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Ankylosaurinae
Taxonomy
Ankylosaurinae was named by Nopcsa (1918). Its type is Ankylosaurus. It was considered monophyletic by Vickaryous et al. (2004).
It was assigned to Acanthopholidae by Nopcsa (1923) and Nopcsa (1923); to Nodosauridae by Nopcsa (1929), Kuhn (1966) and Maryanska (1971); and to Ankylosauridae by Russell and Chamney (1967), Tumanova (1983), Sereno (1986), Tumanova (1987), Tumanova (1993), Vickaryous and Ryan (1997), Sereno (1997), Sereno (1998), Sereno (1999), Ford (2000), Vickaryous et al. (2001), Vickaryous et al. (2004), Han et al. (2014), Arbour et al. (2014), Arbour and Currie (2016), Yang et al. (2017), Arbour and Mallon (2017), Penkalski and Tumanova (2017), Arbour and Evans (2017), Penkalski (2018) and Zheng et al. (2018).
It was assigned to Acanthopholidae by Nopcsa (1923) and Nopcsa (1923); to Nodosauridae by Nopcsa (1929), Kuhn (1966) and Maryanska (1971); and to Ankylosauridae by Russell and Chamney (1967), Tumanova (1983), Sereno (1986), Tumanova (1987), Tumanova (1993), Vickaryous and Ryan (1997), Sereno (1997), Sereno (1998), Sereno (1999), Ford (2000), Vickaryous et al. (2001), Vickaryous et al. (2004), Han et al. (2014), Arbour et al. (2014), Arbour and Currie (2016), Yang et al. (2017), Arbour and Mallon (2017), Penkalski and Tumanova (2017), Arbour and Evans (2017), Penkalski (2018) and Zheng et al. (2018).
Synonymy list
Year | Name and author |
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1918 | Ankylosaurinae Nopcsa |
1923 | Ankylosaurinae Nopcsa p. 126 |
1929 | Ankylosaurinae Nopcsa p. 71 |
1966 | Ankylosaurinae Kuhn p. 105 |
1967 | Ankylosaurinae Russell and Chamney p. 11 |
1971 | Ankylosaurinae Maryanska p. 47 |
1983 | Ankylosaurinae Tumanova p. 110 |
1986 | Ankylosaurinae Sereno |
1987 | Ankylosaurinae Tumanova p. 15 |
1993 | Ankylosaurinae Tumanova p. 119 |
1997 | Ankylosaurinae Sereno p. 443 |
1997 | Ankylosaurinae Vickaryous and Ryan p. 491 |
1998 | Ankylosaurinae Sereno p. 61 |
1999 | Ankylosaurinae Sereno p. 2138 fig. 2 |
2000 | Ankylosaurinae Ford p. 175 |
2001 | Ankylosaurinae Vickaryous et al. p. 1774 |
2004 | Ankylosaurinae Vickaryous et al. p. 364 |
2014 | Ankylosaurinae Arbour et al. |
2014 | Ankylosaurinae Han et al. p. 14 fig. 11 |
2016 | Ankylosaurinae Arbour and Currie p. 405 |
2017 | Ankylosaurinae Arbour and Evans p. 7 |
2017 | Ankylosaurinae Arbour and Mallon p. 768 |
2017 | Ankylosaurinae Penkalski and Tumanova |
2017 | Ankylosaurinae Yang et al. p. 786 fig. 5 |
2018 | Ankylosaurinae Penkalski p. 264 |
2018 | Ankylosaurinae Zheng et al. |
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If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.
Subfm. †Ankylosaurinae Nopcsa 1918
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Tr. †Ankylosaurini Arbour and Currie 2016
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Tr. †Euoplocephalini Penkalski 2018
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G. †Anodontosaurus Sternberg 1929
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†Anodontosaurus inceptus Penkalski 2018
†Anodontosaurus lambei Sternberg 1929
G. †Euoplocephalus Lambe 1910
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†Euoplocephalus tutus Lambe 1902
Invalid names: Stereocephalus Lambe 1902 [replaced]
G. †Scolosaurus Nopcsa 1928 [thorn lizard]
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†Scolosaurus cutleri Nopcsa 1928
†Scolosaurus thronus Penkalski 2018
G. †Minotaurasaurus Miles and Miles 2009
G. †Pinacosaurus Gilmore 1933
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†Pinacosaurus grangeri Gilmore 1933
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Invalid names: Pinacosaurus ninghsiensis Young 1935 [synonym], Syrmosaurus viminocaudus Maleev 1952 [synonym]
†Pinacosaurus mephistocephalus Godefroit et al. 1999
Invalid names: Syrmosaurus Maleev 1952 [synonym]
G. †Saichania Maryanska 1977
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†Saichania chulsanensis Maryanska 1977
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Invalid names: Shanxia tianzhenensis Barrett et al. 1998 [synonym], Tianzhenosaurus youngi Pang and Cheng 1998 [synonym]
Invalid names: Shanxia Barrett et al. 1998 [synonym], Tianzhenosaurus Pang and Cheng 1998 [synonym]
G. †Tarchia Maryanska 1977
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†Tarchia kielanae Maryanska 1977
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Invalid names: Minotaurasaurus ramachandrani Miles and Miles 2009 [synonym]
†Tarchia teresae Penkalski and Tumanova 2017
†Tarchia tumanovae Park et al. 2021
Diagnosis
Reference | Diagnosis | |
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T. A. Tumanova 1987 | Ankylosaurs with wide skull, width of which is slightly less or equal to the length; broad snout; orbits oriented slightly forward; angle of orbital plane with long axis of skull about 30o; anterior wall of pterygoids sometimes inclined somewhat sharply posteriorly, but more often inclined anteriorly; occipital condyle oval; basipterygoid process contacts with pterygoid by suture; orbits in posterior half of skull length. |