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Edmontosaurus annectens

Reptilia - Hadrosauridae

Taxonomy
Claosaurus annectens was named by Marsh (1892). Its type specimen is USNM 2414, a skeleton, and it is a 3D body fossil. Its type locality is Schneider Creek, which is in a Maastrichtian terrestrial horizon in the Lance Formation of Wyoming.

It was synonymized subjectively with Thespesius occidentalis by Lucas (1900), Hay (1902) and Gilmore (1915); it was recombined as Trachodon annectens by Hatcher (1902), Lucas (1904), Osborn (1911), Brown (1913), Brown (1913), Lambe (1913), Brown (1914), Brown (1914), Lambe (1914), Versluys (1923), Lull (1924), Huene (1927) and Drevermann (1928); it was recombined as Thespesius annectens by Gilmore (1924) and Moodie (1926); it was recombined as Anatosaurus annectens by Lull and Wright (1942), Gregory (1948), Vialli (1960), Tatarinov (1964), Russell (1964), Kuhn (1964), Ostrom (1965) and Swinton (1970); it was misspelled as Cliosaurus annectens by Mansel-Pleydell (1902), Beecher (1902), Wieland (1903), Brown (1910), Brown (1912), Lambe (1918), Johnson (1931) and Vickaryous and Ryan (1997); it was recombined as Edmontosaurus annectens by Lambe (1917), Weishampel and Horner (1990), Hunt and Lucas (1992), Horner (1992), Turner (1997), Currie and Padian (1997), Eberth (1997), Godefroit et al. (1998), Horner et al. (2004), Godefroit et al. (2004), Lund and Gates (2006), Prieto-Márques et al. (2006), Manning et al. (2008), Evans et al. (2009), Prieto-Márquez (2010), McDonald et al. (2010), Prieto-Marquez and Salinas (2010), Prieto-Marquez (2010), Prieto-Márquez and Wagner (2010), Campione and Evans (2011), Prieto-Márquez (2011), Brown and Druckenmiller (2011), Prieto-Márquez (2012), Godefroit et al. (2012), Ramírez-Velasco et al. (2012), Zheng et al. (2013), Prieto-Marquez (2013), Murphy et al. (2013), Bolotsky et al. (2014), Ramírez-Velasco et al. (2014), Bell (2014), Prieto-Márquez and Wagner (2014), Campione (2014), Bell et al. (2014), Gates et al. (2014), Bell et al. (2014), Lehman et al. (2016), Prieto-Márquez and Gutarra (2016), Mori et al. (2016) and Cruzado-Caballero and Powell (2017).

Synonyms
Synonymy list
YearName and author
1890Trachodon longiceps Marsh
1892Claosaurus annectens Marsh p. 435
1893Claosaurus annectens Hatcher p. 143
1896Claosaurus annectens Marsh figs. Pl. 72-74
1898Claosaurus annectens Williston p. 70
1898Claosaurus annectens Woodward p. 209
1899Claosaurus annectens Lambe p. 188
1900Claosaurus annectens Nopcsa p. 557
1900Hadrosaurus longiceps Nopcsa p. 557
1902Claosaurus annectens Beecher p. 313
1902Trachodon annectens Hatcher p. 385
1902Trachodon longiceps Hay p. 504
1902Cliosaurus annectens Mansel-Pleydell p. lxxxiv
1903Claosaurus annectens Wieland p. 216
1904Trachodon annectens Lucas p. 317
1910Claosaurus annectens Brown p. 270
1911Trachodon annectens Osborn p. 7
1912Claosaurus annectens Brown p. 105
1913Trachodon annectens Brown p. 387
1913Trachodon annectens Lambe p. 25
1914Trachodon annectens Brown p. 374
1914Trachodon annectens Lambe pp. 133-134
1917Edmontosaurus annectens Lambe
1918Claosaurus annectens Lambe p. 139
1923Trachodon annectens Versluys p. 1
1924Thespesius annectens Gilmore p. 14
1924Thespesius edmontoni Gilmore pp. 14-15 figs. Pl. 2-5
1924Trachodon annectens Lull p. 270
1926Thespesius annectens Moodie p. 74
1927Trachodon annectens Huene p. 260
1927Trachodon longiceps Huene p. 260
1927Thespesius edmontoni Huene p. 261
1928Trachodon annectens Drevermann p. 202–203
1930Thespesius edmontoni Riabinin p. 14
1930Thespesius edmontoni Russell p. 151
1931Claosaurus annectens Johnson p. 374
1933Thespesius edmontoni Russell p. 129
1942Anatosaurus annectens Lull and Wright pp. 154-156
1942Anatosaurus edmontoni Lull and Wright pp. 161-162 fig. 57
1942Anatosaurus longiceps Lull and Wright pp. 163-164
1948Anatosaurus annectens Gregory p. 30
1960Anatosaurus annectens Vialli p. 176
1964Anatosaurus annectens Kuhn p. 14
1964Anatosaurus edmontoni Kuhn p. 14
1964Anatosaurus longiceps Kuhn p. 15
1964Anatosaurus edmontoni Russell p. 15
1964Anatosaurus annectens Russell p. 17
1964Anatosaurus edmontoni Tatarinov p. 562 fig. 609
1964Anatosaurus annectens Tatarinov p. 564
1965Anatosaurus annectens Ostrom p. 39
1965Anatosaurus longiceps Ostrom p. 39
1966Anatosaurus edmontoni Russell p. 16
1967Edmontosaurus edmontoni Russell and Chamney p. 11
1970Anatosaurus annectens Swinton p. 216
1970Anatosaurus edmontoni Swinton p. 221
1979Edmontosaurus edmontoni Brett-Surman p. 561 fig. 1
1980Anatosaurus edmontoni Drake p. 196
1990Edmontosaurus annectens Weishampel and Horner p. 557
1992Edmontosaurus annectens Horner p. 119
1992Edmontosaurus annectens Hunt and Lucas p. 228
1992Anatotitan longiceps Olshevsky p. 230
1997Edmontosaurus annectens Currie and Padian p. 369
1997Edmontosaurus annectens Eberth p. 201
1997Edmontosaurus edmontoni Eberth p. 201
1997Edmontosaurus annectens Turner p. 788
1997Anatosaurus annectus Vickaryous and Ryan p. 490
1998Edmontosaurus annectens Godefroit et al. p. 26
2004Edmontosaurus annectens Godefroit et al. p. 147
2004Edmontosaurus annectens Horner et al. p. 440
2006Edmontosaurus annectens Lund and Gates p. 271
2006Edmontosaurus annectens Prieto-Márques et al. p. 937
2008Edmontosaurus annectens Manning et al. p. 647
2009Edmontosaurus annectens Evans et al. p. 1316
2010Edmontosaurus annectens McDonald et al. p. 33 fig. 39
2010Edmontosaurus annectens Prieto-Marquez p. 845
2010Edmontosaurus annectens Prieto-Marquez and Salinas p. 833
2010Edmontosaurus annectens Prieto-Márquez p. 3
2010Edmontosaurus annectens Prieto-Márquez and Wagner p. 1243 fig. 3
2011Edmontosaurus annectens Brown and Druckenmiller p. 1350
2011Edmontosaurus annectens Campione and Evans p. 9
2011Edmontosaurus annectens Prieto-Márquez p. 63
2012Edmontosaurus annectens Godefroit et al. p. 18 fig. 19
2012Edmontosaurus annectens Prieto-Márquez p. 524
2012Edmontosaurus annectens Ramírez-Velasco et al. p. 392
2013Edmontosaurus annectens Murphy et al. p. 282
2013Edmontosaurus annectens Prieto-Marquez
2013Edmontosaurus annectens Zheng et al.
2014Edmontosaurus annectens Bell p. 575
2014Edmontosaurus annectens Bell et al. p. 70
2014Edmontosaurus annectens Bell et al. p. 178
2014Edmontosaurus annectens Bolotsky et al. p. 320
2014Edmontosaurus annectens Campione p. 208
2014Edmontosaurus annectens Gates et al. p. 160
2014Edmontosaurus annectens Prieto-Márquez and Wagner p. 591
2014Edmontosaurus annectens Ramírez-Velasco et al. p. 349
2016Edmontosaurus annectens Lehman et al. p. 343
2016Edmontosaurus annectens Mori et al. p. 17
2016Edmontosaurus annectens Prieto-Márquez and Gutarra p. 144
2017Edmontosaurus annectens Cruzado-Caballero and Powell p. 12 fig. 13

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
OlfactoresJefferies 1991
subphylumVertebrata
Gnathostomata()
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
Romeriida
Diapsida()
Eosuchia()
Neodiapsida
RankNameAuthor
SauriaGauthier 1984
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Ornithischia()
Neornithischia()
Ornithopoda()
Iguanodontia()
Dryomorpha
Ankylopollexia
Styracosterna
Hadrosauriformes
Hadrosauroidea()
familyHadrosauridae
subfamilyHadrosaurinaeLambe 1918
tribeEdmontosauriniGlut 1997
genusEdmontosaurus
speciesannectens()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
N. E. Campione and D. C. Evans 2011Hadrosaurine hadrosaurid, which at large size (skull length .1 m) is differentiated from Edmontosaurus regalis by the following characteristics: presence of a very long prenarial region of skull; weakly excavated caudodorsal corner of narial vestibule; weakly developed postorbital fossa that results in a strait dorsal process of jugal.