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Merycoidodon culbertsoni

Mammalia - Artiodactyla - Merycoidodontidae

Taxonomy
Merycoidodon culbertsonii was named by Leidy (1848) [genotype]. It is a 3D body fossil. Its type locality is Northwestern End of Seamen Hills (Chadronian), which is in a Chadronian terrestrial horizon in Wyoming.

It was recombined as Oreodon culbertsoni by Leidy (1852), Leidy (1857), Leidy (1869), Gaudry (1878), Cope (1884), Scott (1890), Osborn and Wortman (1894), Stewart (1897), Weber (1904), Zittel and Schlosser (1911), Moodie (1915), Barbour and Cook (1917), O'Harra (1920) and Zittel and Schlosser (1923); it was recombined as Oreodon culbertsonii by Leidy (1853); it was corrected as Merycoidodon culbertsoni by Douglass (1906) and Stevens and Stevens (1996); it was considered a nomen vanum by Lander (1998).

Synonyms
Synonymy list
YearName and author
1848Merycoidodon culbertsonii Leidy p. 49
1851Cotylops speciosa Leidy
1851Oreodon priscum Leidy
1851Oreodon robustum Leidy
1852Oreodon culbertsoni Leidy
1853Oreodon culbertsonii Leidy
1857Oreodon culbertsoni Leidy p. 89
1869Oreodon culbertsoni Leidy
1878Oreodon culbertsoni Gaudry
1884Oreodon culbertsoni Cope p. 512–513
1890Oreodon culbertsoni Scott
1894Oreodon culbertsoni Osborn and Wortman
1897Oreodon culbertsoni Stewart
1902Oreodon robustum Douglass p. 264
1903Oreodon macrorhinus Douglass
1904Oreodon culbertsoni Weber
1906Merycoidodon culbertsoni Douglass p. 565
1907Merycoidodon macrorhinus Douglass p. 822
1908Merycoidodon culbertsonii Lambe
1910Merycoidodon culbertsonii Osborn
1911Oreodon culbertsoni Zittel and Schlosser
1912Merycoidodon culbertsonii Stromer
1913Merycoidodon culbertsonii Scott
1915Oreodon culbertsoni Moodie
1917Oreodon culbertsoni Barbour and Cook
1920Oreodon culbertsoni O'Harra
1921Merycoidodon culbertsonii Thorpe
1921Merycoidodon macrorhinus Thorpe
1923Merycoidodon culbertsonii Thorpe
1923Oreodon culbertsoni Zittel and Schlosser
1924Merycoidodon culbertsonii Loomis
1925Merycoidodon culbertsonii Loomis
1928Merycoidodon culbertsonii Weber and Abel
1930Merycoidodon periculorum Bump and Loomis
1937Merycoidodon culbertsonii Thorpe
1940Merycoidodon culbertsonii Scott p. 669
1940Merycoidodon macrorhinus Scott p. 676
1953Merycoidodon culbertsonii Galbreath
1954Prodesmatochoerus meekae Schultz and Falkenbach p. 226 figs. 18, 22, 24-25
1967Merycoidodon lewisi Clark and Beerbower
1968Genetochoerus periculorum Schultz and Falkenbach
1968Merycoidodon macrorhinus Schultz and Falkenbach
1968Otionohyus vanderpooli Schultz and Falkenbach
1968Merycoidodon culbertsonii Schultz and Falkenbach p. 38 figs. 1-3, 19-23, 51
1968Paramerycoidodon georgei Schultz and Falkenbach p. 87 figs. 6-9, 19-23, 52
1968Otionohyus wardi Schultz and Falkenbach p. 107 figs. 10, 11, 19-23, 52
1968Otionohyus vanderpooli Schultz and Falkenbach p. 115 figs. 10, 19, 22
1996Merycoidodon culbertsoni Stevens and Stevens p. 518 figs. Figures 2B, 3B; Tables 1-7; Appendix B
1998Prodesmatochoerus lewisi Lander
1998Prodesmatochoerus macrorhinus Lander
1998Prodesmatochoerus periculorum Lander
2007Merycoidodon culbertsonii Stevens and Stevens p. 158

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
OlfactoresJefferies 1991
subphylumVertebrata
Gnathostomata()
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
RankNameAuthor
Amniota
Synapsida()
Therapsida()
infraorderCynodontia()
Epicynodontia
infraorderEucynodontia
Probainognathia
Mammaliamorpha
Mammaliaformes
classMammalia
orderArtiodactyla()
suborderTylopoda()
superfamilyOreodontoidea
familyMerycoidodontidaeThorpe 1923
subfamilyMerycoidodontinae
genusMerycoidodon
speciesculbertsoni()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
C. B. Schultz and C. H. Falkenbach 1954 (Prodesmatochoerus meekae)SKULL: Slightly larger than in holotype of P. natronensis, slightly smaller than in cotypes of Subdesmatochoerus socialis; exoccipital foramina elongated; infraorbital foramen above P3.
MANDIBLE: Postsymphysis below anterior portion of P8
DENTITION: Series longer than those of the holotype of P. natronensis.
LIMBS: Larger than in P. natronensis.
M. S. Stevens and J. B. Stevens 1996nial characterization defines the genus-subgenus. The quite variable Subnasal L. averages 49% of the Nasal L., the premo- lars are unreduced with an average pl_P41pl_M3 of 50% (Table 3), the APM3/TM3 ratio averages 0.99 (Table 5), the M3 is brachydont-submesodont with an average APM3/Ht. M3 ratio of 1.72 (Table 4), and TM3/Ht. M3 averages 1.75 (Table 6). M. culbertsoni differs from Merycoidodon presidioensis by a signifi- cantly smaller size (based on P 1-M 3 ,P1-M3,and associated measurements), more inclined premaxillae, wider, lower muzzle, and more posterior infraorbital foramina . M. culbertsoni differs from M. bullatus by small, instead of inflated auditory bullae, and a slightly but significantly narrower P.O.C. and lower crowned M3, otherwise similar.