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Leptocyon vafer

Mammalia - Carnivora - Canidae

Taxonomy
Canis vafer was named by Leidy (1858). It is a 3D body fossil. Its type locality is Niobrara River (coll. Hayden 1857), which is in a Miocene terrestrial horizon in the Loop Fork Formation of Nebraska.

It was recombined as Tephrocyon vafer by Matthew (1909); it was recombined as Vulpes vafer by Stirton (1935) and Macdonald (1948); it was recombined as Leptocyon vafer by Matthew (1918), Galbreath (1953), Storer (1975), Voorhies (1990) and Tedford et al. (2009).

Synonymy list
YearName and author
1858Canis vafer Leidy p. 21
1890Canis vafer Scott and Osborn p. 69
1909Tephrocyon vafer Matthew
1918Leptocyon vafer Matthew
1919Canis vafer Merriam
1935Vulpes vafer Stirton
1948Vulpes vafer Macdonald
1953Leptocyon vafer Galbreath
1975Leptocyon vafer Storer
1990Leptocyon vafer Voorhies
2009Leptocyon vafer Tedford et al. p. 34 figs. 3, 7, 9C–E, G, H, 14G–H, 15A–G, 16A–G, 17, 52, 61A; appendices 2, 4

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
OlfactoresJefferies 1991
subphylumVertebrata
Gnathostomata()
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Synapsida()
Therapsida()
infraorderCynodontia()
RankNameAuthor
Epicynodontia
infraorderEucynodontia
Probainognathia
Mammaliamorpha
Mammaliaformes
classMammalia
subclassTribosphenida()
infraclassEutheria()
Placentalia
Laurasiatheria
Scrotifera
Ferae()
CarnivoramorphaWyss and Flynn 1993
CarnivoraformesFlynn et al.
orderCarnivoraBowditch 1821
suborderCaniformiaKretzoi 1943
superfamilyCanoideaSimpson 1931
familyCanidaeFischer 1817
subfamilyCaninaeGill 1872
genusLeptocyon
speciesvafer()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
R. H. Tedford et al. 2009Distinguished from L. leidyi by its significantly larger size (95% confidence interval for first lower molars of these taxa nearly separated;fig. 7), m2 shorter relative to m1 (fig. 7) with enlarged ante- rolabial cingulum and large metaconid, cranium lacks postparietal foramina, shows reduced cerebellar exposure, and nasals do not extend posterior to maxillary-frontal suture. Distinguished from L. tejonensis in longer muzzle and more widely spaced and anteroposteriorly longer premolars with stronger accessory cusps. L. vafer differs from L. matthewi in significantly smaller size, with no overlap of m1 95% confidence intervals (fig. 7) nasals do not extend poste- rior to maxillary-frontal suture, M2 lacks postprotocrista, m1 lacks hypoconulid shelf, and entoconid not enlarged.