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Copemys longidens

Mammalia - Rodentia - Cricetidae

Taxonomy
Peromyscus longidens was named by Hall (1930). Its type specimen is UCCVP 28502, a partial skull (with M1 and mandible dext. carrying complete dentition), and it is a 3D body fossil. Its type locality is Rodent Hill, which is in a Barstovian terrestrial horizon in the Barstow Formation of California.

It was recombined as Copemys longidens by Clark et al. (1964) and Lindsay (1972).

Synonyms
  • Miochomys niobrariensis was named by Hoffmeister (1959) [genotype]. Its type specimen is UCMP 36103, a maxilla (fragment maxilla sin. with M1), and it is a 3D body fossil. Its type locality is Fort Niobrara, which is in a Barstovian terrestrial horizon in the Valentine Formation of Nebraska. It is the type species of Miochomys.

    It was synonymized subjectively with Copemys longidens by Lindsay (1972); it was recombined as Copemys niobrariensis by Voorhies (1990).
Synonymy list
YearName and author
1930Peromyscus longidens Hall p. 315 figs. Figs 5-7
1959Miochomys niobrariensis Hoffmeister p. 679 figs. fig. 1A
1964Copemys longidens Clark et al.
1972Copemys longidens Lindsay
1990Copemys niobrariensis Voorhies

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RankNameAuthor
kingdomAnimalia()
Bilateria
EubilateriaAx 1987
Deuterostomia
phylumChordataHaeckel 1874
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
subclassSarcopterygii()
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda
Reptiliomorpha
Anthracosauria
subclassAmphibiosauriaKuhn 1967
Cotylosauria()
Amniota
subclassSynapsida
Therapsida()
infraorderCynodontia()
Mammaliamorpha
Mammaliaformes
RankNameAuthor
classMammalia
Theriamorpha(Rowe 1993)
Theriiformes()
Trechnotheria
Cladotheria
Zatheria
subclassTribosphenida()
subclassTheria
Eutheria()
Placentalia
Boreoeutheria
EuarchontogliresMurphy et al. 2001
GliriformesWyss and Meng 1996
Glires()
Simplicidentata()
orderRodentiaBowdich 1821
infraorderMyodontaSchaub 1958
superfamilyMuroideaIlliger 1811
Eumuroida
familyCricetidaeFischer von Waldheim 1817
genusCopemys
specieslongidens()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Copemys longidens Hall 1930
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Invalid names: Miochomys niobrariensis Hoffmeister 1959 [synonym]
Diagnosis
ReferenceDiagnosis
E. R. Hall 1930Size large; tooth row, and individual teeth, long and relatively narrow; molars short crowned and without accessory cusps; Mm3 relatively large; medial border of m3 with a posterior pit as well as an naterior reentrant valley; space between m3 and coronoid process of medium width for Peromyscus, relatvely flat and with one large foramen.
D. F. Hoffmeister 1959 (Miochomys niobrariensis)A cricetid rodent in which the cheek teeth have both the lingual and
labial cusps elevated and of nearly the same height; cingular portion of the cusps wear least; reentrant angles of two sides of tooth
nearly opposite and nearly continuous as transverse valleys across low central axis; the two transverse valleys across tooth give
M1 the appearance of three separate segments,
namely (1) anterocone, (2) protocone-paracone, (3) hypocone-metacone (for terminology, see Hoffmeister, 1951 :11); anterocone
tends to be isolated from other cusps; mesoloph very weakly developed, but extending almost to cingulum; posteroloph
apparently present, but, if so, weakly developed and fusing with posterior border of metacone.
Maxilla.- The ventral two-thirds of the left infraorbital plate and that portion of the maxilla bearing M1-2 are preserved. The
anterior border of the infraorbital plate projects increasingly anteriad as it ascends dorsalIy, and in this detail resembles the
plate in Oryzomys more than that of any other Nearctic genus. The anterior border of the plate is only slightly more medial
than its posterior border, and if the structure is similar otherwise to Recent forms,this would result in a wide infraorbital fissure.
The outer surface of the plate is more concave than in other forms, being similar to Oryzomys in th is respect. No tubercle or
fossa is present ventral to the infraorbital
plate for attachment of M. masseter superficialis.
Probably this muscle was weakly developed;
it did not necessarily attach elsewhere.
The absence of this tubercle is quite in distinction to the condition in Reithrodontomys (subgenus Reithrodontomys), Reithrodontomys
(subgenus Aporodon), and Baiomys. The first molar is relatively farther posterior, in relation to the infraorbital
plate, than in ot her Recent genera of Nearctic cricetid rodents.