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Titanis walleri

Reptilia - Avetheropoda - Phorusrhacidae

Taxonomy
Titanis walleri was named by Brodkorb (1963). Its type specimen is UF 4109, a set of limb elements, and it is a 3D body fossil. Its type locality is Santa Fe River 1, which is in a Blancan terrestrial horizon in Florida.

Synonymy list
YearName and author
1963Titanis walleri Brodkorb p. 112
1998Titanis walleri Emslie
2003Titanis walleri Alvarenga and Holfing p. 76
2005Titanis walleri Gould and Quitmyer p. 205
2007Titanis walleri Bertelli et al. p. 415 figs. Table 2

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
OlfactoresJefferies 1991
subphylumVertebrata
Gnathostomata()
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
Eosuchia()
Neodiapsida
SauriaGauthier 1984
RankNameAuthor
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Saurischia()
Theropoda()
Neotheropoda
AverostraPaul 2002
Tetanurae
orderAvetheropoda
suborderCoelurosauria
Maniraptora
Paraves
classAves
suborderCariamaeFürbringer 1888
familyPhorusrhacidae()
subfamilyPhorusrhacinaeAmeghino 1899
genusTitanis
specieswalleri

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
P. Brodkorb 1963Referable to the family Phorusrhacidae,subfamily Phorus- rhacinae, on the basis of: gigantic size; distal foramen of tarsometatarsus bifurcate,withforamenforextensorbrevisdigitiIV openingthroughdistal end of shelf bounding outer intermetatarsalspace (confluent with foramen for tibialis anticus artery on plantar surface of shaft in Brontornithidae, separate but on plantar face in Rheidae); lower end of shaft not bilaterally expanded (flaring toward roots of trochleae in Rheidae); plantar surface of shaft flat above trochleae (concave in comparablegroups); trochleae nearly on a plane (strongly arched in Psilopteridae and Rheidae); poste- rior wings of trochleae obsolete (well developed in Psilopteridae)