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Grallator (Eubrontes)

Reptilia - Eubrontidae

Subtaxa
Agialopous wyomingensis, Anchisauripus (Grallator), Anchisauripus (Otouphepus) minor, Anchisauripus australis, Anchisauripus bibractensis, Anchisauripus exsertus, Anchisauripus gwyneddensis, Anchisauripus milfordensis (syn. Atreipus acadianus), Anchisauripus poolei, Anchisauripus thomasi, Anchisauripus tuberosus (syn. Brontozoum loxonyx, Brontozoum validum, Plesiornis mirabilis), Brontozoum expansum, Brontozoum isodactylum, Brontozoum thianschanicum, Deuterotrisauropus deambulator, Eubrontes (Brontozoum), Eubrontes (Grallator), Eubrontes (Ichnites), Eubrontes (Ornithichnites), Eubrontes cursorius (syn. Ornithichnites parallelus), Eubrontes fieldi, Eubrontes giganteus, Eubrontes glenrosensis, Eubrontes lyellii (syn. Anomoepus major), Eubrontes monax, Eubrontes pareschequier, Eubrontes platypus, Eubrontes sillimani (syn. Ornithichnites cuneatus, Anchisauripus dananus), Eubrontes tuberatus, Eubrontes veillonensis, Eubrontes xiyangensis, Eubrontes zigongensis, Fulicopus minor, Gigandipus caudatus, Gigandipus hei, Grallator (Anchisauripus) hitchcocki, Grallator (Anchisauripus) minusculus (syn. Plesiornis giganteus), Grallator (Eubrontes) divaricatus, Grallator (Grallator), Grallator andeolensis, Grallator angustidigitus, Grallator crassidigitus, Grallator cuneatus, Grallator damanei, Grallator deambulator, Grallator digitigradus, Grallator emeiensis, Grallator formosus, Grallator gracilis, Grallator graciosus, Grallator lacunensis, Grallator lambereshei, Grallator leribeensis, Grallator lescurei, Grallator limnosus, Grallator limosus, Grallator magnificus, Grallator matsiengensis, Grallator maximus, Grallator microiscus, Grallator minimus, Grallator minor, Grallator mokanametsongensis, Grallator molapoi, Grallator morijensis, Grallator moshoeshoei, Grallator olonensis, Grallator palustris, Grallator princeps, Grallator rapidus, Grallator rectilineus, Grallator sauclierensis, Grallator socialis, Grallator ssatoi, Grallator sulcatus, Grallator tenuis, Grallator variabilis, Grallator wuhuangensis, Grallator yemiaoxiensis, Otouphepus declivis, Plastisauropus ingens, Platysauropus robustus, Prototrisauropus deambulator, Prototrisauropus gardonensis
Species lacking formal opinion data
Sister taxa
Anchisauripus (Grallator), Eubrontes (Brontozoum), Eubrontes (Grallator), Eubrontes (Ichnites), Eubrontes (Ornithichnites), G. (Grallator)

View classification of included taxa

Synonyms
Synonymy list
YearName and author
1845Eubrontes Hitchcock p. 23
1845Fulicopus Hitchcock p. 23
1847Brontozoum Hitchcock p. 50
1848Brontozoum Hitchcock p. 169
1858Brontozoum isodactylum Hitchcock pp. 69-70
1858Grallator Hitchcock p. 72
1858Gigantitherium Hitchcock p. 93
1858Hyphepus Hitchcock pp. 97, 180
1861Brontozoum Owen p. 327
1889Brontozoum isodactylum Eyerman p. 32
1894Brontozoum isodactylum Lyman p. 205
1898Brontozoum Hitchcock p. 401
1898Gigantitherium Hitchcock p. 401
1898Grallator Hitchcock p. 401
1898Hyphepus Hitchcock p. 401
1902Brontozoum Mansel-Pleydell p. lxxviii
1904Otouphepus Cushman p. 155
1904Anchisauripus Lull pp. 486-487
1904Gigandipus Lull p. 492
1904Hyphepus Lull p. 493
1904Grallator Lull p. 494
1904Fulicopus Lull p. 505
1904Eubrontes Lull p. 510
1915Anchisauripus Lull p. 180
1915Otouphepus Lull p. 187
1915Gigandipus Lull p. 191
1915Hyphepus Lull p. 193
1915Eubrontes Lull p. 195
1915Grallator Lull p. 200
1917Anchisauripus Lull p. 114
1917Otouphepus Lull p. 114
1917Eubrontes Lull p. 115
1917Gigandipus Lull p. 115
1917Hyphepus Lull p. 115
1917Grallator Lull p. 116
1932Agialopous Branson and Mehl
1952Grallator Bock p. 402
1952Gigandipus Bock p. 403
1952Anchisauripus Bock p. 406
1952Otouphebus Bock p. 407
1953Grallator Lull p. 153
1953Anchisauripus Lull p. 166
1953Otouphepus Lull p. 176
1953Eubrontes Lull p. 178
1953Gigandipus Lull p. 184
1953Hyphepus Lull p. 186
1954Grallator Baird p. 182
1957Grallator Baird p. 453
1957Anchisauripus Baird p. 462
1958Anchisauripus Kuhn p. 22
1958Eubrontes Kuhn p. 22
1958Grallator Kuhn p. 22
1958Gigandipus Kuhn p. 23
1958Hyphepus Kuhn p. 23
1958Otouphepus Kuhn p. 23
1958Agialopous Kuhn p. 24
1963Grallator Kuhn p. 77
1963Anchisauripus Kuhn p. 79
1963Eubrontes Kuhn p. 83
1963Gigandipus Kuhn p. 84
1963Hyphepus Kuhn p. 85
1963Agialopous Kuhn p. 89
1965Deuterotrisauropus Ellenberger
1965Prototrisauropus Ellenberger
1967Eubrontes Lapparent and Montenat p. 33
1967Grallator Lapparent and Montenat p. 33
1969Agailopous Haubold p. 98
1969Anchisauripus Haubold p. 99
1969Eubrontes Haubold p. 99
1969Gigandipus Haubold p. 99
1969Grallator Haubold p. 99
1969Hyphepus Haubold p. 99
1970Qemetrisauropus Ellenberger p. 344
1970Prototrisauropus Ellenberger p. 345
1970Platysauropus Ellenberger p. 348
1970Kainotrisauropus Ellenberger p. 350
1971Eubrontes (Brontozoum) Haubold
1971Agialopous Haubold p. 64
1971Anchisauripus Haubold p. 65
1971Anchisauripus (Otouphepus) Haubold p. 65
1971Grallator Haubold p. 66
1971Eubrontes Haubold p. 75
1971Gigandipus Haubold p. 77
1971Hyphepus Haubold p. 77
1972Anchisauripus Demathieu and Haubold p. 806
1972Eubrontes Demathieu and Haubold p. 806
1972Grallator Demathieu and Haubold p. 806
1974Qemetrisauropus Ellenberger p. 16
1974Aetonychopus Ellenberger p. 17
1974Plastisauropus Ellenberger p. 17
1974Kainotrisauropus Ellenberger p. 73
1974Aetonychopus Ellenberger p. 79
1979Eubrontes Leonardi p. 244
1984Grallator Olsen and Galton p. 97
1984Grallator (Eubrontes) Olsen and Galton p. 97
1984Grallator (Anchisauripus) Olsen and Galton p. 99
1986Grallator Zhen et al. p. 2
1986Paracoelurosaurichnus Zhen et al. p. 4
1986Eubrontes Zhen et al. p. 9
1986Youngichnus Zhen et al. p. 10
1987Anchisauripus Weems p. 15
1987Eubrontes Weems p. 17
1987Chonglongpus Yang and Yang
1987Gigandipus Yang and Yang p. 19
1987Chuanchengpus Yang and Yang p. 23–24
1988Grallator Haderer p. 8
1989Grallator Zhen et al. p. 187
1989Eubrontes Zhen et al. pp. 192-193
1989Paracoelurosaurichnus Zhen et al. p. 192
1989Youngichnus Zhen et al. p. 193
1991Grallator Gierlinski p. 137
1991Grallator (Eubrontes) Gierlinski p. 137
1992Anchisauripus Lockley p. 272
1992Eubrontes Lockley p. 272
1992Grallator Lockley p. 272
1994Grallator Gierlinski and Ahlberg p. 101
1994Eubrontes Vignaud et al. p. 497
1994Grallator Vignaud et al. p. 497
1996Grallator Gierlinski p. 120
1998Grallator Hunt et al. p. 522
2000Anchisauripus Leonardi and Mietto p. 197
2000Eubrontes Leonardi and Mietto p. 197
2000Grallator Leonardi and Mietto p. 197
2004Grallator (Eubrontes) Milàn and Gierlinski p. 71
2004Anchisauripus Niedzwiedzki and Pienkowski p. 335
2006Eubrontes Lucas et al. p. 86–87
2006Grallator Lucas et al. p. 86–87
2006Eubrontes Lucas et al. p. 114
2006Grallator Lucas et al. p. 114
2006Eubrontes Weems p. 372
2006Grallator Weishampel p. 132
2006Anchisauripus Weishampel p. 133
2006Eubrontes Weishampel p. 137
2007Weiyuanpus Gao p. 342
2007Eubrontes Gao p. 345
2007Youngichnus Gao p. 345
2009Grallator Gierlinski et al. p. 227
2010Eubrontes Li et al. p. 726
2011Anchisauripus Niedźwiedzki p. 295
2011Grallator Niedźwiedzki p. 295
2011Eubrontes Niedźwiedzki p. 296
2016Eubrontes Salisbury et al. p. 36

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RankNameAuthor
kingdomAnimalia()
Triploblastica
Nephrozoa
Deuterostomia
phylumChordataHaeckel 1847
OlfactoresJefferies 1991
subphylumVertebrata
Gnathostomata()
Osteichthyes()
Sarcopterygii
subclassDipnotetrapodomorpha(Nelson 2006)
subclassTetrapodomorpha()
Tetrapoda()
Reptiliomorpha
Anthracosauria
Batrachosauria()
Cotylosauria()
Amniota
Sauropsida
classReptilia
subclassEureptilia()
RankNameAuthor
Romeriida
Diapsida()
Eosuchia()
Neodiapsida
SauriaGauthier 1984
Archosauromorpha(Huene 1946)
Crocopoda
ArchosauriformesGauthier 1986
Eucrocopoda
Archosauria()
informalAvemetatarsalia
Ornithodira
Dinosauromorpha
Dinosauriformes
Dinosauria()
Saurischia()
Theropoda()
familyEubrontidae
subgenusGrallator (Eubrontes)()
subgenusEubrontes()

If no rank is listed, the taxon is considered an unranked clade in modern classifications. Ranks may be repeated or presented in the wrong order because authors working on different parts of the classification may disagree about how to rank taxa.

Diagnosis
ReferenceDiagnosis
E. Hitchcock 1848 (Brontozoum)Foot tridactylous, pachydactylous, tubercular-clawed; inner toe shortest; all of them directed forward. Phalangeal expansions on the inner toe, two; on the middle toe, three; on the outer toe, four; corresponding to the number of phalanges, except the distal expansion, which was probably made by the two extreme phalanges. Lower extremity of the tarso-metatarsal bone double-headed; rarely making a distinct impression through the cushion beneath. Cushion sloping upwards posteriorly. Claws on the lateral toes a little outside of their axes; on the middle toe, a little towards its inner side.
R. S. Lull 1904 (Anchisauripus)Bipedal, tetradactylous; hallux rotated to the rear so as to lie in lin with the claw of the second digit, insistent, that is, inserted high on the leg so that only the tip of the claw impresses and that only when the weight is borne on the hinder part of the foot. The hallux evidently bore a well developed, subfunctional, grasping claw. Foot of moderate size with well marked phalangeal pads. Limbs of medium length. Manus and tail never impressing.
R. S. Lull 1915 (Anchisauripus)Those of the family.
R. S. Lull 1915 (Gigandipus)Those of the family, the most readily recognized being the small hallux which lies at right angles to digit II, instead of being rotated far to the rear as in Anchisauripus.
R. S. Lull 1915Those of the family.
R. S. Lull 1915 (Hyphepus)Bipedal, tetradactylous, hallux incumbent, curved sharply backward. Phalangeal pads obscurely defined; anterior digits apparently clawless.