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Paramerychyus harrisonensis

Mammalia - Artiodactyla - Merycoidodontidae

Taxonomy
Merychyus harrisonensis was named by Peterson (1907). Its type specimen is CM 1341, a partial skull (a fairly well preserved skull which has been somewhat crushed vertically and is without the lower jaws), and it is a 3D body fossil. Its type locality is Van Tassel (Lower), which is in a Harrisonian terrestrial horizon in Wyoming. It is the type species of Paramerychyus.

It was recombined as Ticholeptus harrisonensis by Loomis (1923); it was recombined as Merycoides harrisonensis by Lander (1998); it was recombined as Paramerychyus harrisonensis by Schultz and Falkenbach (1947) and Stevens and Stevens (2007).

Synonyms
Synonymy list
YearName and author
1907Merychyus harrisonensis Peterson
1923Ticholeptus harrisonensis Loomis
1935Merychyus harrisonensis Schlaikjer
1947Paramerychyus harrisonensis Schultz and Falkenbach p. 248 figs. 1, 9
1950Hypsiops luskensis Schultz and Falkenbach p. 122 figs. 4-6, 11, 13-15
1980Phenacocoelus luskensis MacFadden p. 93
1998Merycoides harrisonensis Lander
2007Paramerychyus harrisonensis Stevens and Stevens p. 164

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
OlfactoresJefferies 1991
subphylumVertebrata
Gnathostomata()
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
RankNameAuthor
Amniota
Synapsida()
Therapsida()
infraorderCynodontia()
Epicynodontia
infraorderEucynodontia
Probainognathia
Mammaliamorpha
Mammaliaformes
classMammalia
orderArtiodactyla()
suborderTylopoda()
superfamilyOreodontoidea
familyMerycoidodontidaeThorpe 1923
subfamilyMerychyinae
genusParamerychyus
speciesharrisonensis()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
C. B. Schultz and C. H. Falkenbach 1947SKULL:Larger than any known examples of Merychyus; smaller than known specimens of Ticholeptus; approximate size of P. relictus; lacrimal fossa moderately large and deep; prelacrimal vacuity present (in the holotype the very large prelacrimal vacuity may be due to damage of the area; in the referred skull the vacuity is much smaller); malar moderately robust, with a slight but gradual upward trend posteriorly; zygomatic arch of medium construction with shallow rise posteriorly; orbits round, looking forward and upward; postorbital pillar quite heavy; bulla covering large area (Peterson described the bulla as "flask-like in form with a conical swelling on the posterior-external surface"); palatal surface vaulted.
MANDIBLE:Unknown.
DENTITION:C/ heavy but not long; pre- molars not crowded; external styles of molars prominent.
LIMBS: Unknown.
C. B. Schultz and C. H. Falkenbach 1950 (Hypsiops luskensis)SKULL: Approximately same length but narrower than H. breviceps, smaller than other species of genus; nasals retracted to area above P 1 ; anterior nasal-maxilla contact above anterior portion of P2 ; zygomatic arch elongated posteriorly and not extending upward as high as in other species of this genus, but similar to those of Phenacocoelus kayi and P. stouti (see discussion, p. 104); infraorbital foramen above anterior portion of P4; small, round, facial vacuity; other characters similar to those of H. brachymelis and H. brachymelis petersoni.
MANDIBLE: Characters and outline typical of genus; postsymphysis below Pa.
DENTITION:Shorter and lighter than in H. breviceps; smaller than examples of other known species of genus; p1-p 3 each with anterior intermediate crest; PrPa each with posterior intermediate crest.
LIMBS:Similar to those of H. brachymelis; considerable individual variation in length but within the expected variation of a species.