Basic info Taxonomic history Classification Relationships
Morphology Ecology and taphonomy External Literature Search Age range and collections

Euoplocephalus tutus

Reptilia - Ankylosauridae

Synonymy list
YearName and author
1902Stereocephalus tutus Lambe p. 55 figs. Pl. 10, 11, 21
1905Stereocephalus tutus Lambe p. 23
1908Stereocephalus tutus Brown p. 197
1910Euoplocephalus tutus Lambe p. 151
1914Ankylosaurus tutus Brown p. 378
1914Euoplocephalus tutus Lambe p. 132
1915Palaeoscincus tutus Hennig p. 14
1915Euoplocephalus tutus Lambe p. 117
1917Stereocephalus tutus Sternberg p. 210
1919Euoplocephalus tutus Lambe p. 39
1921Stereocephalus tutus Gilmore p. 589
1923Euoplocephalus tutus Gilmore pp. 47-48
1924Euoplocephalus tutus Parks p. 7
1927Euoplocephalus tutus Huene p. 261
1928Euoplocephalus tutus Sternberg p. 100
1930Euoplocephalus tutus Russell p. 149
1964Ankylosaurus tutus Kuhn p. 45
1964Euoplocephalus tutus Russell p. 13
1965Europlocephalus tutus Simmons p. 70
1966Euoplocephalus tutus Russell p. 19
1978Euoplocephalus tutus Coombs p. 148 fig. 3
1986Euoplocephalus tutus Coombs, Jr. p. 170
1986Euoplocephalus tutus Sereno
1987Euoplocephalus tutus Tumanova
1997Euoplocephalus tutus Eberth p. 201
1997Euoplocephalus tutus Vickaryous and Ryan p. 490
2000Euoplocephalus tutus Currie p. 277
2000Euoplocephalus tutus Ford p. 175
2001Euoplocephalus tutus Barrett p. 40 fig. 2.8
2001Euoplocephalus tutus Carpenter p. 457
2001Euoplocephalus tutus Eberth et al. p. 58
2001Euoplocephalus tutus Ford and Kirkland p. 254
2003Euoplocephalus tutus Garcia and Pereda-Suberbiola p. 161
2004Euoplocephalus tutus Vickaryous et al. p. 365
2005Euoplocephalus tutus Currie p. 5
2005Euoplocephalus tutus Ryan and Evans p. 315
2006Euoplocephalus tutus Sullivan and Fowler p. 259
2006Euoplocephalus tutus Vickaryous p. 1011
2007Euoplocephalus tutus Arbour p. 41A
2008Euoplocephalus tutus Burns p. 1107
2008Euoplocephalus tutus Longrich p. 994
2009Euoplocephalus tutus Osi and Makádi p. 235 fig. 6
2011Euoplocephalus tutus Burns and Sullivan p. 177
2012Euoplocephalus tutus Arbour et al. p. 55
2013Euoplocephalus tutus Arbour and Currie p. 31
2013Euoplocephalus tutus Kirkland et al. p. 7
2013Euoplocephalus tutus Penkalski and Blows p. 178
2014Euoplocephalus tutus Arbour et al. p. 2
2014Euoplocephalus tutus Han et al. p. 12 fig. 7
2018Euoplocephalus tutus Penkalski p. 264

Is something missing? Join the Paleobiology Database and enter the data

RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
subphylumVertebrata
superclassGnathostomata
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
RankNameAuthor
Eosuchia()
Neodiapsida
SauriaGauthier 1984
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Ornithischia()
Genasauria
Thyreophora()
Ankylosauria()
familyAnkylosauridae
subfamilyAnkylosaurinaeNopcsa 1918
tribeEuoplocephalini
genusEuoplocephalus
speciestutus()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
T. A. Tumanova 1987Premaxillary beak very wide, anterior blunt, width equal to or exceeding space between rear [maxillary] teeth; maxillary shelves well developed; plane surface of palatal bone tilted laterally; anterior wall of medial section of pterygoids slanted forward; paroccipital processes laterally oriented; occipital condyle broad oval, oriented posteroventrally; quadrate and paroccipital processes not fused; occipital plane tilted slightly forward.
V. M. Arbour and P. J. Currie 2013Differs from Anodontosaurus lambei and Scolosaurus cutleri in lacking subcircular caputegulae at the bases of the quadratojugal and squamosal horns (postocular caputegulae); differs from Anodontosaurus lambei in lacking interstitial osteoderms at the bases of the primary osteoderms of the first cervical half ring, and in having semicircular major osteoderms in dorsal view on the tail club; differs from Dyoplosaurus acutosquameus in having laterally-directed sacral ribs; differs from Scolosaurus cutleri in having oval to subcircular-based keeled medial and lateral primary half ring osteoderms and in having a proportionately shorter postacetabular process of the ilium; differs from Ankylosaurus magniventris in having anteriorly-directed nares, and in lacking a continuous keel between the squamosal horn and supraorbitals.
P. G. Penkalski 2018A moderate-sized (5 m) ankylosaurine with well-defined scales in the frontal region; erect squamosal boss with central apex and kinked (sigmoidal) keel; skull trapezoidal in dorsal view; nasal plate large, centrally convex; some skull plates with incipient nodes or keels, mostly in nasofrontal region; supraorbital bosses pointed in dorsal view and distinct, especially in lateral view; prominent quadratojugal horn with central apex; premaxillary palate divided; humerus less robust than in other euoplocephalins; deltopectoral crest does not extend past humeral midpoint; six oval, sharply keeled, fused osteoderms with central apices on h.r.1; keels on medial and lateral osteoderms sigmoidal (kinked); free osteoderms weakly ornamented, with fine (1-2 mm) pitting; many osteoderms oval and sharply keeled; some osteoderms with offset, L-shaped or folded keel and/ or double-edged perimeter.