.0 - 182.0Luxembourg Symphyta indet., Pseudoxyelocerus bascharagensis (105427)
Toarcian182.7 - 174.1Germany Xyelula benderi, Brigittepterus brauckmanni, Liadobracona raduhna (104056)
Toarcian182.7 - 174.1India (Andhra Pradesh) Xyelula alexandri (79229)
Toarcian182.7 - 174.1India (Adilabad) Kotaphialtites frankmortoni (117538)
Toarcian182.7 - 174.1Kyrgyzstan (Batken) Onokhoius sogdianus, Proapocritus praecursor, Sagulyda magna, Sagulyda ferganica, Ferganolyda sogdiana, Ferganolyda radialis, Ferganolyda cubitalis (105003) Sagulyda arcuata (110817) Shurabisca liassica, Sepulenia syricta, Liasirex sogdianus (105124)
Toarcian182.7 - 174.1Russian Federation (Irkutsk) Liadoxyela iensis (104995)
Bajocian - Bathonian170.3 - 166.1Mongolia (Bayan-Khongor) Parabakharius parvus (124340) Xyelula bakharensis (114615)